THE ANNALS
AND
MAGAZINE OF NATURAL HISTORY,
INCLUDING
ZOOLOGY, BOTANY, AND GEOLOGY.
( BEIKG A CON'J'DlUATION OP THE 'AN"N"ALS' CO�IRJKED W[Tll LOUDON AND
CIIARLESWORTll•S I MAGAZINE OF NATURAL HISTORY,')
CONDUCTED BY
AU!El T C. L. G. GUNTHER, M.A., M.D., Ph.D., F.R.S.
WII,LIAM CARRUTHERS, F.R.S., F.L.S., F.G.S.,
ANO
WILLIAM FRANCIS, Ph.D., F.T,.S.
VOL. XVIII.-SIX' II SBHIES.
LONDON:
PRINTED AND PUilLISIIED IlY TAYLOR AND FR. NCIS.
:OLD BY SIMPKIN, MARSIIALL, JIA�ULTON, KENT, A: "D CO., LD.;
WHITTAJ{lU • . ND CO.: Il. lLLd:nE, PAIUS:
MACLACHLAN .AND STEWART, EDl�BURGII:
UODGKS, FIOGIS, AND CO., DOOLIN: AND ASll.ETI, BERLIN.
1896.
THE A�NALS
AND
MAGAZINE OF NATURAL HISTORY.
[SIXTH SERIES.]
No. 105. SEPTE�IBER 1890.
Ann. & Ma,q. N. His . Se . 6. Vol. x iii. 14
Ma mna{sji·om Celebes, Bon,eo, and he Philii>pines. :241
XXXVIIL-0n 1llam111als om Gelebes, Bo neo, and lie
Pl,ilippines ecenll!J ,·ece ed a ! e B i ish illuseum. By
OLDFIELD 'l'HOMAS.
'l'HE specimens e e ed o in he p esen pape we e mos ly
collec ed Ly � ess s. Cha les and E nes Hose in N. Celebes
an<l by M . A l cd E e e a he ex eme sou h o he same
island, and bo h a abou he same da e, Oc obe o
Decembe 1895. One o he chie objec s o hese na u alis s
was o ob ain o ou Na ional .Mnseum specimens o he
nume ous small mammals ha ha e been desc ibed o la e
yea s om he islaud o Celcbcs and ha e hi he o been
un ep esen ed in he B i ish Museum. This objec has
242 M . 0. Thomas on Mammals om
happily been e y success ully accomplished, and Celebes is
no, no longe he wo s - ep esen ed pa o he wo ld in he
i'lluseum collec ions, as was o me ly he case-, n imp o e
neu wholly due o hose who ha e al eady made No he n
Bo neo he bes - ep esen ed pa o he wo ld in he same
collec ions.
1n spi e o he many Celebean species p e iously desc ibed,
mos o which ha e now been ob ained, no le» han i e new
ones ! a e also been disco e ed-a esul lie mo e su p ising
as he Hoses we e wo king in wha was supposed o be he
bes -known pa o Celebes.
Besides he Hose and E e e specimens, a ba ob ained by
Go e no C eagh a San<lakan, N: Bo neo, is also desc ibed,
as also is a e y ema kable new ui -ba which occu s in
U . John Whi ehcad's las collec ion om he Philippines.
Bonda ne11adensis, sp. n.
Size o Boneia bide11s, Jen . Ea s o medium leng h,
s! a ply poin ed abo e, hei ou e edge wi h a small lobule
basally. Wings om he sides o he back as in ypical
Xan lw pyia. Colou abo e o head <la k smoky b own, o
nape dull yellow, o back <la k eddiRh b own, g adually
becoming pale and b igh e on he ump. Sides o neck
wi h da k golden gland- u oi. Chin, hma , and belly pale
g eyish b own. Vings and limb;; naked, excep jus on he
p oximal pa o he o ea m and on he emo a and neigh
bou ing pa s o he in e emo al memb ane.
Skull wi h a b oad hea y muzzle and s ong widely ex
panded zygoma a. P emaxill e sepa a ed in on .
Uppe inciso s wo in numbe , sho , cylind ical, equi
dis an om each o he and he canine:3. Canines long,
g oo ed in on . Fi s uppe p emola small, bu no
specially c ushed in be ween he cauine and secoud p e
mola . Lowe inciso s in an e en cu ed ow be ween he
canines, he ou e ou o i e imes as s ou as he inne ones,
Canines sho , s ou , and s ongly sla11 ed ou wa ds.
Dimensions o he ype (an adul male skin) :-
Fo ea m 97 millim. ( =3·8 inches); head and body (ap
p oxima e) 1511; ea (d ied) 20; lowe leg 53.
:;kull: ex eme leng h om nasion o occipu -!11·7; g ea es
b ead h 25; ip o nasals o sup ao bi al o amina 20; in e
o bi al b ead h 7·5; ip o ip o pos o bi al p ocesses 16;
on o uppe canine o back o � 15·1.
I-Jab. Menado, Celebes.
Coll. C. and E. Hose, Oc . 1895.
Celebes, Bo neo, and he Philippines. 243
This ba is p obably he N. Celebes ep esen a i e o he
sou he n B. bidens, om which i may be dis inguished by
he dis ance be ween he do sal o igins o he wings, i s
poin ed ea s, and di e en colou , no ably he b own ins ead
o " golden-yellow " c own.
HARPYI0�YCTERIS, gen. no .
Index wi h a claw. , ings om he sides o he hai y
back, inse ed behind a he junc ion o he i s and second
oes. No ail. Hin<l limbs appa en ly e y sho . In e
emo al memb ane obsole e, bu ied in hick u .
Den i ion.-!. 10�0, C. , P. ¾, M. ix 2=28 o 30.
Uppe inciso s la ge, ouching each o he and he canines,
shaped, when iewed in on , almos like hose o Desmodus,
each wi h a long oblique cusp ouching i s ellow in he
micldle line o he skull, bu in sec ion each is b oadly ian
gula , wi h a b oad pos e io basal ledge. Canines wi h a
la ge pos e io seconda y cusp abou hal as high as he main
cusp, and wi h a b oad pos e o-in e nal basal ledge, bu no
addi ional in e nal cusps; i s di ec ion much mo e slan ing
o wa ds han usual, as is he lowe canine also, so ha he
wo c oss each o he nea ly a igh angles, ins ead o being
app oxima ely pa allel. Fi s wo p emola s abou as in
<..:ynop e ls. :llola s oblong in sec ion and o a peculia
cuspida e cha ac e , he la e al longi udiual walls o he usual
median g oo e b oken up in o se e al minu e cusps, none o
which a c a all specially leng hened. Below, he inciso s
a e p ac ically obsole e, being minu e and almos c owded
ou * by he la ge canines, which ouch each o he in he
middle line, and each ha e an an e o-in e nal and a pos e o
ex e nal seconda y cusp an<l a b oad pos e io ledge.
l is di icul o say wi h ce ain y o wha p e iously
known genus his ema kable o m is mos nea ly allied. I s
peculia canines o a ce ain ex en ecall hose o llai-pyia,
bu his esemblance may be ei he acciden al o due o hei
common descen om he (p esumably) cuspida e- oo hed
ances o s o he P e opodid e . On he whole i may be
mos con enien ly placed nea Xan ha pyia and Boneia,
wi h which i sha es ce ain ex e nal cha ac e s, an indical
claw, and he cheek- oo h o mula o P. {, Jl. �; bu he
*In be single ype specimen one lowe inciso only is p esen , he
o he ha in" allen.
See P. Z. S. ll:i88, p. 473.
2-14 ; . 0. Thomas on Jlammals o111
unique inciso s, he sho bi- and icuspiuale canines, and he
mul icuspida e mola , sepa a e i widely e en om hese,
and ende i one o he mos isola ed o all he gene a o
he g oup.
l a pyionyc e is TVl1i eheadi, sp. n.
Size abou as in Xan ha pyia ample. :icauda a. Fu so ,
close and woolly, especially pos e io ly. Gene al colou o
l n m ;ill o e abo e and below a uni o m chocola e-b own,
a li le da ke on he ace, and a li le ligh e on he nape and
shoulde s. " Ying-memb anes da k, wi h a ew whi ish spo s
sca e ed abon hem. Ea s o medium leng h, ounded a
hei ips. Fu o he back ex ending hinly on o he o e
a ms and co e ing he hind limbs densely Jo1Yn o he oo s
o he claws. J n e emo nl memb ane ba ely a en h o an
inch wide, wholly bu ied in he u .
Dimensions o he _ype (an adul skin o doub ul sex) :
Fo ea m 84 uillim. (=3·3 inches); liead and body 140;
ea 17; index- inge and claw 60; hi d inge , me aca pal
59, i s phalanx 44, second phalanx 54.
Skull: basal leng h 37·5; g ea es b ead h 2:3•8; in e
o bi al L ead h, ip o ip o pos o bi al p oeesses 6·9. F on
o canine o back o m-2 17.
J lab. l lindo o, Philippines, al . 5000 ee .
Coll. J. Whi ehead, Decembe 1895.
l l . 'Whi ehead is o be cong a ula ed on his in e es in
a<ldi ion o he splendid disco e ies he has al eady made i�
he Philippine Islands.
Rliinolophus C,·eaglii, sp. n.
Size medium. Nose-lea widely di e en om ha o any
o he species by he ac ha while usually, in Dobson's
wo ds, he uppe ma gin o he connec ing p ocess o he
sella is conjoined wi h he summi o he e ical p ocess a
he same le el, o exceeds i pos e io ly in heigh , in he
p esen species he connec ing p ocess is almos obsole e and
is a o e opped by he an e io e ical p ocess. Abo e
he low connec ing p ocess, on he on ace o he sella,
he e is a p ominen u o b own hai s, p ojec ing o wa ds
and o e hanging he an e io e ical p ocess. The la e
p ocess is spa ula e, na owe below, b oadening ou in he
middle, poin ed abo e. . Iu e na ial lappe � ounded. Ho i
zon al nose-lea b oad, jus abou co e ing he muzzle.
Lowe lip wi h h ee g oo es. Ea s la ge, sha ply poin ed
Celebes, Bo neo, and he Philippi'nes. 245
a hei ips. ·wings om he ibi o nea ly hal an inch om
be ankle. Colou , as usual, dull b own abo e and below·.
Fi s uppe p emola be ween he well-sepa a ed canine
and second p emola . Second lowe p emola en i ely absen .
Dimensions o he ype (an adul emale in spi i ):
Fo ea m 49 millim. ( = l ·95 inch) ; head and body 51;
ail 17·5; ca om no ch 2:!; nose-lea 15 x 10; lowe
leg 23.
Hob. Sandakan, B i ish N. Bo neo.
P esen ed and eol oe ed by Go e no C. V. C eagh, C.U.G.,
in wl,ose liouou I ha e en u ed o name lie species.
This e y in e es ing ba di e s om all he o dina y
membe s o he genus bo h by he absence o he second
lowe p emola and by he cba ac e s o i s nose-lea , in
which la e espec i shows a ce ain endency owa ds he
unique and peculia Rh. ca;lophyllus, Pe e s. To he bes o
my belie , no species as ye desc ibed esembles Rh. C ea_qhi
closely enough o need de ailed compa ison wi h i .
Felis donies ica, L.
·wi h he a ious ema kable )!u ines sen by : [ . Yhi e
head om Luzon in 1895 he e was a ca ound li ing wild
a a conside able al i ude in he moun ains, As i was
possible l,a his was a domes ic ca un wild, i was pu
aside o u he e idence on he sul�jcc . Now, again,
}J . E e e sends om .i loun Bon hain, S. Celebes, a ca
ob ained a 6500 ee which p ecisely ag ees in e e y espec
wi h }J . Whi chead's Luzon animal. On compa ing hese
wo moun ain skins wi h wo specimens ob ained a :IIakassa
by 1 ' all ace, and conside ed e e since as F. domes ica, I ind
he e is jus he di e ence which migh be expec ed o occu
be ween ame and e al indi iduals. Indeed, one o T
allace's
is almos p ecisely like E,·e e 's, while he o he is ob iously
a domes ica ed specimen, and he compa ison o he whole
se ies shows conclusi ely ha he moun ain-ca , l,o h o
Luzon and Celebes mus be ega ded as e al indi iduals o
he o dina y ,'Jalay do,ues ic ca . The same conclusion is o
be d awn om he skulls, which ag ee e y closely wi h
hose o he domes ic ca , while qui e di e en om hose o
Felis clwus, o which, i a genuine wild species, he Luzon
and Celcbcs ca would by i s colou be mos nea ly a !ined.
Among he Mu id e ob ained by :lless s. C. and E. Hose
he ollowing known species occu , mos o hem ha ing been
[J e iou,;ly un ep esen ed in lie �I useum collec ion :-.llus
246 lll . 0. Thomas on .iJ anw,als om
Hellwaldi, Jen ., lll. Mu.,schenb oecki, Jen ., 11[. Meye i,
Jen ., .I . celebensis, G ay, !J[. xan hu u.,, G ay, 11[. calli
iclws, Jen ., ll . 11eglec 1 s, ,Ten . (?), JI. epl ippium, J cn .,
and C,·au o h,·i:c * leucu a, G ay. The skins o he las
named qui e ag ee wi h he ype o "Ecldo l, i:c leucu a," and
con i m D . J en ink's eco d o Celebcs as he ue locali y
o his ema kable a .
Jn addi ion o hese, wo new species o !J us we e ob ained,
whose desc ip ions ollow :-
!J us a o um, sp. n.
Size abou as in ,l u., a us. Fu o medium leng h, so ,
wi hou longe b is les in e mixed. Ea s la ge, e enly
ounded. Gene al colou abo e b ownish g ey, inely
sp inkled ' 'i h dull yellowish. Head a he pale , bu he
ci cum e ence o each eye sligh ly da ke . Unde su ace
di y g eyish yellow, he bases o he hai s sla y g ey, hei
ips dull yellowish; line o dema ca ion on sides li le
ma ked. Hind ee a he elonga e ; i h hind oe eaching
o he middle o he i s phalanx o he ou h; uppe su ace
o hands and ee sil e y whi e. Tail somewha sho e han
head and body, inely scaled (abou 11 scales o he cen i
me e), i s p oximal wo hi ds abo e b own, i s end and he
whole o i s unde su ace whi e.
Skull wi h a he a small c anial and long acial po ion.
Sup ao bi al beads dis inc , al hough no s ong. F on o
zygoma- no li le p ojec ed o wa ds. An e io pala al
o amina o medium leng h, no eaching back o he le el o
.!!::..1
, lllola s la ge and hea y. Bull o small.
Dimensions o he ype (an adul male in skin) :-
Head and body 193 nilli n.; ail 157; hin<l oo (mois
ened) 38·8; ea 20.
Skull : basal leng h 37·5; basila leng h o henselion 34·3;
g ea es b ead h 21 ·2 ; nasals U)·4 x 4·6; in e o bi al b ead h
6'4, in e pa ie al 5x 9·5; dias ema 12; pala e leng h om
benselion 19·2; pala al o amina 7·4 x 3· ; leng h o uppe
mola se ies 7·6; b ead h o pala e ou side �-' 9·2, inside
m.l 4•5,
Jlab. Ru ukan, Celebes, 3500 ee .
Coll. Cha les and E nes Hose.
Type killed Oc . 8, 1895.
* Nom. no .
Ech.io h i: , G ay, P. Z. S. 1867, p. 69(), nee
Echino h , Pe e s, MB. Akad. Be l. 1853, p. 487 (Echinode ma a).
Celebes, Bo neo, and he Philippines. 247
Besides he ype, he e a e se e al o he specimens om
Ru uka11, wo om l Ienado, and one om l loun i Iasa ang.
This species is appa en ly mcs closely allied o .JI. ch yso
comus, Ilol m., also a na i e o Celebes, bu di e s om i by
i s la ge size, beaded sup ao bi al edges, and much hea ie
mola s.
1llus minahass o, sp. n.
Gene al appea ance ha o he Mus 1lla ga1·eUa; o Sa awak,
bu he s nc u e o he ee qui e as in no mal Mus. Size
small. Fu long- and so . Ea s o medium leng h. Gene al
colou abo e u ous, dulle on he back, b igh e on he sides.
Unde su ace whi e, bases o he hai s g ey ; line o dema
ca ion on sides well-ma ked. : Ie apodials b own, inge s
and oes whi ish. Thumb wi h a la ge nail, o he digi s
wi h sho , sha p, cu ed claws. Hallux no opposable, wi h
a claw, eaching wi hou i s claw o he base o he second
oe; i h oe eaching o he middle o he second phalanx
o he ou h. Soles naked; pads appa en ly six in numbe
as usual, no s ia ed, he pos e io one elonga ed. Tail long,
slende , e y inely scaled (23-25 scales o he cen ime e),
sho -hai ed ( he ex eme ip is gone, bu he e is no sign o
he commencemen o a e minal pencil), uni o mly b nwn
abo e and below.
::lkull sugges ing ha o Chi opodomys b_y i s disp opo
ionally la ge b ain-case and small ace. l luzzlc sho and
slende . In e o bi al egion na ow, la abo e, i s edges
squa e, ain ly beaded; pa ie als la ge, con ex; in e pa ie al
la ge; zygoma a e y weak and slende , g ea es b ead h o
skull jus a hei pos e io oo s; an e io zygoma- oo no
p ojec ed o wa ds. Pala al o amina small. Mola s sho ,
b oad and onuded.
Dimensions o be ype (an adul emale in skin):-
Head and body 77 millim.; ail (ex eme ip gone) 105 ;
hind oo (mois ened) 20; heel o on o las oo -pad 9·3;
ea (d ied) 1:M.
Skull: basal leng h 18·6; basila leng h om henselion
17·2; g ea es b ead l1 12·3; nasals 7·1 x 2·5; in e o bi al
b ead h 3·8; b ain-case, leng h 13·2, b ead h U ·8; in e
pa ie al 4,5 x 8·5; dias ema 6·1; an e io pala al o amina 2·9;
leng h o uppe mola se ies 3·2.
llab. Runikan, 11Iinahassa, N. Celebes.
Coll. Cha les and E nes Hose, No embe 1895.
This e y p e y li le mouse is eadily dis inguishable
om all o he s by i s Chi opodomys-like appea ance and
skull, combined wi h i s s ic ly .Mus-like hands and ee .
