MAMMALS COLLECTED BY DR. W. L. ABBOTT ON BORNEO
AND SOME OF THE SMALL ADJACENT ISLANDS.
By Ma cus Wa d Lyon, J .,
Assis an Cu a o , Di ision o Mammals, U. 8. Na ional Museum.
INTRODUCTION.
Be o e collec ing on he mainland o Bo neo, D . W. L. Abbo
made wo expedi ions o adjacen islands, he mammalian auna o
which is closely allied o ha o Bo neo. One o hese expedi ions
was o he Na una Islands, no h o Bo neo, and was made du ing he
sp ing and summe o 1900; he o he was o he Ka ima a Islands,
o he wes coas , du ing Augus and Sep embe , 1904. Lis s o
he mammals ob ained on hese expedi ions we e published by M .
Ge i S. Mille , j ., in 1901 and in 1906. x Mo e ecen ly Doc o
Abbo has isi ed he mainland o Bo neo i e imes, on each
occasion s opping a some o he adjacen islands. An accoun o
he mammals collec ed on he i s ip o he mainland, co e ing
he Kapuas Ri e egion, was published by me in 1907.2 The p esen
pape aims o gi e an accoun o he mammals ob ained on he
emaining ou expedi ions — wo o sou hwes e n Bo neo and wo
o sou heas e n Bo neo — as well as hose collec ed in he nea -by
islands. Fo he sake o comple eness, I ha e included in he
p esen pape he species collec ed in he Kapuas Ri e egion, as
well as hose om Ka ima a Islands, he la e ha ing been isi ed
a second ime. Fou sho pape s 3 ha e appea ed ecen ly which
i P oc. Wash. Acad. Sci., ol. 3, pp. 111-138, Ma ch 26, 1901. P oe. U. S. Na . Mus., ol. 31, pp. 55-66,
July 23, 190d.
2 P oc. U. S. Na . Mus., ol. 33, pp. 547-572, Decembe 24, 1907.
3 Lyon, Desc ip ion o a new squi el o he Sciu us p e os ii g oup om Pulo Temaju, wes coas o
Bo neo. Smi hs. Misc. Coll., ol. 4S, pp. 275-270, Feb ua y 4, 1907.
Lyon, Rema ks on he Insec o o es o he genus Gymnu a. P oc. U. S. Na . Mus., ol. 36, p. 449,
May 27, 1909. Doc o Abbo 's Bo nean specimens lis ed.
Ellio , On he genus P esbylis Esch., and Le Ta sie Bu ion, wi h desc ip ions o wo new species o
Ta sius. Bull. Ame . Mus. Na . His ., No. 28, pp. 151-154, May 27, 1910. Desc ibes he a sie om
he Kapuas Ri e egion.
Ellio , Desc ip ions o some new species o monkeys o he gene a Pi hecus and Pyga h ii collec ed by
D . W. L. Abbo and p esen ed o he Uni ed S a es Na ional Museum. P oc. U. S. Na . Mus., ol.
38, pp. 313-352, Augus 6, 1910. Desc ibes he common macaques om Ka ima a, he Kapuas Ri e
egion, Bawean Island, and Pulo Mala Si i.
P oceedings U. S. Na ional Museum, Vol. 40— No. 1809.
54 PROCEEDINGS OF THE NATIONAL MUSEUM. ol.40.
deal in pa wi h he mammals included in he p esen pape , bu
he collec ions as a whole a e he e published o he i s ime.
A lis o he locali ies isi ed by Doc o Abbo is gi en below,
oge he wi h his geog aphical and na u al his o y ield-no es. Many
o hese locali ies a e no o be ound on he o dina y maps, and
e e ence should be made o he maps on pages 55 and 57.
LIST OF LOCALITIES, WITH GEOGRAPHICAL AND NATURAL HISTORY
FIELD-NOTES BY DR. W. L. ABBOTT.
Wes Bo nean mainland, June 6-Sep embe 16, 1907.
Sukadana (o SuJcudana). — A Sukudana a mass o hills 1,000 o 2,000 ee high ise
igh om he seasho e. They a e well o es ed, bu he e a e many plan a ions o
du ians and o he ui upon hei slopes and many clea ings in he neighbo hood.
The sea is e y shallow nea he coas ; o he wise i would ha e been a good collec ing
g ound o me,1 as animal li e is abundan in he neighbo hood. The inhabi an s
a e all Malays excep a ew Chinese ade s and small plan e s.
Sempang Ri e . — The coun y along he Sempang Ri e is low and swampy, e y
li le abo e high wa e , as is also nea ly all wes Bo neo nea he coas . The lowe
pa o he p incipal a luen , he Semandang, is also low and la . The headwa e s
o bo h i e s a e among he hills. The low coun y is o he mos pa hea y o es ,
wi h a s ip o clea ing along he i e banks. The hilly coun y is inhabi ed by
Dyaks and he e he e is compa a i ely li le i gin o es emaining. The g ea e
pa o he su ace is co e ed wi h seconda y jungle. A ew hinoce oses a e said o
inhabi he uppe Sempang abou Ba u Dayeu (o Dajeuh). A much la ge ca
han Felis nebulosa occu s, called by Malays and Dyaks " imow." I mus be e y
a e, as e y ew people whom I me had e e seen i . The big ed pig (Sus ga gan ua )
I hea d o om bo h Dyaks and Malays, especially abou he uppe Sempang Ri e ,
bu no epo ed a all common.
Ma an (o Ma ai) Ri e . — The Sungei Ma an en e s he Sempang a he ajah's
kampong ( illage) called Sempang. I camped abou 12 miles up (A hou s) a a place
called Ma an. Fo me ly he e was a la ge kampong he e bu i was abandoned
wen y o hi y yea s since. Mos o he neighbo hood is la ge seconda y jungle.
The hill called Ma an was close by. Animal li e was e y abundan , and o ang-u ans
plen i ul. The e a e no pe manen illages now upon he Sungei Ma an, excep
on he slopes o he hill called Sepunchok, abou 2 hou s abo e Sempang, whe e
he e a e eigh amilies.
Moun Palung. — Palung is 1,110 me e s in heigh . I ascended Pan i and emained
one nigh ; saw no mammals high up. Pan i [no shown on map] is one o he hills o
Palung. I is somewhe e abou 3,500 ee . Rhinoce oses a e said o inhabi he
lowlands abou he base o Palung. No api o ban ing (Bos) in his pa o Bo neo.
Sou hwes Bo nean mainland, June 17-Sep embe 29, 1908.
Kendawangan Ri e . — The coun y along he lowe Kendawangan Ri e , as else-
whe e in wes Bo neo, is mos ly swamp, jus abo e high-wa e ma k. Occasional
highe pa ches and ac s occu which always emain d y and a e called "pema ong,"
abou wha we call "islands" in swamps o "hummocks" in Flo ida. The e a e no
hills close o he lowe i e excep a Mankol. A Lanchu , 30 miles om he mou h,
he e is qui e a la ge ac o d y land ex ending back o Moun Kedio, 4 miles om
he i e . Abou Kalang Anye , a Malay kampong o h ee houses, 70 miles om he
mou h, he banks become highe , wi h bu li le swamp. A ew miles highe up he
hilly and olling coun y is eached, he Dyak coun y, whe e mos o he o iginal
jungle has been des oyed and now co e ed wi h seconda y jungle, sc ub, and "alang
i Doc o Abbo a els abou in a schoone , and he ancho age p obably was poo .
no. 1809. MAMMALS FROM BORNEO AND VICINITY— LYON. 55
alang" (long coa se g ass). On he uppe pa o he la ges ibu a y, he Mambuluh,
he e a e many "danau, " o lakes, which, howe e , d y up in ime o d ough . The e
a e se e al hund ed Malays li ing along he lowe Kendawangan, Lobo Ba il, 18 miles
Fig. 1.— Map o Sou hwes e n Bo neo.
om he mou h, being he highes pe manen ly inhabi ed place. The a an and
gu a ga he e s wande all o e he o es s and he uninhabi ed bel lying be ween he
Malay se lemen s and he Dyak coun y on he hilly g ound, and hey all ca y guns
56 PROCEEDINGS OF THE NATIONAL MUSEUM. ol.40.
wi h hem. On he uppe Kendawangan among he Dyaks he e a e abou 200 Malays,
mos ly abou Ma o. Nea ly e e y man, bo h Dyak and Malay, has a gun; so game is
ema kably sca ce. Luckily, powde is di icul o ob ain. The Du ch ha e no
in e e ed wi h he na i es much as ye , and ha e no aken away hei i ea ms, as
hey a e g adually doing all o e hei colonies, much o he ad an age o he animal
li e, o Dyaks de ou e e y hing hey can kill. I mysel saw e y much less li e
han on he Sempang, and a less han in sou heas Bo neo, whe e he na i es we e all
disa med se e al yea s ago. The Dyaks kill a mo e wi h hei "be an ik, " o spea
aps, han by shoo ing, excep in he case o o ang-u an. The Dyaks a e ex emely
ond o he mea o hese, and i is useless o look o o angs anywhe e in he neighbo -
hood o a Dyak se lemen . I could hea no hing o Sus ga gan ua, and p obably i
does no occu in ha pa o Bo neo, no could I hea any hing o he " imau, " o
ige , which was said o inhabi he Sempang dis ic . A ew hinoce oses a e said o
inhabi he neighbo hood o Moun Kedio and a ew sapi u an (Bos sondaicus) on i s
wes slopes on he headwa e s o he li le i e Tinga . This is he only place on o
nea he wes coas o Bo neo whe e wild ca le occu . The Malays old me hey also
exis ed in some places on he uppe Pasaguan Ri e — ha is he nex la ge i e no h
o he Kendawangan. The coun y abou Moun Kedio is uninhabi ed, and he e is
said o be much animal li e he e. Fi y o six y yea s ago he dis ic was inhabi ed
by Dyaks, bu hese, becoming in ol ed in wa wi h he Malay ajah, le hei homes
and led in o he in e io . All o e his co ne o Bo neo occu sligh ly ele a ed
sandy ac s co e ed wi h small ees (o smalle ees han he su ounding o es );
hese a e locally called ' ' padang, ' ' which is Malay o meadow. Much o his land and
a good deal o he d ie o es was bu ned o e six o se en yea s ago, du ing a pe iod o
excessi e d ough . No ain ell o ou o i e mon hs, a mos unusual occu ence in
any pa o Bo neo, whe e o dina ily ain alls e e y 'mon h in he yea , and he e is
no p ope d y season. The ollowing animals, in addi ion o some al eady men ioned,
we e well known o he na i es, bu none was ob ained du ing his ip: Mydaus,
called bobo by Malay and kalinsida by Dyaks; Rei h osclu us; Fells nebulosa,
imau dahan; Fells ma mo a a? , a wild ca wi h a la ge ail, was caugh by a Malay
in a sna e, bu o some eason was no b ough o me; Pa adoxu us philippinensis;
Gymnu a, local name angkis.
Ba u Ju ong.— Ba u Ju ong is he sou he ly poin o a ange o hills which s e ch
no h-no heas o Mankol on he Kendawangan Ri e . I ancho ed in a small s ai
be ween Pulo I as and he mainland. Two o h ee Malay amilies li ed he e and had
hei clea ings. Pigs we e plen i ul; kijang (mun jacs) and usa we e also common,
bu he Malays we e con inually a e hem wi h hei dogs. Sou h o his he e a e no
inhabi an s in he sou hwes co ne o Bo neo, and un il ecen ly none on he sou h
coas wes o he Sungei Jelai. La ely, howe e , abou a dozen amilies o Malays
ha e made clea ings on he lowe cou se o he Sungei Aye Hi am Besa . Animals
we e said o be plen i ul, especially usa, abou Tanjong Kepala, whe e he e is said
o be la ge ac s o sho g ass. I saw wo o angs and he sa ongs (nes s) o many mo e
abou 2 miles eas o Ba u Ju ong, whe e I sho one emale.
Mankol. — Mankol lies along he Kendawangan Ri e o abou 2 miles. The e
a e abou 25 o mo e houses al oge he . A his poin he no h end o he ange o
hills (400 o 800 ee high) app oaches he i e . Animals we e sca ce, as mos o he
people had guns, and he jungle in he hills and along he base was ull o old je a s and
paga s ( aps and sna es). A ew o angs we e said o occu , bu I saw no aces.
Lanchu . — Lanchu is now wi hou inhabi an s. I was o me ly he mos impo -
an illage on he i e and he esidence o he ajah. The g ound along he i e
is qui e high and d y and co e ed wi h seconda y jungle and long g ass o some dis-
ance back om he i e . Fou o i e miles due eas ises Moun Kedio, in an
uninhabi ed dis ic co e ed wi h hea y o es .
no. 1809. MAMMALS FROM BORNEO AND VICINITY— LYON. 57
Klumpang Bay, sou heas e n Bo neo, Janua y 8-Ma ch 13, 1908, and Ap il
18-19, 1909,
Tanjong Ba u lies a he no h en ance o Klumpang Bay. The hill is 3 o 4
miles long and 1,200 ee a i s highes poin . I is co e ed wi h hea y o es ,
Fig. 2.— Map o Sou heas e n Bo neo.
excep a he sou h end, whe e he e a e some peppe ga dens and a good many old
clea ings co e ed wi h sc ub and coa se g ass. The imbe is e y good, con aining
much bilian (i on wood). A ange o lowe hills ex ends no h o his o Buki Ba u
nea he Sampanahan Ri e and Pamukang Bay. This ange o hills is e iden ly an
58 PROCEEDINGS OF THE NATIONAL MUSEUM. ol.40.
ancien island, joined o he mainland by he ele a ion o he land and also by sil ing
up a wide ac o mang o e swamp lying in he in e ening space. As one ascends
he i e , a ew miles om he bay, e idences o ele a ion become e e ywhe e isible
in he masses o limes one co al p ojec ing om he swampy su ace. In many cases
he la su ace o he ancien ee is bu a ew inches below he mud o ac ually on he
su ace. A li le a he an i egula line o limes one hills, mos ly wi h p ecipi ous
and wa e wo n sides, uns in a gene al di ec ion pa allel wi h he coas . The
unclea ed land is all hea y o es . Many old clea ings and some new exis in he
neighbo hood o he i e s. The popula ion o he coas is made up o Bugis, Banje ,
and o he Mohammedan Malay ibes. Inland is a la ge Dyak (pagan) popula ion.
The Du ch annexed his dis ic in 1905.
Pangkallahan (o Bangkallaan) Ri e . — Kampong Pangkallahan is abou 7 miles
up he i e o he same name and is he esidence o he mankoh, o head o he Dyaks
o he dis ic . The coun y is co e ed wi h splendid o es , wi h only a ew clea -
ings. The e a e anges and sca e ed hills o p ecipi ous limes one ock. Animals
seemed sca ce, as usual in Dyak dis ic s, e e y hing hun ed o o d i en away by'
he Dyaks and hei dogs. They ha e e y ew guns and no powde . Abou a hal
mile abo e he illage is he lowe en ance o he Temmelung o unnel o he Pang-
kallahan. This unnel ca e, h ough which he i e lows, swa ms wi h ba s as well as
wi h edible bi ds'-nes swi s, hund eds o housands o hei nes s being collec ed
annually. I could no de ise any way o ca ching he ba s, as I had no ne and had
no b ough a gun o ea o a capsize, he e being a dange ous apid nea he lowe
end o he unnel. The Temmelung mus be abou 2 miles long, judging by he ime
we ook passing h ough i . I is h ough limes one ock. I do no hink he oo
was anywhe e e y hick, a some places only 2 o 3 ya ds. Tha pa o he coun y
is ull o limes one hills and ocks all honeycombed wi h ca es and passages. Bu
ba s did no seem o be p esen in he hund eds o holes I isi ed, excep an occasional
indi idual which I could no secu e. Judging om he noise, o he ligh om he
o ches did li le bu accen ua e he gloom, he e mus be hund eds o housands in
he Temmelung. Ano he unnel occu s a he up he same i e , bu I did no
isi i , as i was said o be e y di icul o passage excep by a e y small canoe.
Sa a ok Ri e . — The Sa a ok is a small i e lowing in o Klumpang Bay. Two
Dyak houses s ood in a small new clea ing. Back o his and ex ending eas wa d o
nea ly 2 miles was a la ge ac o alang alang ( all, coa se g ass) co e ing se e al hun-
d ed ac es. The ; appea ed o be a good many usa he e, bu he g ass, 4 o 5 ee
high, made i e y di icul o shoo any hing. When he g ass is bu ned o in he d y
season he place is said o be equen ed by a he d o sapi u an (Bos sondaicus). The e
we e no acks o hese a he ime o my isi . Mos o my collec ing was done nea
some limes one hills and ocks abou a mile wes wa d in he mids o splendid o es .
Only he ed P esby is was seen he e; nea he coas only he black one exis ed.
Besides he animals sho in Klumpang Bay I saw he long- ailed and he pig- ailed
macaques and P esby is c is a a.
Pamukang o Tjengal Bay, Ma ch 17-Ap il 7, 1908, and Ma ch 23-Ap il 13, 1909.
Musangs we e gene ally sca ce excep he angalunga, which was common. I
le mos o hose go which I caugh in aps. The Mydaus was well known, bu no
specimens we e ob ained. Gymnu a mus be e y common, as one o en smel hem
in he jungle, al hough none we e ob ained. The pig- ailed macaque was wice me
wi h in d o es, bu none we e secu ed. I me wi h Rei h osciu us wice on Buki
Ba u. I was unning on he g ound in hea y o es . I s mo emen s we e so quick
I could no shoo i . I s big bushy ail, ca ied high o e i s back, made he animal
e y conspicuous. I was much su p ised o ind ha i is a g ound squi el. The
headman o he Bajaus a Sungei Manungul, Pamukang Bay, said he e used o be
some hinos in ha locali y, bu he had seen no aces o yea s.
Pasi Ri e , Decembe 31, 1908-Janua y 22, 1909.
no. 1809. MAMMALS FROM BORNEO AND VICINITY— LYON. 59
Balik Papan Bay, Feb ua y 1-Feb ua y 24, 1909. Balik Papan Bay ex ends
nea ly 18 miles in a no he ly di ec ion. Se e al i e s emp y in o i . The eas e n
sho e is high and hilly o he mos pa , he wes e n low and co e ed wi h mang o es.
The e we e e y ew inhabi an s when he Royal Du ch Oil Company i s es ab-
lished i s headqua e s he e abou i een yea s ago. Now he e is a la ge se lemen ,
wha es, oil e ine ies, pa a in wo ks, e c., and abou 6,000 inhabi an s, si ua ed on
he eas e n en ance o he bay. I is an excellen ha bo . The sho es o he bay
a e s ill mos ly hea y o es . The line o he bay p obably coincides wi h ha o a
aul . The hills a e mos ly ed la e i e, bu a line o limes one s e ches sou hwes -
wa d no a om he head o he bay. Animal li e is ai ly plen i ul. Ban ing (Bos
sondaicus) we e said o occu especially a ound he head o he bay.
Pulo Lamuko an, May 7-10, 1907. Pulo Lamuko an is he la ges o he Bu ong
Islands, and lies abou 10 miles o he mainland o Bo neo. I is abou 4 miles long
by 1 mile wide. I consis s o a long idge wi h a low ock nea he middle o he
island. The highes poin is nea he sou he n end and is nea ly 1,000 ee in heigh .
Mos o he su ace is now clea ed and he lowe pa s en i ely plan ed wi h coconu s,
which a e e y ine and heal hy. The only hea y o es emaining co e s he highes
summi and he c es o he idge on he sou he n pa o he island, and a smalle
piece on he summi o he no he n idge. The soil seems e ile and he su ace is
no e y ocky, excep upon he sho e. The o es ees a e e y la ge in he emain-
ing jungle. The o he islands in he g oup, ou in numbe , a e smalle and ha e
been en i ely clea ed and plan ed wi h coconu s. The inhabi an s a e Malays
om Sambas. A Sciu as i a us, a Macacus cynomolgus, a la ge pig, and h ee o
mo e a s cons i u e he mammalian auna. Pigs a e no now e y nume ous, ha ing
been la gely hun ed o by Chinese om he mainland, who employ dogs. Pigs we e
o me ly e y abundan upon he o he islands o he g oup (Pena a and Kebun),
bu ha e been en i ely ex e mina ed upon hese since he clea ing o he jungle.
Ra s seemed p e y common. My aps we e se in he jungle on he idge nea he
highes peak. F ui pigeons and Nicoba pigeons we e ai ly common.
Pulo Temaju, May 5-6, 1907. (Fi s isi ed by Doc o Abbo in 1905.) Pulo
Temaju is now mos ly clea ed and plan ed wi h coconu . The only hea y o es
emaining co e s he summi and uppe slopes o he highes peak. The e a e no
mammals excep Sciu us p ose pinx and a s. The coconu ees a e no e y p o-
duc i e and seem much diseased.
Pulo Da u, May 2-4, 1907. Pulo Da u lies 21 miles om he coas o Bo neo. I
is abou 1| miles long by h ee- ou hs mile wide. I is e y hilly (moun ainous)
and ocky, and ises o abou 1,000 ee . I is co e ed wi h hea y o es , excep
a some places on he eas side, whe e some clea ings ha e been made and a ew
coconu s plan ed. The clea ings a e now o e g own wi h la ge seconda y jungle,
as he a s swa m o such an ex en i is impossible o g ow any hing. Squi els
(Sciu us i a us g oup) we e plen i ul. No monkeys. I pu ou abou o y a
aps one nigh . Nea ly e e y one was sp ung in he mo ning, bu many o he a s
had been de ou ed by land o he mi c abs. Whi e ui pigeons and Nicoba pigeons
we e common. The e a e no sand beaches, bu plen y o esh wa e lows ou benea h
he ocks, on he eas side a any a e, be ween high and low wa e ma ks. Mus
" a us" appea ed o be mo e abundan hen M. "lingensis," bu he he mi c abs
showed ma ked p e e ence o he la e and spoiled mos o hose caugh .
Pulo Panebangan, May 16-26, June 2-3, and Sep embe 20-21, 1907. Pulo
Panebangan lies 8 miles om Pulo Maja, which is p ac ically he mainland o Bo neo.
I is abou 4 miles long by 2 o 3 wide, con aining abou 6,000 o 7,000 ac es, I is
e y hilly, sca cely any le el g ound. The highes poin is said o be 1,700 ee .
I is uninhabi ed and co e ed wi h dense jungle. Ra ans (o an almos alueless
a ie y) abound, and he jungle is so ma ed up wi h a ans and hei lagella as
o be impene able. In some places he jungle seems o ha e been clea ed a some
60 PROCEEDINGS OF THE NATIONAL MUSEUM. ol.40.
p e ious pe iod, as a he head o he small bay on he no h side. Macacus cyno nolgus
was he only monkey seen. The e is no lo ong (P esby ia). The a u a is p obably
ai ly common, as I equen ly hea d i . I only ac ually saw he h ee which we e
sho . I hough I saw and hea d Nannosciu us on one occasion, bu am no ce ain.
The e is no napu o la ge agulus. No dee occu s; no o e acks seen; and he
angalunga apped was he only musang seen. The small Hipposide os was e y
common, lying abou in he day ime; he la ge one was less common. The e we e
se e al o he species o ba s lying abou in he o es in dayligh , bu I ailed o
secu e any. No Rhinolophus we e seen. Ano he musang is said o occu and also
a ed lying squi el. Pigs we e plen i ul. All hose no iced upon he beach seemed
o be males. We did no once see a small one ou o wo o h ee dozen indi iduals.
Possibly he old males app op ia e he beaches and ee s, which a e he bes eeding
g ounds, and d i e away in ude s. The pigs we e all e y lean and gaun . Up o
he p esen he e a e no inhabi an s, al hough he e a e many du ians, mangos, and
chempadak. We hea d ha some Ka ima a people p opose o come in July, 1907,
and o m plan a ions.
Pulo Pelapis, May 29-June 1, 1907. Pelapis, a g oup o ou islands, lies 3^ miles
sou hwes o Panebangan and 17 miles om Ka ima a. The islands a e all hilly and
ocky and co e ed wi h o es . The highes poin is abou 1,200 ee . The o al a ea
is abou 5,000 ac es. Sou h Island (also called Pelapis Tengah) is he la ges and is
nea ly 2| miles long. All collec ing was done upon his island. Some yea s ago some
Malays and Ka ima a people se led upon Sou h Island and a med some plan a ions
o coconu s, e c., bu h ee yea s ago an epidemic b oke ou , many died and he es
o he se le s led, and o-day sca ce a ace o human occupancy can be seen. Besides
he animals ob ained, he lying lemu and he common long- ailed macaque occu .
No agulus, no musangs, o no a u as occu . The pigs come ou upon he sand
beaches and ee s a low ide, and can be sho wi hou much di icul y. Ba s we e
lying abou in he o es on Pelapis, bu no nea ly so many as upon Panebangan.
None we e ob ained.
Ka ima a Islands, Telok Eda , Ka ima a Island, Oc obe 4-7, 1908. (Fi s isi ed
by D . W. L. Abbo in 1905.)
Pulo Juan a, Sep embe 10-11, 1907. Pulo Juan a is a small island, 6^ miles
om he mainland, abou 1 kilome e long by wide. I is abou 300 ee high.
The g ea e pa is co e ed wi h hea y o es , bu a clea ing was made abou ou
yea s ago and an a emp made o plan coconu s, bu no hing could be g own owing
o he a s and -pigs. Abou hi y pigs we e killed, bu many emained. The e a e
s ill a ew on he island, bu I could nei he shoo no ap hem. The e a e no squi -
els on he island.
Pulo Bauwal, June 12-16, 1908. Pulo Bauwal (o Rendez ous Island) lies nea
he sou hwes co ne o Bo neo. I is abou 15 miles om Tanjong Samba , which
is he ex eme sou hwes poin . Bauwal is abou 6 geog aphic miles long by 5 in
ex eme wid h and con ains abou 12,000 ac es. The s ai sepa a ing i om he
mainland is om 4 o 6 miles wide and 4 o 12 a homs deep. The island is su -
ounded by wide co al ee s and ha d ocks. The island is composed o ha d ed
ock (i on o e?) and is a he low. The e a e wo o h ee hills, ising o abou 300
ee o he ops o he ees. The su ace is olling and co e ed wi h o es . The e
is some mang o e swamp. A house, inhabi ed by Pon ianak Malays, is on he eas
coas , and i e houses inhabi ed by O ang Lau om Ka ima a we e es ablished nea
he no hwes co ne abou h ee yea s ago. The auna is a he peculia in ha he e
a e no squi els, aguli, usa, o pigs. Mun jacs and angalungas a e common.
Solombo (no shown on map), Decembe 4-G, 1907. The island o Solombo, o
Masolombo Besa , is abou 4 miles long by abou 2 wide. The su ace is mos ly a he
low and olling, and he e is one hill abou 250 ee high. The ock seems o be nea ly
all olcanic, excep , o cou se, he co al a ound he sho e. The island is su ounded
no. 1809. MAMMALS FROM BORNEO AND VICINITY— LYON. 61
by a co al ee . I lies midway be ween Madu a and Bo neo, 84 miles o each. The e
is qui e a la ge popula ion, 300 o 400 o Bugis and Madu ese. The island has been
se led abou o y yea s. Mos o he hea y o es has been clea ed, excep abou he
hill. The soil, da k ed, wi h many s ones and ocks o la a upon he su ace, is e y
e ile, and p oduces la ge c ops o paddy and maize. The only mammals I saw we e
lying oxes, which we e qui e common. Ra s we e said o be plen i ul. The e a e
no monkeys o squi els. The e a e said o be many sapi o e al ca le (Bos sondaicus),
and he na i es we e e y anxious o me o go and shoo some, bu I did no ha e
ime o do so. They a e said o ha e been unning wild a long ime. Bi ds a e e y
plen i ul. The anxie y as o he sa e y o my schoone p e en ed me om doing as
much as I would ha e liked, besides cu ing sho my s ay, o he wind wen a ound
o he no hwes and we had o ge ou on sho no ice. I should like o ha e pu in
se e al mo e days and isi ed Pulo Solombo Ki chil. The e a e no people upon he
la e and i is s ill unclea ed o es . Bi ds a e said o be e y plen i ul he e.
Bawean Island, No embe 24-27, 1907. (No shown on map.) Bawean Island
lies abou 60 miles no h o he S ai s o Madu a. I is abou 11 miles long by 10
wide, he a ea being abou 100 squa e miles. The su ace is moun ainous, se e al o
he hills ising om 2,000 o 2,200 ee . The island is olcanic, he ocks being mos ly
la a and basal , wi h some limes one. The e a e ex ensi e co al ee s a ound he
coas . Many olcanic cones a e sca e ed abou and he e a e se e al ho sp ings.
A beau i ul lake o abou 15 ac es called he Telaga occupies he ex inc c a e o he
moun ain o he same name. The island is densely inhabi ed a ound he coas s,
he popula ion being abou 50,000. Mos o he men go o Ja a, he S ai s, and
Suma a is sea ch o wo k. Nea ly all he saises in Singapo e and Penang a e
Beyanese. One sees bu ew males be ween he ages o 18 and 40 in Bawean. The
women wea e he ma s o pandanus, o which he island is amous, and which a e
expo ed all o e he a chipelago. Animal li e is no plen i ul. Rema kably ew
bi ds a e o be seen. As o mammals, pigs a e e y common. The only ones ob ained,
howe e , we e young — oo small o be o any use as specimens. The usa (Ce us
kuhli)mis no nume ous, and is only ound in a ew locali ies. The only specimens
ob ained we e h ee pai s o ho ns om he kampong o Tombak on he no h side
o he island. I is mo e nume ous on he hills behind Tombak han a any o he
place. The e is also a po cupine, possibly wo species, a musang, a Mam's, and wha
appea s o be an o e , bu hey do no seem o be common, and none we e ob ained.
P e opus was e y common, bu no o he ba s we e seen. The ca le a e he ame
a ie y o Bos sondaicus, which has been in oduced om Bali. They a e no used
o milk, bu a e employed o some ex en o d a and plowing. A ew bu aloes
a e also kep . The e is bu li le i gin o es le upon Bawean. The la ges piece
lies upon he no he n slopes o he cen al moun ain mass, especially Gunong Besa
and he Telaga. The e is also a ac upon he wes coas which I did no isi ,
howe e , and he e a e some small pa ches on he eas e n and sou he n slopes o he
moun ains. The es o he island is o a g ea ex en co e ed wi h small sc ub and
gian bamboos. Mangos and jack ui escaped om cul i a ion o m much o he
jungle. The sc ub is e e ywhe e a e sed by he ails o med by he nume ous
ca le.
A ends (o Ke amian") Island, No embe 24, 1908. (See map, p. 57.)
Pulo Ma a Si i, Decembe 7-11, 1907, and No embe 25-Decembe 1, 1908.
Ma a Si i is he la ges o he Lau o o Lau Ki chil Islands. I is 7£ miles long by
1J b oad, is 1,400 ee high. The e is sca cely any le el g ound, he whole island
being hilly, consis ing o a long idge di iding in o wo peninsulas a i s no heas
end, inclosing he bay o Telok Sungei. I is a good, sa e ha bo , and we lay he e in
secu i y, al hough he wes e ly monsoon was blowing s ong a he ime. The o he
wo islands o he g oup, Kalambau and Kadapangan, a e each abou one-hal he
eize o Ma a Si i, and each consis s o one long idge. The islands a e o g ani e
68 PROCEEDINGS OF THE NATIONAL MUSEUM. ol. 40.
has he long maxilla y oo h- ow o T. ca ima se. Six een specimens
om Ka ima a in 1904.
' Measu emen s o he adul , Ca . No. 145352, male : Head and body,
475 mm.; ail, 78; hind oo , including hoo s, 125; weigh , 2.2 kilos;
condylo-basal leng h o skull, 90.2 ; zygoma ic wid h, 44.1 ; maxilla y
oo h- ow, al eoli, 35.
Measu emen s o adul kanchils.
1 Collec o 's measu emen s.
2 Measu ed by w i e .
3 Collec o 's measu emen s in pounds and qua e s compu ed o kilog ams.
4 Measu emen s only app oxima e.
6 Type o T agulus i gicollis.
6 Skulls only.
no. 1809. MAMMALS FROM BORNEO AND VICINITY— LYON. 69
RUSA BROOKEI (Hose).
1893. Ce us b ookei Hose, Ann. Mag. Na . His ., se . 6, ol. 12, p. 206.
1906. Rusa b ookei, Lyon, P oc. U. S. Na . Mus., ol. 31, p. 585, Decembe 18,
1906.
1907. Rusa b ookei, Lyon, P oc. U. S. Na . Mus., ol. 33, p. 550, Decembe 24,
1907.
Twen y- h ee indi iduals o Rusa h ookei a e ep esen ed in Doc o
Abbo 's ecen collec ions om sou hwes e n and sou heas e n Bo -
neo, six een o hem by an le s ob ained om na i es, and ou by
skins and skulls, and he emaining h ee by skulls and scalps. Two
specimens we e ob ained along he Kapuas Ri e in 1905. (Fo
exac locali ies and he ex e nal, c anial, and an le measu emen s
see ables, p. 70.)
I is possible ha mo e han one species is ep esen ed by his
ma e ial. The skulls o he male specimens om sou heas e n Bo -
neo a e age somewha smalle han he skull o a ully adul male
om he Sempang Ri e . The skins o hose om sou heas e n
Bo neo a e decidedly di e en om ha ob ained in sou hwes e n
Bo neo, bu he di e ence is p obably one o pelage. I is seen mos
clea ly on he lowe back. In he Sempang skin he colo appea s
uni o mly da k b ownish, al hough he hai s a e e y ligh in colo
a he base; none o he hai s a e annula ed. In he specimens om
sou heas e n Bo neo he colo is e y b igh and " eddish," and a
dis inc g izzling is conspicuous. The indi idual hai s a e almos
whi ish a hei bases, g adually changing o a da k b ownish abou
he middle po ion, ollowed by a well-ma ked b igh hazel ing and
a blackish e minal band. A mo e o less well- denned blackish-b own
s ipe ex ends along he back o he animal. The pelage di e ence
appea s o be independen o season. The h ee specimens showing
he g izzling we e collec ed in Feb ua y, Ap il, and No embe ,
espec i ely; he da k ung izzled skin was collec ed in Augus .
The single specimen om Pulo Lau shows no no ewo hy di e -
ence om he mainland indi iduals.
Gene ally common, especially abou Pamukang Bay. Those ound on Pulo Lau
had much ine ho ns han hose on he mainland. Two heads ob ained a Pamukang
Bay ha e ex emely poo ho ns, bu hey a e ce ainly no imma u e. P obably
he ood is be e on Lau . The e is good pas u age he e. The ca casses o dee
om bo h places a e abou he same size. — W. L. A.
70 PROCEEDINGS OF THE NATIONAL MUSEUM. ol. 40.
Ex e nal and c anial measu emen s o Bo nean samba s.
RUSA KUHLII (Miille and Schlegel).
1839-44. Ce us kuhlii Mulle and Schlegel, Ve h. Na . Gesch.
Bezi ., p. 223.
Nede l.
Th ee pai s o an le s o his small dee we e b ough back by
Doc o Abbo , om he island o Bawean, he ype-locali y. (Fo
measu emen s see he las h ee specimens men ioned in he able
below.)
Measu emen s o an le s o Samba s om Bo neo and Pulo Bawean.
Locali y.
Uppe Sempamg Ri e
Do
Pulo Bauwal
Kenda angan Ri e . . .
Do
Do
Do
Do
Do
Do
Do
Mambulah Ri e
Do
Mankol
Fulo Lau
Do
Pamukang Bay
Do
Uppe Pasi Ri e
Sa a ai Ri e
Bawean Island
Do
Do
Ca a-
logue.
No.
145309
145370
153773
153774
153775
153776
153777
153778
153779
153780
153781
153782
153783
153784
151856
151857
« 151860
151861
154413
154414
151853
151854
151855
Ci cum-
e ence o
an le
abo e
bu .
mm.7
80 (8)
154 143
146 147
175 178
104 100
130 135
112 111
172 177
145 146
145 142
171 75
163 163
220 (»)
200 204
167 163
157 167
98 98
101 100
172 173
150 149
95 95
112 115
102 105
Ci cum-
e ence o
an le
abo e
b ow ine.
mmJ
65 (8)
109 102
107 107
131 130
118 115
100 95
93 90
102 105
122 116
100 103
122 (ii)
108 103
178 (13)
167 170
148 140
110 110
82 81
85 81
139 139
120 112
02 64
75 75
70 70
Leng h
o an le
along
con exi y
o cu e.
mm.7
231 (8)
370 406
412 410
409 395
435 442
394 405
435 423
415 407
416 405
490 407
450 195
466 440
445 (13)
432 435
545 502
430 435
319 341
287 274
438 435
472 485
324 315
372 374
354 358
Bu o
ip o
b ow ine
along con-
exi y.
mm.7
(9) (8)
122 115
234 230
128 122
208 217
215 212
167 101
200 187
235 239
147 188
215 (ii)
245 248
218 (")
205 233
200 195
240 230
08 60
105 95
(is) 255
185 172
118 110
192 105
142 136
Tip o
apical
ine o i s
angle
wi h
main
unk o
an le .
(9)
67
30
50
75
(■0)
51
52
108
43
(8)
70
37
80
67
15
46
50
80
39
1270 (ii)
70 78
140 (13)
72 60
108 125
72 76
78 5
(15) (15)
70 65
136 111
73 80
60 66
90 83
i Collec o 's measu emen s.
2 Collec o 's igu es in pounds compu ed o kilo
g ams.
3 Las mola s abou hal way h ough al eoli.
* Emb yo in u e us.
6 Es ima ed by collec o .
6 Gu ed.
7 Fi s igu e, le an le ; second, igh an le .
8 B oken o .
8 Imma u e, las mola s jus coming h ough, b ow ine alone p esen
i° Absen .
u Spike an le on igh side.
■2 A wo poin ed ine.
'3 A le d opped an le only.
H Fully adul as judged by oo hwea , bu no e
small size o an le s.
16 An le s de o med.
i6 B oken.
no. 1809. MAMMALS FROM BORNEO AND VICINITY— LYON. 71
MUNTIACUS PLEIHARICUS Kohlb ugge.
1896. Ce ulus pleiha icus Kohlb ugge, Na uu k Tijdsch . Nede landsch-
Indie, ol. 55, p. 192, and pla e acing p. 260. Type-locali y, Pleiha i,
sou heas e n Bo neo.
In 1906 x I desc ibed as new a mun jac om he island o Banka,
compa ing i wi h specimens o Mun iacus moscha us om Suma a,
and wi h h ee Bo nean mun jacs. A ha ime I assumed ha
he e was only one species o mun jac on Bo neo and used Kohl-
b ugge's name pleiha icus o i , hinking ha he an le s he igu ed
we e sligh a ia ions om he no mal ype and ha ing seen a pai
o simila looking an le s om Tenasse im. Two o hose Bo nean
specimens a e emales and he hi d is a pai o an le s om he
Sakaiam Ri e . Wi h he much mo e abundan ma e ial ha has
been collec ed by Doc o Abbo in Bo neo, I qui e ag ee wi h
Kohlb ugge ha wo dis inc o ms o mun jac occu on Bo neo,
and now conside ha he wo emale specimens we e co ec ly
iden i ied as M. pleiha icus in 1906, bu ha he single pai o an le s
do no ep esen M. pleiha icus bu belong o a species o be desc ibed
below and o which he Banka animal is me ely a sligh ly smalle
geog aphic ace.
Doc o Abbo secu ed six specimens o Mun iacus pleiha icus, h ee
om sou hwes e n and h ee om sou heas e n Bo neo; one young
emale, wo imma u e males, and h ee adul males. The an le s o
he adul males ep esen a much mo e ex eme ype han do hose
igu ed by Kohlb ugge. I is no imp obable, howe e , ha hey
a e newly- o med an le s, al hough he bone o ming hem looks
ully ma u e and as i hey had no been ecen ly co e ed wi h el e .
The ollowing is a desc ip ion o M. pleiha icus based on Doc o
Abbo 's specimens:
Gene al colo o he animal a ligh , dull och aceous, somewha like
och aceous-bu , da ke and b igh e along he middle line o he
back and neck, whe e he e is a conside able admix u e o a b own
somewha like P ou 's o ming an indis inc do sal line; ea s dull
b ownish on he ou side, whi ish on he inside; bases o ea s simila
o adjacen po ions o head; chin, unde side o neck, inne side o
o e legs, and inguinal egion whi ish o c eam-bu ; hind legs and
ou e side o o e legs simila o he gene al pale och aceous head and
body colo , excep o a sligh admix u e j b ownish along he
lowe legs, mos ma ked on he o e legs; ail whi ish, wi h a na ow
do sal s ipe o da k b ownish sligh ly mixed wi h och aceous.
Compa ed wi h he o he species o Bo nean mun jac, Ca . No.
151863, adul male om Pamukang Bay, M. pleiha icus is e y di -
e en . The gene al colo o he o he is a b igh eddish-b own,
i P oc. U. S.Na . Mus., ol. 31, p. 582, Decembe 24, 1906.
72 PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. 40.
e y simila o Ridgway's e uginous, wi h a ine black g izzle, while
he legs a e da k b ownish, in ma ked con as o he och aceous
legs o M. pleiha icus.
The e a e h ee good adul male skulls o M. pleiha icus, each o
hem wi h li le sho spike an le s e mina ing much longe , slende
pedicles. Unlike he an le s igu ed by Kohlb ugge, none o hem
show bu s. The leng h o he an le s a y be ween 22 and 35 mm.,
decidedly sho e han hose igu ed by Kohlb ugge. The skulls o
he o he wo males a e imma u e, showing long, slende pedicles
o so , spongy bone, wi h as ye no di e en ia ion o he ip in o
ha d an le subs ance.
The mos s iking cha ac e s o he skull o M. pleiha icus a e i s
smalle size and sho e and much slende e an le pedicles. In
addi ion o hese, i di e s in se e al mino de ails, such as ex ensi e
a icula ion o he uppe ex emi y o he p emaxilla wi h he nasal
ins ead o jus mee ing he nasal; supe io po ion o he lach ymal,
abo e he la ge pi , much na owe , he an e io supe o-ex e nal
su ace o he mala na owe and mo e poin ed; he a ch o e he
pos e io na es no smoo h and ounded bu ma ked by a mo e o
less well-de ined, ounded idge, pa o he basisphenoids; and in e -
pa ie al abou wice as wide as i is long, ins ead o abou h ee
imes. The ee h o he wo o ms o Bo nean mun jac do no show
any s iking di e ences; hey a e abou he same size in he wo
species, hence ela i ely la ge in M. pleiha icus. (See pi. 1, igs. 2
and 4, and pi. 2, uppe igu e.)
(Fo ex e nal and c anial measu emen s, see able below.)
Ex e nal and c anial measu emen s o Bo nean mun jacs.
1 Collec o 's measu emen s.
2 Collec o 's measu emen s in pounds and qua e s compu ed o kilog ams.
3 Second and hi d mola s no h ough al eoli.
4 Las mola s no h ough al eoli.
6. Gu ed.
6 Type.
7 U e us con ained a nea ly ma u e emb yo.
no. 1809. MAMMALS FROM BORNEO AND VICINITY— LYON. 73
MUNTIACUS RUBIDUS, new species.
1907. Mun iacus plciha icus, Lyon, P oc. U. S. Na . Mus., ol. 33, p. 550, Decem-
be 24, 1907. (One pai o an le s om he Sakaiam Ri e .)
Type.— -Skin and skull o adul male, Ca . No. 151863, U.S.N.M.,
collec ed a Pamukang Bay, sou heas e n Bo neo, Ma ch 20, 1908, by
D . W. L. Abbo . O iginal numbe , 5907.
Diagnos ic cha ac e s. — A medium-sized membe o he genus, much
la ge and da ke in colo han Mun iacus pleiha icus o Bo neo,
smalle han M. mosclia us o Suma a, and closely esembling M.
bancanus1 o Banka, bu sligh ly la ge and wi h da ke and longe
legs.
Colo . — Type: Gene al colo o uppe pa s o body hazel, iche
and da ke in he median line and o e he hind qua e s, e e ywhe e
wi h a ine blackish g izzle, on he shoulde s he hazel paling almos o
och aceous; neck a mix u e o dull awny och aceous and blackish,
he wo colo s abou equally mixed on he sides, and he blackish
in excess abo e; ace dull b ownish; op o head b igh hazel; ou e
su ace o ea s dull hazel, excep blackish ma gins; inne su ace o
ea s whi ish; co e ing o pedicles hazel, wi h a conspicuous black
line unning down hem and con inued down on he ace o e he
eyes; sides o head simila o sides o neck; chin and uppe po ion o
h oa whi ish; unde side o neck gene ally clay colo wi h da ke
g izzling; middle line o ches and uppe belly b ownish, some hing
like hai b own, ligh ening la e ally o d ab, which g adually blends
wi h he g izzle o awny och aceous and blackish o he sides o he
body; inguinal and axilla y egions whi ish, he ligh colo ex ending
pa way down he inne side o he legs; ou e side o o elegs black-
ish b own, a na ow a ea on inne side awny och aceous, a small
indis inc awny och aceous spo abo e each hal hoo ; hind legs
somewha da ke han o elegs, he inne side being ligh e only as a
as he heel, a e y ill-de ined awny spo abo e each hal hoo ; ail
deep ich hazel abo e, whi e on he sides and below.
An le s. — No mal o he genus, hose on he ype a he smalle
han usual, and he le one inju ed. (See pi. 3, igs. 1 o 7, pi. 1,
igs. 1 and 3, and pi. 2, lowe igu e.)
STcull and ee h. — The skull o Mun iacus ubidus is a once dis-
inguished om ha o M. pleiha icus by i s much la ge size, s ou e
an le pedicles, and a ma ked conca i y on i s do sal aspec a he
pos e io end o he nasals. I is, howe e , e y simila o he skull o
M. bancanus, he main dis inguishing cha ac e s being la ge size
and sho e an e o-pos e io diame e o he in e pa ie al. The ee h
in M. ubidus ha e abou he same size ha hey do in M. bancanus
1 Lyon, P oc. U. S. Na . Mus., ol. 31, p. 582, Decembe 18, 1900.
74 PROCEEDINGS OF THE NATIONAL MUSEUM. ol.40.
and hence a e ela i ely smalle . (See pi. 1, igs. 1 and 3, and pi. 2,
lowe igu e.)
Measu emen s. — See able, p. 72.
Specimens examined. — Two, he ype and an adul emale, Ca . No.
154383, om Pulo Ma a Si i, and he on le s and an le s o wen y
specimens, one om he Sakaiam Ri e and nine een om he
Kendawangan Ri e .
Rema ks. — Mun iacus ubidus is e y dis inc om he o he
Bo nean mun jac, and is e y eadily sepa able om he la ge
Suma an animal. I closely app oaches he mun jac o Banka, o
which i is sca cely mo e han a subspecies. This is ano he illus a-
ion o he close ela ion exis ing be ween he animals o Bo neo and
Banka. I ha e e e ed he adul emale om Pulo Ma a Si i o
M. ubidus, because he skulls o he wo specimens a e almos exac ly
alike. The skin o he Ma a Si i specimen, howe e , is ligh e in
colo , and he legs a e no nea ly so da k. I can no belie e ha he
mun jac on Ma a Si i is indigenous, bu hink i mus ha e been
b ough a he ecen ly o he island om he mainland o Bo neo.
STJS "VITTATUS."
1906. Sus w a us, Mille , P oc. U. S. Na . Mus., ol. 30, p. 748, June 13, 1906.
Two pigs o he Sus i a us g oups we e collec ed on Bawean
Island. Nei he o hem is ully ma u e. The olde , Ca . No.
151841, has he las mola s no qui e le el wi h he es o he oo h-
ow; he younge , Ca . No. 151840, has he las mola s jus b eaking
h ough he al eoli. The skull measu emen s o he wo specimens
a e espec i ely: Uppe leng h o skull, 330, 290; zygoma ic wid h,
132, 120; pa ie al cons ic ion, 37, 31; maxilla y oo h ow, 108,
; m2, 20 x 16, 20 x 16; m3, 31 x 18, .
SUS BARBATUS Mulle .
1839. Sus ba ba us Mulle , Tijdsch . Na uu l. Gesch. Physiol., ol. 5, p. 149.
(Type-locali y, Banje massing.)
1906. Sus ba ba us, Mille , P oc. U. S. Na . Mus., ol. 30, p. 739, June 13, 1906.
1907. Sus ba ba us, Lyon, P oc. U. S. Na . Mus. , ol. 33, p. 550, Decembe 24, 1907-
In addi ion o he six skulls o Sus ba ba us ob ained om he
na i es along he Landak Ri e in 1905, Doc o Abbo has since
ob ained wen y-nine adul specimens o his pig om Bo neo, and
adjacen islands. (See ables on pp. 76 and 77 o a lis o hese speci-
mens, wi h hei ex e nal and c anial measu emen s.) The ou
adul specimens om Pulos Pelapis and Panebangan a e age some-
wha smalle han specimens om he mainland, and hei ee h
show ela i ely mo e wea in p opo ion o hei age, as judged by
skull su u es. I can ind no sa is ac o y s uc u al cha ac e s by
which hey may be dis inguished om mainland animals. The
no. 1809. MAMMALS FROM BORNEO AND VICINITY— LYON. 75
excessi e wea o he ee h on hese island specimens is p obably due
o some di e ence in he cha ac e o he ood on he island as com-
pa ed wi h he Bo nean mainland.
One o he emales had ou e uses in u e o, ano he se en, one nine, one en, and
ano he ele en. One emale had se en young wi h he .
Ve y common abou Pamukang Bay. Ha e ne e been in a place whe e pigs we e
as common as in some locali ies in his dis ic . Fo some eason I go only h ee
boa s, bu he sows wi h hei young simply abounded. Twice while he schoone
was ancho ed a Tanjong K ame d o es o pigs a emp ed o swim ac oss Klumpang
Bay. I was absen up i e a he ime. My Malay skippe wen a e hem in a
small boa and bagged an old boa wi h a club. On he o he occasion hi een pigs
swam close o he essel, bu no boa was a ailable and hey passed wi hou moles a-
ion. Some imes hey ge in o kelongs ( ish ykes) a low ide. The sows appea o
go in gangs o h ee o i e, bo h when p egnan and when hey ha e hei young. I
appea s o ake mos o hei ime o p ocu e su icien ood when suckling, o I me
wi h hese gangs oaming abou nea ly all day. Sus ga gan ua does no appea o occu
in his dis ic (Klumpang Bay egion). No one had e e seen o hea d o i . A
small black hai y pig is said o be locally plen i ul. The Dyaks call i "By wangi"
(li e ally moon-pig). I saw no hing o i . I was said o be e y small, he males,
wi h usks, he size o goa s o less. I is p obably some so o Sus i a us. — W. L. A.
76 PROCEEDINGS OF THE NATIONAL MUSEUM. ol. 40.
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84 PROCEEDINGS OF THE NATIONAL MUSEUM. ol.40.
Skull and ee h. — These show no s iking peculia i ies, bu he
audi al bullae a e dis inc ly mo e in la ed han hey a e in he closely
ela ed Sciu us ca ima se, especially no iceable in he an e io segmen
o he bullae.
Measu emen s. — Fo ex e nal and c anial measu emen s o he ype
and se ies, see able on p. 83. S. pelapius is one o he smalle mem-
be s o he p e os ii g oup, abou he same size as S. ca ima se.
Specimens examined. — Th ee, all om Sou h Island, Pelapis.
Rema ks. — Sciu us pelapius is e y closely ela ed o he nea -by
S. ca ima se, he only dis inguishing cha ac e s being i s da ke and
" edde " shoulde and la ge audi al bullae. Two specimens in he
ca ima se se ies ha e shoulde s nea ly as da k as hose o pelapius,
bu none ha I ha e seen shows he mo e in la ed bullae.
SCIURUS DULITENSIS (Bonho e).
1901. Sciu us i a us duli ensis Bonho e, Ann. Mag. Na . His ., 8e . 7, ol. 7,
p. 451, May, 1901.
1907. Sciu us duli ensis, Lyon, P oc. U. S. Na . Mus., ol. 33, No. 1577, p. 555,
Decembe 24, 1907. (Kapuas Ri e egion.)
Since he collec ed nine specimens o plan ain squi els om he
Kapuas Ri e egion D . W. L. Abbo has secu ed ele en addi ional
specimens in sou hwes e n Bo neo, eigh in sou heas e n Bo neo, and
one on Pulo Lau . As a as he gene al colo a ion o he uppe
pa s and side s ipes is conce ned all he Bo nean specimens a e
ai ly cons an . They a y conside ably, howe e , as o he b igh -
ness o he unde pa s, as may be seen e en in specimens om one
locali y. The single specimen om Pulo Lau does no seem o be
sepa able om he o he s.
I is no ewo hy ha nea ly all he specimens om he Kapuas Ri e
egion, as well as se e al in he U. S. Na ional Museum om no he n
Bo neo, ha e he en i e unde side o he ails dis inc ly eddish o
u ous, while nea ly all he o he s om Bo neo ha e he en i e unde
side o he ail gene ally g ayish. This cha ac e is no absolu ely
cons an and a bes could only be conside ed a subspeci ic one. The
di e ence can no be a ibu ed o season, sex, o age. Wi hou
mo e specimens, especially well-p epa ed skins om no he n Bo neo,
i does no o he p esen seem ad isable o ecognize his di e ence
by name.
I may no be wi hou in e es o poin ou how easily he " ed"
colo o he unde pa s o he Sciu us i a us g oup o squi els may
become al e ed. The e a e in he museum wo specimens o Sciu us
duli ensis om no h Bo neo, collec ed by C. F. Adams in 1887. A
ha ime hey we e p ese ed by some pickling solu ion, p obably
sal and alum. The unde pa s a e now a dull inaceous-cinnamon,
ins ead o he usual b igh och aceous colo s. Some o he specimens
no. 1809. MAMMALS FROM BORNEO AND VICINITY— LYON. 85
collec ed in Sa awak by Hose, which ha e ecen ly had hei skulls
emo ed, and ha e been made o e in o mode n s udy skins, ha e he
o iginal colo s o he unde pa s mo e o less dulled and ende ed
somewha pinkish. The luid used o elax hese skins in he
emaking was p obably alcohol and wa e . In o de o elax he
ee in he la ge numbe o squi els o he i a us g oup ecen ly
collec ed by D . W. L. Abbo , he unlabeled hind oo and adjacen
pa o leg o each specimen was soaked o abou wen y- ou hou s
in ap wa e placed in a gal anized-i on ecep acle. In a g ea many
o he specimens he b igh och aceous- u ous colo s o he inne side
o he leg ha e become dulled and mo e pinkish on he oo and leg
ha was soaked. In he las case i can ha dly be p obable ha he
plain wa e b ough abou he change in colo . I is mo e likely due
o he p ese a i e, p obably a senic o a mix u e o a senic and
alum, used on he inside o he skin, and which had come h ough he
skin o he hai s by solu ion and osmosis.
SCIURUS LAMUCOTANUS, new species.
Type. — Skin and skull o adul emale, Ca . No. 145405, collec ed
on Pulo Lamuko an, o wes coas o Bo neo, May 8, 1907, by D .
W. L. Abbo . O iginal numbe , 5201.
Diagnos ic cha ac e s. — A " ed" bellied, " ed" oo ed membe o
he Sciu us i a us g oup, di e ing om S. duli ensis o he neighbo -
ing mainland o Bo neo in ha ing a gene ally mo e och aceous one
on he uppe pa s, ligh e unde pa s, b oade bu less clea da k
sides ipe and mo e bu y ligh side s ipe.
Colo . — Type: Uppe pa s o head and body, a ine g izzle o
och aceous-bu and blackish, he wo colo s abou equally p omi-
nen ; uppe su ace o ail simila bu g izzle coa se , and he
och aceous-bu da kening o och aceous and awny och aceous a
he ip; unde side o ail wi h he och aceous-bu eplaced by
och aceous and awny och aceous, he la e colo s being in excess
o he black; ligh side s ipe (5 by 65 mm.) bu y; da k side s ipe
(12 by 80 mm.) blackish, sp inkled wi h a ew ligh hai s; unde -
pa s and inne sides o o e and hind legs och aceous-bu ; ou e
sides o o e and hind legs simila o adjacen pa s o body; o e and
hind ee simila o ou e side o legs, bu mo e och aceous-bu ;
inne side o ea s dull awney och aceous; ou e side simila o head;
o bi al ing and egion abou mou h och aceous-bu . The e is
some li le di e ence shown in ce ain indi iduals o he se ies as o
b igh ness o unde pa s, bu all a e essen ially like he ype.
Skull and ee h. — These show no special cha ac e is ics and appa -
en ly do no di e om hose o examples o Sciu us duli ensis om
Bo neo.
86 PROCEEDINGS OF THE NATIONAL MUSEUM. ol.40.
Measu emen s. — Fo ex e nal and c anial measu emen s see able,
pages 90 and 91.
Specimens examined. — Eigh , all om Pulo Lamuko an.
Rema ks. — While Sciu us lamuco anus is su icien ly dis inc om
he Bo nean S. duli ensis o be ega ded as a sepa a e species, i is
no , howe e , a highly di e en ia ed o m.
SCIURUS SERUTUS Mille .
1906. Sciu us se u us Mille , P oc. U. S. Na . Mus., ol. 33, p. 58, July 23, 1906.
One specimen collec ed on Pulo Se u u, Ka ima a Islands, in 1904.
SCIURUS DATUS, new species.
Type.— Skin and skull o adul emale, Ca . No. 145393, U.S.N.M.,
collec ed on Pulo Da u, o wes coas o Bo neo, May 2, 1907, by
D . W. L. Abbo . O iginal numbe , 5153.
Diagnos ic cha ac e s. — A " ed" bellied, " ed" oo ed membe o
he Sciu us i a us g oup, simila o S. duli ensis o Bo neo, bu
dis inc ly smalle , and wi h wide da k side s ipes.
Colo . — Type: Uppe pa s o head and body a ine g izzle o
blackish and a dull ligh c eam-bu , in some ligh s appea ing dull
oli e-bu , he ou e ligh annula ions o he hai s being ligh e han
he inne mo e bu y ones; uppe su ace o ail simila in colo o
uppe pa s o body, bu g izzling much coa se , unde side o ail
wi h he c eam-bu la gely eplaced by och aceous, especially in-
middle line, whe e i is he p edomina ing colo ; ligh side s ipe
(4 by 65 mm.) dull c eam colo ; da k side s ipe (10-15 by 80 mm.)
blackish, sp inkled wi h a ew ligh hai s; unde pa s and inne
sides o o e and hind legs och aceous-bu , a he b igh ; ou e side
o o e and hind legs simila o adjacen pa s o body; o e and hind
ee gene ally simila o legs, bu mo e och aceous-bu y; ou side o
ea s, like op o head, inside mo e och aceous bu y; o bi al ing
pinkish-bu . The se ies o specimens o Sciu us da us is e y
uni o m in colo a ion, and di e s in no essen ials om he ype.
Two examples ha e he hai s o he unde pa s a he wo n, allow-
ing he bases o show and da ken and dull he och aceous bu .
Skull and ee h. — Aside om hei dis inc ly smalle size hese
show no no ewo hy cha ac e is ics.
Measu emen s. — Fo ex e nal and c anial measu emen s see able
page 90.
Specimens examined. — Six, all om Pulo Da u.
Rem,a ks. — In he colo 'o he unde pa s and o he side s ipes
Sciu us da us closely esembles S. lamuco anus; in he colo o he
uppe pa s i mo e nea ly esembles S. duli ensis om Bo neo. I
can be eadily dis inguished om ei he , howe e , by i s dis inc ly
smalle size.
no. 1809. MAMMALS FROM BORNEO AND VICINITY— LYON. 87
SCIURUS SIRIENSIS, new species.
Type.— Skin and skull o adul male, Ca . No. 151768, U.S.N.M.,
collec ed on Pulo Ma a Si i, Ja a Sea, Decembe 7, 1907, by D . W. L.
Abbo . O iginal numbe , 5580.
Diagnos ic cha ac e s. — A e y pale o m o he " ed" oo ed, " ed"
bellied Sciu us i a us g oup o squi els, gene ally simila o S.
lau ensis Mille ,1 bu sligh ly mo e pallid, and wi h la ge audi al
bullae.
Colo . — Uppe pa s o body a ine g izzle o c eam-bu and
blackish, he c eam-bu being in excess; ail bo h abo e and below
simila , bu he g izzle much coa se ; unde pa s, including h oa
and inne side o o e and hind legs, pale och aceous bu ; ligh
side s ipe (17 by 60 mm.) whi ish c eam colo ; da k side s ipe
abou he same leng h and wid h, dull pale blackish; op o head a
g izzle o bu y and blackish, he o me in excess; sides o head gen-
e ally bu y; ou side o hind legs simila in colo o back; ou side o
o elegs simila o adjoining pa s o body, bu mo e bu y; o e
and hind ee gene ally bu y o och aceous bu y; ea s simila o
adjoining pa s o body; whiske s black.
Skull and ee h. — These show no special peculia i ies and e y
closely esemble hose o Sciu us lau ensis. The bullae, howe e , a e
dis inc ly la ge in S. sinensis, so ha he wo species a e easily
dis inguished.
Measu emen s. — See able, page 90.
Specimens examined. — Six, all om Pulo Ma a Si i.
Rema ks. — The occu ence o wo species so closely esembling each
o he as Sciu us lau ensis and S. sinensis on islands o opposi e coas s
o Bo neo is in e es ing. I is p obably he esul o simila en i-
onmen s, bu I ha e been unable o ind any de ails conce ning he
condi ions on he wo islands.
SCIURUS ARENDSIS, new species.
Type.— Skin and skull o adul male, Ca . No. 154276, U.S.N.M.,
collec ed on A ends Island, Ja a Sea, No embe 24, 1908, by D .
W. L. Abbo . O iginal numbe , 6251.
Diagnos ic cha ac e s. — A membe o he Sciu us i a us g oup o
squi els, dis inguished by i s small size, ligh unde pa s, and
whi ish axilla y hai s; a he closely ela ed o i s geog aphic
neighbo , Sciu us sinensis, om Pulo Ma a Si i (see abo e), bu
smalle , wi h mo e whi ish hai s on unde pa s, and wi h da ke
uppe pa s.
Colo . — Type: Uppe pa s o head, neck, and body, and ou e
su aces o o e and hind legs and uppe su ace o ee , a ine g izzle
i P oc. Wash. Acad. Sci., ol. 3, p. 128, Ma ch 26, 1901, Pulo Lau . Na una Islands.
88 PROCEEDINGS OF THE NATIONAL MUSEUM. ol. 40.
o pale och aceous bu and blackish, he wo colo s abou equally
mixed; ail, bo h abo e and below simila , bu he g izzle coa se and
he och aceous bu p edomina ing on he unde side; unde pa s, and
inne side o hind legs, a colo closely app oaching Ridgway's pink
bu , becoming whi ish c eam colo in axilla y egion and abou chin,
and nea ly as ligh on inne side o o e legs; ligh side s ipe (6 by
50 mm.) bu y c eam colo , ill de ined; black side s ipe (12 by 45
mm.) blackish, s ongly lined wi h whi ish, and many o he hai s
wi h na ow och aceous bu ings; egion o nose, sides o head and
neck simila o uppe pa s, bu he och aceous bu deepe and mo e
conspicuous; a a he poo ly denned eye ing; ea s, inside and ou ,
simila o uppe pa s.
STcull and ee h. — Aside om i s sligh ly smalle size and ela i ely
hea ie ee h, he skull and ee h o Sciu us a endsis show no di e -
ences om hose o nume ous ela ed o ms.
Measu emen s. — See able, page 90.
Specimens examined. — Fi e, all om Pulo A ends.
Rema ks. — Sciu us a endsis is simply ano he o he nume ous
aces o Sciu us i a us, p oduced by isola ion on a ious Malayan
islands. I closely esembles i s geog aphic neighbo s, S. sinensis,
om Pulo Ma a Si i, and S. duli ensis, om he Bo nean mainland.
I di e s om S. sinensis in i s smalle size, da ke uppe pa s, and
in he possession o nume ous whi e hai s on he unde pa s, as in
S. i a us albescens (Bonho e) * om no he n Suma a. F om he
mainland S. duli ensis i is dis inguished by i s smalle size, ligh e
colo a ion, bo h abo e and below, and less conspicuous side s ipes.
SCIURUS POLIOPUS, new species.
Type.— Skin and skull o adul male, Ca . No. 151789, U.S.N.M.,
collec ed a Pamukang Bay, sou he n Bo neo, Ap il 5, 1908, by D .
W. L. Abbo . O iginal numbe , 5923.
Diagnos ic cha ac e s. — A membe o he i a us g oup dis inguished
by a e y dull " ed" belly and conspicuous g ay legs and ee .
Colo . — Uppe pa s o head and body, a ine g izzle o pale dull
bu -yellow and blackish, he wo colo s abou equally mixed; highs
and shoulde s simila , bu he bu y colo becoming g adually
eplaced by a ligh g ay on he ou e side o he o e and hind legs
and on ee , lowe pa s o inne side o o e and hind legs also g ay-
ish; ail, simila o uppe pa s o body bu he g izzle e y coa se,
appea ing somewha annula ed, and wi h he bu y colo e y ligh
abo e, bu da ke unde nea h; unde pa s, a e y dull och aceous
colo , anging om almos clay colo in he ype o a e y da k
1 Ann. Mag. Na . His ., se . 7, ol. 7, May, 1901, p. 446.
no. 1809. MAMMALS FROM BORNEO AND VICINITY— LYON. 89
awny och aceous in he specimen om Klumpang Bay; inne side
o he uppe po ions o o e and hind legs simila o unde pa s o
body; ligh side s ipe (abou 75 by 7 mm.), dull c eam colo ; da k
side s ipe abou he same leng h, bu nea ly wice as wide, blackish,
ai ly clea in colo o sp inkled wi h a ew bu y hai s. A sugges-
ion o he g ayish colo o he ee is seen abou he ex eme an e io
po ion o head.
Skull and ee h. — These show no special peculia i ies, and I ha e
been unable o ind cha ac e s in hem by which hey may be dis-
inguished om ela ed o ms.
Measu emen s. — See able, page 91.
Specimens examined. — Th ee, he ype om Pamukang Bay, one
om Klumpang Bay, and one om he Sa a ok Ri e .
Rema ks. — Sciu us poliopus needs no compa ison wi h any o he
o he known membe s o he i a us g oup. The combina ion o
" ed" belly and g ay ee occu s in no o he membe s o he g oup
so a as I am awa e, wi h he excep ion o he species o be desc ibed
below.
SCIURUS MARINSULARIS, new species.
Type.— Skin and skull o adul male, Ca . No. 151777, collec ed on
Pulo Lau , o sou heas e n Bo neo, Decembe 17, 1907, by D . W. L.
Abbo . O iginal numbe , 5619.
Diagnos ic cha ac e s. — A membe o he i a us g oup, wi h a ed
belly and g ay ee simila o Sciu us poliopus, desc ibed abo e, bu
wi h he unde pa s och aceous- u ous and wi h less g ay appea ing
on po ion o legs adjacen o ee .
Colo — Type: Uppe pa s o head and body and ou e su aces
o o e and hind legs a ine g izzle o blackish and a colo in e media e
be ween a ligh awny-oli e and bu -yellow, he wo colo s abou in
equal p opo ions; unde pa s, including inne sides o o e and hind
legs, och aceous- u ous; uppe su aces o o e and hind ee , wi h he
bu y colo o he uppe pa s eplaced by ligh g ayish, ail simila
o he uppe pa s, bu he bu y po ions o he hai ligh e and he
g izzle coa se , he ail in ce ain ligh s and po ions appea ing annu-
la ed; ligh side s ipe (abou 75 by 10 mm.), bu ; da k side s ipe
(abou 75 by 5 mm.), blackish. A sugges ion o he g ayish colo o
he ee is seen abou he ex eme an e io po ion o he head.
Skull and ee h. — These show no special peculia i ies, and I ha e
been unable o ind cha ac e s in hem by which hey may be dis-
inguished om ela ed o ms.
Measu emen s. — See able, page 91.
Specimens examined. — Six om Pulo Lau and wo om Pulo
Sebuku.
90 PROCEEDINGS OF THE NATIONAL MUSEUM. ol. 40.
Rema ks. — Sciu us ma insula is needs compa ison only wi h S.
polio-pus, desc ibed abo e. The wo species may be mo e closely
ela ed han he desc ip ions would indica e. The specimens ep e-
sen ing he mainland species a e e iden ly in no so esh a pelage,
which migh accoun , in pa a leas , o he s iking di e ence in
colo o he unde pa s o he wo o ms. The g ea e ex ension o
g ay abou he ee and legs o he mainland animal se es o dis in-
guish hem eadily, and would no appea o be p oduced by wea .
I is un o una e ha no c anial o den al cha ac e s can be ound o
sepa a e he g ay- oo ed squi els om he bu y- oo ed o ms. I
ha dly seems p obable ha he di e ence in colo o he ee and legs
can be dimo phic. Among he many squi els o he i a us g oup
collec ed by Doc o Abbo i has ne e been no iced be o e. The
only o he g ay- oo ed squi els o his g oup in he U. S. Na ional
Museum a e specimens o he g ay-bellied Sciu us no a us om Ja a.
All he o he g ay-bellied and " ed "-bellied species ha e he ee in
gene al concolo wi h he uppe pa s, o o en mo e bu y o och a-
ceous in colo .
Measu emen s o squi els o he i a us g oup.
1 Collec o 's measu emen s.
a Measu ed by w i e a e ee had been elaxed by soaking in wa e 24 hou s.
8 Type.
no. 1809. MAMMALS FROM BORNEO AND VICINITY—LYON. 91
Measu emen s o squi els o he i a us g oup — Con inued.
Name. Locali y. Ca a-
logue
No. Sex and age.
S. lamuco anus
Do
Do
Do
Pulo Lamuko an.
....do
do
....do
Do
duli ensis.
Do
Do
Do
Do
Do
....do
Sukadana
....do
....do
....do
Sempang Ri e .
Ba u Ju ong
.do.
Do.
Do.
Do.
Do.
Do.
Do.
Do.
Do.
Do.
Do.
Do.
.do.
.do.
Kendawangan Ri e .
....do
Klumpang Bay
....do
....do
Tjan ung
Balik Papan Bay
do
....do
Do
Do
S. poliopus
Do
Do
S. ma insula is .
Do
Do
....do
Pulo Lau
Klumpang Bay
Sa a ok Ri e
Pamukang Bay
Pulo Lau , eas side.
Pulo Lau
....do
Do.
Do.
Do.
Do.
Do.
....do
do »..
....do
Pulo Sebuku .
....do
145406
145402
145403
145404
1145405
145407
145408
145409
145410
145411
153075
153G70
153G77
153078
153079
153080
151785
1517S4
L51786
151787
L54289
154290
154291
154288
151779
151783
151788
1 151789
151774
151776
•151777
151780
151778
151775
151781
151 7S2
Male adul
Female, adul .
....do
Female, young
adul .
Female, adul .
Male, adul
....do
....do
Female, adul .
....do
Male, imma-
u e.
Male, young
adul .
Male, adul
Male
Female, adul .
....do
Male
Female, adul .
....do
Male, adul . . .
....do
....do
Male, young
adul .
Female, adul .
Female
Male
....do
Male, adul
Male
....do
Male, young
adul .
Male
Female
Female, adul .
....do
Female
;;/ in
214
220
225
216
224
I'll i
212
215
215
221
185
190
206
213
215
200
190
205
215
203
203
203
206
210
205
195
200
205
212
210
205
205
210
211
205
205
m m .
198
178
l.-.s
190
196
175
L90
185
185
182
180
m in
51
50
50
51
51
48
50
47
49
50
48
47
50
49
47
49
49
49
47
48
50
51
53
mm.
51.3
50.6
50.8
49.8
50.4
49.4
48.3
49.0
48.8
51.4
47.3
45.6
49.0
mm.
17.0
17.5
17.2
18.7
16.8
17.7
16.4
17.5
17.3
18.7
16.3
15.2
17.2
47.9
49.0 18.4
10.5
49.2
49.8
48.0
49.1
49.7
50.7
50.0
49.7
45.5
48.0
48.6
17.3
16.5
17.4
18.6
18.0
17.7
17.9
17.5
19.0
18.0
17.8
46.8
48.6 18.3
48.8
47.3
48.0
18.0
18.0
17.4
17.2
18.0
mm.
29.8
30.2
30.0
29.6
29.0
29.0
28.5
28.4
29.5
30.5
27.5
27.2
28.1
28.3
29.8
28.4
30.0
28.7
28.8
29.4
27.7
28.5
30.8
29.8
30.3
30.2
29.5
28.7
29.8
1 Type.
SCIURUS LOWII Thomas.
1892. Sciu us lowii Thomas, Ann. Mag. Na . His ., se . 6, ol. 9, p. 253, Ma ch, 1892.
Two specimens we e secu ed, an adul male om he Kenda-
wangan Ri e , sou hwes e n Bo neo, and an adul emale om he
Sa a ok Ri e , sou heas e n Bo neo.
(Fo measu emen s, see able, p. 92.)
SCIURUS PARVUS Mille .
1901. Sciu us pa us Mille , P oc. Biol. Soc. Wash., ol. 14, p. 33, Ap il 5, 1901.
Type-locali y, Nulu, Sa awak, Bo neo.
Th ee small squi els, membe s o he enuis g oup, we e secu ed
along he Sempang Ri e . They do no di e essen ially om
specimens o Sciu us pa us om no he n Bo neo.
(Fo measu emen s, see able, p. 92.)
92 PROCEEDINGS OF THE NATIONAL MUSEUM. ol. 40.
SCIURUS BORNEENSIS (G ay).
1867. Mac oxus u ogas e , a . bo neensis G ay, Ann. Mag. Na . His ., se . 3,
ol. 20, 1867, p. 283.
1901. Sciu us hippu us g ayi Bonho e, Ann. Mag. Na . His ., se . 7, ol. 7,
Feb ua y, 1901, p. 171, oo no e.
1907. Sciu us g ayi, Lyon, Smi hs. Misc. Coll., ol. 50, Ap il 8, 1907, p. 28.
One specimen, an adul male (Ca . No. 154293), om Balik Papan
Bay, is appa en ly indis inguishable om specimens o Sciu us bo -
neensis in he U. S. Na ional Museum collec ion om No h Bo neo.
(Fo measu emen s, see able, below.)
SCIURUS HIPPURELLUS Lyon.
1907. Sciu us hippu ellus Lyon, Smi hs. Misc. Coll., ol. 50, p. 27, Ap il 8, 1907.
1907. Sciu us hippu ellus, Lyon, P oc. U. S. Na . Mus., ol. 33, p. 557, Decem-
be 24, 1907.
Two om he Landak Ri e and one om he Kapuas, collec ed
in 1905.
Measu emen s o squi els.
RATUFA EPHIPPIUM (Mulle ).
1838-39. Sciu us ephippium Mulle , Tijdsch . Na uu l. Gesch. Physiol., ol. 5,
p. 147.
Doc o Abbo secu ed se en een specimens, p ac ically opo ypes,
o R. epliij^pium om he egion o Klumpang and Pamukang bays.
Mulle 's igu e3 is an excellen ep esen a ion o his species as
shown by unbleached specimens o he p esen se ies. The side o
head and op o nose in he pla e, howe e , a e oo " ed." In Doc-
o Abbo 's specimens hose po ions o he head a e simila in
1 Collec o 's measu emen s.
2 Measu ed by w i e a e elaxing in wa e o 24 hou s.
8Ve h. Na . Gesch. Nede l. Bezi ., pi. 13, 1839-1844.
no. 1809. MAMMALS FROM BORNEO AND VICINITY— LYON. 93
colo o he base o whiske s in M ille 's pla e. The g izzling seen
along he edge o he do sal s ipe in he pla e is less no iceable in
he specimens.
Rega ding he ype-locali y o his species, M . Ge i S. Mille , j .,
who examined in Leiden, in 1904, he ype, an adul male moun ed,
ema ked: "No locali y can be gi en beyond sou heas e n Bo neo, in
he low coun y."
(Fo measu emen s, see able, p. 96.)
RATUFA COTHURNATA, new species.
Type. — Skin and skull o adul emale, Ca . No. 145378, collec ed
a oo o Moun Palung, nea Sukadana, wes e n Bo neo, Feb ua y
9, 1907, by D . W. L. Abbo . O iginal numbe , 5537.
Diagnos ic cha ac e s. — A membe o he ephippium g oup, di -
e ing om ue ephippium in ha ing he da k do sal a ea less
sha ply de ined and gene al colo a ion no so b igh , mos closely
ela ed o Ra u a ephippium ba amensis Bonho e,1 bu di e ing in
ha ing he highs concolo wi h unde pa s o body.
Colo . — Type: Uppe pa s o head and body a mix u e o awny
och aceous and blackish, he o me mo e conspicuous an e io ly
and la e ally, he la e mo e p ominen in he do sal line and pos-
e io ly. The mix u e occu s as a g izzle, excep o e he shoulde s,
whe e he awny-och aceous is essen ially clea and unmixed, and
on he lowe back, whe e he blackish is unmixed and sp eads ou
la e ally nea ly as a as he emo al spo s. On he head be ween
he ea s he blackish is qui e pu e. Cheeks and sides o neck a he
da k awny-och aceous. Unde pa s bu o bu y. Fo e and hind
legs and ee bu o och aceous-bu , he inne sides o he legs
ligh e han he ou e . Gene al colo o ail abo e blackish, he
bases o he hai s whi ish, ma ginal po ions o unde side o ail
blackish ; cen al po ion, including sho app essed hai s, och aceous-
bu . Ea s inside and ou dull och aceous. The ype is one o he
b igh es and eshes -looking in he se ies. The majo i y o speci-
mens a e dulle and ligh e in colo , he awny-och aceous becoming
och aceous-bu o bu y and he unde pa s co espondingly ligh e ,
he blackish becoming b ownish. One specimen, Ca . No. 145381,
om Kumbang Ri e , is e y much bleached and wo n, he pelage
in places ha ing a singed appea ance.
Skull and ee h. — Appa en ly he e a e no cha ac e s by which he
skulls and ee h o R. co hu na a can be dis inguished om hose o
R. ephippium.
Measu emen s. — The ex e nal and c anial measu emen s o R.
co hu na a a e essen ially he same as hey a e in R. ephippium, he
hind oo and g ea es leng h o skull a e aging, howe e , a i le
less. The hind oo in R. co hu na a anges om 81 o 87 mm. and
i Ann. Mag. Na . His ., se . 7, ol. 5, p. 496, June, 1900. Type-locali y, Ba aia dis ic , Sa awak.
100 PROCEEDINGS OF THE NATIONAL MUSEUM. ol.40.
Measu emen s. — Fo ex e nal and c anial measu emen s o he
ype and se ies, see able, pages 101 and 102. The leng h o hind
oo o his subspecies uns om 38 o 44 mm. nea ly always o e
40 mm., o he ypical subspecies, 35 o 39, always unde 40.
EPIMYS NEGLECTUS LAMUCOTANUS, new subspecies.
Type.— Skin and skull o adul emale, Ca . No. 145497, collec ed
on Pulo Lamuko an, o wes coas o Bo neo, May 10, 1907, by
D . W. L. Abbo . O iginal numbe , 5224.
Diagnos ic cha ac e s. — A o m o Epimys neglec us Jen ink, cha -
ac e ized by a longe maxilla y oo h ow.
Dis ibu ion. — The islands o Lamuko an and Panebangan, o he
wes coas o Bo neo, and Pulo Ma a Si i, o he sou heas coas o
Bo neo.
Colo . — Type: Uppe pa s and sides o head, neck, and body, a
mix u e in abou equal p opo ions o blackish b own and a colo
be ween och aceous bu and clay colo ; unde pa s, c eam bu ;
ou side o legs simila o uppe pa s ; inne side simila o unde pa s ;
ail ligh b ownish, scan ily hai ed, h ee b ownish hai s o each scale,
and each hai abou he leng h o one and one-hal scales ; ou side o
ea s wi h sho b ownish hai s, inside wi h ligh dull bu y hai s.
Pelage. — As in Epimys neglec us ducis abo e.
Shull and ee h. — These a e o he same gene al o m and size as in
he alexand inus- a us g oup o a s. The maxilla y oo h ow
a e ages longe , om abou 7 o 7.6 mm., han i does in ypical
Epimys neglec us, abou 6.5 o 7.
Measu emen s. — Fo ex e nal and c anial measu emen s o he
ype and se ies, see able, page 101. The leng h o hind oo o his
subspecies uns om 35 o 40 mm., nea ly always unde 40, abou
he same as in he ypical subspecies.
Rema ks. — The Ma a Si i skins ha e on he a e age sligh ly longe
hind ee han do skins om Panebangan and Lamuko an. Some o
hem ha e ee qui e as la ge as do he membe s o he p eceding
subspecies, bu hey seem closes o he p esen o m.
no. 1809. MAMMALS FROM BORNEO AND VICINITY— LYON. 101
Measu emen s o Epimys neglec us.
1 Collec o 's measu emen s. 2 Skull only, bu specimen measu ed in lesh by collec o . 3 Type.
102 PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. 40.
'Type. 2 Skull only, bu specimen measu ed in lesh by collec o .
EPIMYS SEBUCUS, new species.
Type. — Skin and skull o adul male, Ca . No. 151964, collec ed
on Pulo Sebuku, o sou heas e n coas o Bo neo Janua y 2, 1908,
by D . W. L. Abbo . O iginal numbe , 5717.
Diagnos ic cha ac e s. — A membe o he Ej imys i mus 3 g oup,
di e ing om he ypical o m in a mo e eddish b own gene al col-
o a ion.
Colo . —Type: Uppe pa s and sides o head, neck, and body
and ou e sides o legs a coa se g izzly mix u e o blackish and a
colo some hing be ween usse and awny oli e, he usse mo e
conspicuous pos e io ly; unde pa s and inne sides o legs, bu o
bu y, he hai s p ac ically he same colo o hei bases; ail uni-
o mly b ownish h oughou ; ee simila , ea s da k b ownish, e y
scan ily hai ed wi h sho da k hai s.
Pelage.— Pelage o h ee kinds o hai s, sho , uni o mly g ayish
colo ed unde u ; longe hai s wi h da k bases, a usse - awny oli e
middle po ion and blackish ips; and long, slende g oo ed b is les,
uni o mly blackish in colo ; he h ee ypes named in he o de c
hei abundance. Middle po ion o ail wi h en scales o he cen-
1 Mille , P oc. Acad. Na . Sci. Phila., 1902, p. 155, June 11, 1902.
no. 1809. MAMMALS FROM BORNEO AND VICINITY— LYON. 103
ime e , each scale sub ended by h ee hai s, equalling a scale o a
scale and a qua e in leng h.
Slcull and ee h. — These show no special cha ac e s, and I can
ind no essen ial di e ences o dis inguish hem om he skulls
and ee h o Epimys Ji mus.
Measu emen s. — Fo measu emen s o he ype and se ies see
able, page 104.
Specimens examined. — Ele en, all om Pulo Sebuku.
Rema ks. — Epimys sebucus a bes can only be conside ed a sligh ly
di e en ia ed o m o he widely dis ibu ed Epimys Ji mus g oup,
o which he e a e many examples in he museum om Suma a
and adjacen islands and he Rhio-Linga A chipelago. No membe s
o he g oup we e aken on Bo neo.
EPIMYS CRASSUS, new species.
y;/i)e._Skin and skull o adul male, Ca , No. 145471, collec ed on
Pulo Lamuko an, o he wes coas o Bo neo, May 8, 1907, by
D . W. L. Abbo . O iginal numbe , 5190.
Diagnos ic cha ac e s .—A la ge membe o he Epimys ji mus l
g oup, di e ing om he ypical o m in la ge size, la ge and
hea ie skull, and a a he p ominen swelling on he an e io
po ion o he nasal bones.
Colo . Type: Uppe pa s and sides o head, neck, and body,
and ou e sides o legs a coa se g izzly mix u e o blackish and bu ,
sligh ly da ke and a he dulle han ha o Ridgway; unde pa s
and inne sides o legs c eam colo , he hai s essen ially he same
colo h oughou ; ail and ee ligh b ownish; ea s da k b ownish,
e y scan ily co e ed wi h sho , da k hai s.
Pelage.— Pelage o h ee kinds o hai s; sho , uni o mly g ayish
colo ed unde u ; longe hai s wi h da k bases, a bu y middle
po ion, and blackish ips; and long, slende g oo ed b is les, black-
ish in colo ; he h ee ypes named in he o de o hei abundance.
Middle po ion o ail wi h eigh and one-hal scales o he cen i-
me e , each scale sub ended by h ee hai s, equally abou a scale
and a qua e in leng h.
Slcull and ee h.— The skulls o Epimys c assus a e age la ge and
hea ie han do hose o E. Ji mus o like age, and mos o he speci-
mens ha e an elonga ed swelling on he ou e side o he an e io
hal o he nasal bones, which is ba ely indica ed on he skulls o
o he membe s o he Ji mus g oup. The ee h a e age sligh ly
smalle ' han hey do in Epimys sebucus. The bulla?, also, a e a
i le smalle in E. c assus.
Measu emen s— F 'o measu emen s o he ype and se ies see
able, page 104.
Specimens examined .—Fou een, om Pulo Lamuko an.
i Mille , P oc. Acad. Na . Sci. Phila., 1902, p. 155, June 11, 1902.
104 PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. 40.
Rema ks. — Epimys c assus appea s o be a well-de ined membe o
he E. j mus g oup. I is la ge han any*o he membe s o he
g oup in he U. S. Na ional Museum. The swelling o he nasals is
also qui e cha ac e is ic. Ex e nally i can no be di e en ia ed
om ypical jl mus.
Measu emen s o a s o he i mus g oup.
1 Collec o 's measu emen s 1 Type.
EPIMYS SABANUS Thomas.
1887. Mus sabanus Thomas, Ann. Mag. Na . His ,, se . 5, ol. 20, p. 269, 1887.
Six specimens om he mainland o Bo neo, a ully adul emale
om he oo o Moun Palling, a young adul emale om he
Sa a ok Ri e , and ou young adul emales om Balik Pap an Bay,
and a young adul emale om Pulo Lau . The ully adul specimen
ag ees in all essen ial espec s wi h an example om no he n
Bo neo. The younge specimens, howe e , a e dis inc ly smalle ,
mo e so han hei age would indica e. I is possible ha hey
ep esen ano he o m o he Epimys sabanus g oup, bu he ma e-
ial a hand is oo scan y o de e mine his.
(Fo measu emen s see able, p. 106.)
EPIMYS NASUTUS, new species.
Type. — Skin and skull o adul male, Ca . No. 145519, collec ed on
Pulo Panebangan, o wes coas o Bo neo, May 21, 1907, by D . W. L.
Abbo . O iginal numbe , 5270.
no. 1800. MAMMALS FROM BORNEO AND VICINITY— LYON. 105
Diagnos ic cha ac e s. — A membe o he Epimys sabanus1 g oup
cha ac e ized by a sho e ail, somewha smalle skull, and dis-
inc ly hea ie os um.
Colo . — Type: Uppe pa s and sides o head, neck, and body and
ou e sides o legs a mix u e o blackish and o ange bu , he o me
in excess in he median line, and he la e in excess along he sides
and on he legs; unde pa s, including inne sides o legs, c eam colo ,
ligh e on he h oa han elsewhe e; ea s da k b ownish, scan ily
clo hed wi h a ew da k hai s; ail b ownish, no no iceably ligh e
below han abo e.
Pelage. — Pelage composed o h ee kinds o hai s — a a he scan y,
dull, d ab-g ay colo ed unde u ; hai s o no mal ex u e, wi h dull
g ayish bases, succeeded by a da k b ownish ing, hen he con-
spicuous o ange bu band and a small blackish apex; la ened and
g oo ed spines, dull g ayish a base and blackish a ex emi y. On
he unde pa s he spinous hai s a e sho e and weake , and all he
hai s a e uni o mly c eam colo h oughou . Middle po ion o ail
wi h se en and one-hal scales o he cen ime e , each scale sub-
ended by h ee hai s abou a scale and a hal in leng h.
STcuU and ee h. — The skull and ee h a e abou he a e age in size
o he g oup, bu he skull is angula and massi e; he b ain case
and in e o bi al egion being wide han usual, and he os um
unusually hea y.
Measu emen s. — Type: Head and body, 252 (253) 2 mm.; ail, 326
(329); hind oo , 54 (49); condylo-basal leng h o skull, 51.4 (51.7);
zygoma ic wid h, 25.8 (26.2); in e o bi al cons ic ion, 9.3 (9.4);
b ead h o b ain case abo e oo s o zygoma a 19.4 (19.4); g ea es
b ead h o os um, 11 (10.6); dep h o os um nea inciso s, 11.3
(11.5); maxilla y oo h ow, al eoli, 9.3 (9.3). (See able, p. 106.)
Specimens examined. — Th ee, all om Pulo Panebangan, wo skins
wi h skulls and one skull wi hou skin.
Rema ks. — Epimys nasu us di e s by i s hea y os um om all
he a s o he E. sabanus g oup ha I ha e seen wi h he excep ion
o E. balse, om Tana Bala o he Ba u Islands, o he wes coas o
Bo neo. The skull o he Panebangan animal is decidedly la ge ,
howe e , han ha o he a om Tana Bala. The ail o he la e
is a he sho and uni o mly b ownish in colo , like he ail o Epimys
nasu us. The ails o abou hal he membe s o he g oup a e dis-
inc ly bicolo , b ownish abo e and c eam colo benea h.
i Thomas, Ann. Mag. Na . His ., se . 5, ol. 20, p. 2G9, 1S87.
2 Measu emen s in pa en heses a e hose o a pa a ype, Ca . No. 145518, an old adul emale.
106 PROCEEDINGS OF THE NATIONAL MUSEUM. ol. 40.
Measu emen s o long- ailed a s.
EPIMYS WHITEHEADI (Thomas).
1894. Mils whi eheadi Thomas, Ann. Mag. Na . His ., se . 6, ol. 14, p. 457,
Decembe , 1894.
Abou i y specimens o small eddish b own, spiny a s, wi h
bicolo ails, we e collec ed in a ious pa s o Bo neo o on islands
nea he coas by Doc o Abbo . (Fo he exac locali ies see able
o measu emen s, p. 107.) I ha e called hem all Epimys whi eheadi,
al hough hey show much a ia ion, and I ha e been unable o com-
pa e hem wi h opo ypes om no he n Bo neo. Se e al o he
p esen se ies ha e a s ong endency o be mo e b own and less
" ed" on he uppe pa s and o ha e g ay ins ead o och aceous
bellies. Such specimens also a e age la ge ex e nally and c anially
han do he o he s. These specimens a e pe haps ep esen a i es o
Thomas's ecen ly desc ibed Epimys whi eheadi pe lu us,4 bu hey
ha e dis inc ly smalle skulls, sho e ails, and longe hind ee , and
a e ound in sou hwes e n Bo neo and Pulos Bauwal and Sebuku.
The se ies o skins om Balik Papan Bay a e e y uni o m in hei
b igh " eddish" colo a ion and look exac ly like a se ies o opo-
ypes o Epimys aspe (Mille ) 5 om T ong, Lowe Siam.
(Fo measu emen s see able, p. 107.)
1 Collec o 's measu emen s.
2 Type.
3 Skull only.
4 Ann. Mag. Na . His ., se . 8, ol. 7, p. 205, Feb ua y, 1911.
6 P oc. Biol. Soc. Wash., ol. 13, p. 145, Ap il 21, 1900.
no. 1809. MAMMALS FROM BORNEO AND VICINITY— LYON. 107
Measu e ien s o Epimys whi eheadi.
Locali y.
Summi o Moun Palung...
Do
Do
Kendawangen Ri e
Do
Do
Do
Do
Do
Do
Do
Do
Do
Do
Pulo Bauwal
Do
Do
Do
Do
Do
Do
Do
Do
Do
Pulo Sebuku
Do
Do
Do
Do
Do ,
Do
Balik Papan Bay
Do
Do
Do
Do
Do
Do
Do
Do
Do
Do
Do
Do
Do
Do
Do
Ca a-
logue
No. S a e o wea o ee h.
145553
145554
2 145555
155720
155721
155722
155723
2155724
2155725
155720
2155727
2155728
155729
155730
153710
153711
153712
153713
153714
153715
153716
153717
153718
153719
151933
151934
151937
151938
151939
3151940
151941
154321
154322
154323
151324
151325
154326
151327
154328
154329
151330
154331
154332
151333
154335
154336
151337
Male.
Female. | Mode a ely wo n.
..do.
in in .
130
133
128
...do I Unwo n
Sligh ly wo n
Mode a ely wo n I 129
Much wo n 131
Female .
..do....
..do....
Male
Female .
A ale
Female .
Male
Female
...do...
...do...
Female
...do...
...do...
...do...
..do...
Male . . .
Female
Mode a ely wo n.
....do
....do
....do
....do
....do
....do
....do
Sligh ly wo n
....do
Mode a ely wo n.
....do
....do
Sligh ly wo n
Mode a ely wo n.
Much wo n
Mode a ely wo n.
Do 154338
Male....
..do....
..do....
Female .
..do
Male —
Female .
Male
..do...
..do...
...do...
Female
Male. ..
Female
...do...
Male . . .
...do...
Female
Male. . .
Female
Sligh ly wo n
Mode a ely wo n.
....do
Sligh ly wo n
Mode a ely wo n.
....do
....do
....do
m m .
108
125
102
;// in
32
33
30
Male. .-
Female
Mode a ely wo n.
....do
....do
....do
do
do
do
do
do
do
do
Mode a ely wo n.
do
117
125
112
118
120
128
113
122
122
134
US
122
119
128
117
125
mm.
34.' 6
105 27
30.0
30.4
32.2
15.6
S S
6.2
31.5
30.5
33.0
29.7
31.2
30.0
31.0
30.7
31.4
30.9
28.7
30.8
34.0
30.4
14.0
13.0
14.0
14.5
15.6
14.0
14.0
15.3
14.6
14.4
14.4
14.3
14.7
14.0
14.8
15.4
14.4
32.7
34.0
31.4
31.8
30.6
31.5
31.5
33.0
30.6
31.6
31.0
32.4
31.0
32.0
33.1
31.3
15.0
15.4
15.0
14.2
14.8
14.8
14.6
14.5
31.3
14.5
14.7
14.2
14.9
14.7
14.2
14.3
14.2
14.0
5.2
5.2
5.2
4.9
5.7
5.7
5.4
5.9
5.9
5.4
5.3
5.6
5.4
5. 7
5.4
5.5
5.5
5.5
5.3
5.4
5.4
5.8
5.7
5.9
5.0
5.7
5.7
5.3
5.2
5.6
5.8
5.2
5.5
5.3
5.6
5.3
5.4
5.6
5.2
5.8
5.4
5.2
5.6
i Collec o 's measu emen s. 2 Belly g ay. s U P us con ained wo emb yos.
EPIMYS RAJAH Thomas.
1894. Mus ajah Thomas, Ann. Mag. Na . His ., se . 6, ol. 14, p. 451, Decembe ,
1894.
Abou nine y specimens o a s belonging o he Epimys ajah
g oup we e collec ed on Bo neo and many o he adjacen islands.
Abou hal o hem, emb acing all hose om he mainland o Bo neo,
and om Pulos Lamuko an, Da u, and Bauwal, may be ega ded as
ep esen ing he ypical o m. Those om he o he islands a e
e e ed o ou di e en o ms, h ee o which a e desc ibed below
as new. The specimens om sou heas e n Bo neo a e age b igh e
108 PROCEEDINGS OF THE NATIONAL MUSEUM. ol. 40.
in colo han do hose om he wes e n pa o he island, in his
espec esembling he o m occu ing on Pulo Lau .
(Fo measu emen s and locali ies, see able, pp. 110 and 111.)
EPIMYS SERUTUS.
1906. Mus se uVus Mille , P oc. U. S. Na . Mus., ol. 31, p. 59, July 23, 1906.
Twel e specimens collec ed on Pulo Se u u, Ka ima a Islands.
EPIMYS CAR1MAT.E Mille .
1906. Mus ca ima se Mille , P oc. U. S. Na . Mus., ol. 31, p. 59, July 23, 1906.
Type-locali y, Ka ima a Island.
Six a s o he Epimys ajali-su i e g oup collec ed on Pulo
Pelapis do no show any ma e ial di e ences om Epimys ca ima se,.
The e a e mo e specimens inclined o be da k in he Pelapis se ies
han in he Ka ima a se ies, bu none o hem a e da ke han he
da kes o hose om Ka ima a. Ele en specimens we e collec ed
on Ka ima a in 1904.
(Fo measu emen s, see able, p. 110.)
EPIMYS PERFLAVUS, new species.
Type.— Skin and skull o adul male, Ca . No. 151918, collec ed
on Pulo Lau , o sou heas coas o Bo neo, Decembe 25, 1907, by
D . W. L. Abbo . O iginal numbe , 5684.
Diagnos ic cha ac e s. — A membe o he Epimys ajah-su i e g oup
wi h he uppe pa s and sides s ongly colo ed wi h b igh och aceous
bu .
Colo .- — Type : Uppe pa s and sides o head, neck, and body and
ou e sides o legs b igh och aceous bu , spa ingly mixed wi h ligh
b ownish along he middle o he back; unde pa s whi ish o c eam
colo , his colo ex ending pa way down he inne side o he legs;
an indica ion o an och aceous bu colla on h oa (p esen in i e o
he pa a ypes and absen in h ee); ee whi ish; uppe su ace o
ail da k, unde su ace whi ish; ea s da k b ownish, scan ily clo hed
wi h da k hai s.
Skull and ee h. — These show no angible di e ences om he skulls
and ee h o Epimys ajah.
Measu emen s. — See able, pages 110 and 111.
Speci nens examined. — Ele en skins wi h skulls and wo skulls wi h-
ou skins, all om Pulo Lau .
Rema ks. — Epimys anambx Mille 1 shows he nea es app oach in
colo a ion o E. pe la us, bu he uppe pa s in he Anamba a s a e
conside ably da kened by b ownish, and he ex e nal measu emen s
a e no so la ge. Epimys anambx also appea s o ha e a smalle skull,
wi h u ela i ely sho e os um. The skulls o Malayan a s change
i P oc. Wash. Acad. Sci., ol. 2, p. 205, Augus 20, 1900.
no. 1809. MAMMALS FROM BORNEO AND VICINITY— LYON. 109
so much in shape and size wi h inc easing age and p esen so many
appa en indi idual a ia ions ha no e y much eliance can be
placed upon hem as de e mining species.
EPIMYS SATURATUS, new species.
Type. — Skin and skull o adul male, Ca . No. 145523, collec ed
on Pulo Panebangan, o wes coas o Bo neo, May 17, 1907, by
D . W. L. Abbo . O iginal numbe , 5236.
Diagnos ic cha ac e s. — A medium sized membe o he Epimys
ajah-su i e g oup o a s, dis inguished by i s da k o ange-bu and
blackish colo a ion.
Golo . — Type: Uppe pa s o head, neck, and body a mix u e o
o ange-bu , a he da ke han Ridgway's, and blackish, he la e
colo in excess along he back; sides o head, neck, and nea ly all o
legs nea ly clea o ange-bu , only spa ingly mixed wi h blackish;
unde pa s whi ish o c eam colo , ex ending pa way down inne
side o legs; an o ange-bu colla , 10 o 15 mm. wide, be ween h oa
and ches ; ee whi ish; ea s blackish; ail da k abo e, c eam colo
below.
Skull and ee h. — The skull and ee h o Epimys sa u a us do no
appea o di e om hose o he ela ed Epimys ajah and E.
ca ima se.
Measu emen s. — See able, page 111.
Specimens examined. — Twen y- h ee, six skins wi h skulls and
se en een skulls wi h no skins, all om Pulo Panebangan.
Rema ks. — Epimys sa u a us is one o he da kes and iches
colo ed membe s o he E. ajah g oup. O he species a e as da k,
such as he nea -by Epimys se u us om Pulo Se u u o he Ka ima a
Islands, bu he da ke colo is p oduced b}' an inc ease in he
blackish elemen s, while in E. sa u a us he ligh e colo s a e deepened.
EPIMYS UBECUS, new species.
Type. — Skin and skull o adul male, Ca . No. 151931, collec ed
on Pulo Sebuku, o sou heas coas o Bo neo, Janua y 3, 1908, by
D . W. L. Abbo . O iginal numbe , 5727.
Diagnos ic cha ac e s. — A membe o he Epimys ajah g oup,
gene ally esembling ajah, bu dis inc ly smalle .
Colo . — Type: Uppe pa s o head, neck, and body a mix u e o
da k b ownish and och aceous bu , he da ke colo mo e p ominen
in he middle a ea o he back; sides o head, neck, and body, and
ou e sides o legs, och aceous bu , nea ly clea bu sligh ly mixed
wi h b ownish; unde pa s and inne sides o legs whi ish o c eam
colo , he ligh colo no in e up ed a he h oa by a da ke colla ;
ee whi ish; ail da k abo e, whi ish benea h and a he ex emi y.
116 PROCEEDINGS OF THE NATIONAL MUSEUM. ol. 40.
Measu emen s o Vi e a angalunga.
Locali y. Ca a-
logue
No. Sex and age.
Sempang Ri e
Do.
Foo o Gunong Palung
Kumbaiig Ri e
Kendawangan Ri e ...
Do
Do
Klumpang Bay
Do
Do
Do
Sa a ok Ri e
Do
Balik Papan Bay
Do
Do
Do
Do
Pulo Panebangan
Pulo Lau
Pulo Bauwal
14.5338
145339
145340
145341
153842
153843
153844
1518(18
151869
151870
151871
151872
151873
154354
154355
154356
154357
154358
145337
151867
153841
Male, adul
Female, adul
Female, young adul . .
Female, e y young.. .
Male, adul
Female, adul
Male, old
Female, young adul . .
Male, young adul
Female, young adul . .
Male, adul
Female, adul
Male, old
Male, adul
Male
....do
Male, adul
do
Female, old
Female, adul
do
in in
675
lis:,
663
515
635
630
620
606
600
615
665
CHI
662
620
(ill)
600
605
643
651)
610
635
mm
350
325
330
240
300
3 1 5
313
310
335
300
275
300
315
315
300
310
315
320
310
310
305
in in
107
104
103
94
1ill
103
105
100
103
100
101
103
111!)
95
kilos
3.7
4.0
1.8
3.0
3.2
3.2
3.2
2.7
2.5
4.0
3.6
4.0
3.2
3.4
2. 8
3.2
3.6
4.7
2.8
4.0
mm.
117.6
114.3
113.7
98.9
111.0
108.7
112.5
107.5
110.2
109.4
114.9
110.6
115.5
113.0
114.7
m m .
58.0
54.8
54.3
44.6
58.3
57.5
57.7
53.1
54.5
54.0
60.6
56.3
61.4
58.4
58.0
nun.
46.8
43.1
44.0
44.2
42.9
44.7
41.5
42.8
42.6
44.6
43.5
45.3
46.2
45.6
110.0
113.9
111.4
111.8
112.3
57.3
59.6
59.0
53.5
55.0
43.4
44.7
41.7
43.6
43.9
'Collec o 's measu emen s. ' Collec o 's measu emen s in pounds and qua e s compu ed o kilog ams.
PARADOXURUS PHILIPPINENSIS (Jou dan).
1837. Pa adoxu us philippinensis Jou dan, Comp es Rendus, ol. 5, p. 523,
1837. Type-locali y, Philippine Islands, Luzon and Mindanao.
1885. Pa adoxu us philippinensis, Blan o d, P oc. Zool. Soc. London, 1885,
p. 800.
1907. Pa adoxu us philippinensis, Lyon, P oc. U. S. Na . Mus., ol. 33, p. 559,
Decembe 24, 1907. Kapuas Ri e egion.
D . W. L. Abbo has collec ed h ee addi ional specimens o
Pa adoxu us on Bo neo since his i s expedi ion in 1905, wo. om
he Sempang Ri e and one om Klumpang Bay. I ha e ollowed
Blan o d in calling hem P. philippinensis. Jou dan's accoun is
e y meage and he e a e no eally good specimens om he Philip-
pines in he Na ional Museum. Two skins collec ed by D . E. A.
Mea ns in he Philippine Islands, one o hem young and he o he
wi hou a skull, show no s iking di e ences om he Bo nean speci-
mens.
(Fo measu emen s see able, p. 117.)
ARCTOGALIDIA STIGMATICA (Temminck).
1835. Pa adoxu us s igma icus Temminck, Esquisses Zool. Co e de Guine, 1s pa ,
Mamm., p. 120. Type-locali y, Doeson (o Dusan) Ri e , sou he n Bo neo.
1907. A c ogalidia s igma ica, Lyon, P oc. U. S. Na . Mus., ol. 33, p. 559, De-
cembe 24, 1907. Landak Ri e , Bo neo.
Two specimens om Klumpang Bay, sou heas e n Bo neo. None
we e secu ed in he egions o he Kendawangan o Sempang i e s.
Measu emen s. — See able, page 117.
no. 1809. .]/ AMMALS FROM BORNEO AND VICINITY— LYON. 117
Measu emen s o i e ids.
HEMIGALUS HARDWICKII (G ay).
1S30. Vi e a ha dwickii G ay, Spic. Zool., ol. 2, p. 9.
One specimen; an adul male, om Balik Papan Bay. I is uni-
o m in size and colo wi h specimens om no he n Bo neo, as well
as om Suma a and he Malay Peninsula.
(Fo measu emen s see able abo e.)
HERPESTES SEMITORQUATUS (G ay).
1846. He pes es semi o qua us G ay, Ann. Mag. Na . His ., ol. 18, 1846, p. 211.
1907. He pes es semi o qua us, Lyon, P oc. U. S. Na . Mus., ol. 33, p. 559,
Decembe 24, 1907.
Skin and skull o a young male om Sanggau, Kapuas Ri e ;
p obably ep esen s his species, bu i is oo young o de e mine
posi i ely.
HERPESTES BRACHYURUS (G ay).
1837. He pes es b achyu us G ay , Mag. Na . His ., ol. 1, p. 578. Type-locali y,
Indian Islands.
Fou specimens — wo om he Kendawangan Ri e and wo om
Balik Papan Bay.
(Fo measu emen s see able abo e.)
CYNOGALE BARBATUS (Miille .)
1838-39. Po amophilus ba ba us Mulle , Tijdsch . Na uu l. Gesch. Physiol., ol.
5, p. 142.
Six specimens o his in e es ing animal we e secu ed by Doc o
Abbo . Mode n au ho s ha e uni o mly e e ed he Bo nean
Oynogale o he species desc ibed by G ay4 unde he name benne ii,
i Collec o 's measu emen s.
Collec o 's measu emen s in pounds and qua e s compu ed o kilog ams.
3 m3 hal way up.
4 Ann. Mag. Na . His ., ol. 1, 1837, p. 579.
118 PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. 40.
om Suma a. A ca e ul compa ison o a ine old skull o a emale
om Am Bay, Suma a, wi h he p esen se ies om Bo neo con-
inces me ha he Bo nean and Suma an o ms ep esen dis inc
species. The skull o Gynogale la ba us is sligh ly la ge han ha
o 0. benne ii, has mo e sp eading zygoma a, a mo e in la ed b ain-
case and a less conspicuous pos o bi al cons ic ion, he ee h a e
la ge and hea ie , he mos no iceable di e ence being in he las
uppe mola s. Midle 's igu e1 o he skull shows ha his specimen
was o he same ype as he p esen se ies o Bo nean skulls. (Fo a
compa ison o measu emen s o he wo o ms, see able below.)
Ex e nally he wo o ms a e essen ially alike. The wo Suma an
specimens, howe e , a e a he ligh e in colo on he unde pa s
han a e he Bo nean skins.
Th ee emb yos he size o walnu s. — W. L. A.
U e us con ained wo emb yos nea ly ma u e. — W. L. A.
Measu emen s o Cynogale benne ii and C. ba ba us.
LUTRA LOVII Gun he .
1876. Lu a lo ii Gun he , P oc. Zool. Soc. London, p. 736. Type-locali y,
Bo neo, opposi e island o Labuan.
1907. Lu a lo ii, Lyon, P oc. U. S. Na , Mus., ol. 33, p. 560, Decembe 24, 1907.
Two specimens om he Kapuas Ri e egion collec ed in 1905.
> Ve h. Na . Gesch. Nede l. Bezi ., pi. 17, 1839-1844.
2 Measu emen s by collec o .
8 Las uppe mola s jus appea ing h ough al eolus.
* Las milk mola s and canines s ill in place.
no. 1809. MAMMALS FROM BORNEO AND VICINITY— LYOS. 119
AONYX CINEREA (Illige ).
1815. Lu a cine ea Illige , Abh. Akad. Be lin, 1811, 1815, p. 99. Type-locali y,
nea Ba a ia, Ja a.
A specimen o he clawless o e , a young adul emale, Ca . No.
151879, was aken on Pulo Lau . The ee h a e mode a ely la ge,
abou he size ound in a specimen om he Kinaba angan Ki e ,
no he n Bo neo.1 The shape o he ascending amus o he mandible
closely esembles ha o a specimen om Pulo Se oko, Rhio-Linga
A chipelago.1 The skull is smalle han skulls o o he clawless
o e s in he collec ion, bu i is no so old.
Measu emen s. — Head and body, 460 mm.; ail, 310; hind oo ,
95; weigh , 5J pounds, equals 2.5 kilos; basal leng h o skull, 79.3;
zygoma ic wid h, 58.9; maxilla y oo h ow (al eoli), 29.6.
PUTORIUS NUDIPES Desma es .
1822. Mus ela nudipes Desma es , Mammalogie, p. 537, 1822. Type-locali y,
Ja a.
1842. Pu o ius nudipes, F. Cu ie , His . Na . Mamm., Table me hod., p. 3, 1842.
One specimen was secu ed a Tjan ung, sou heas e n Bo neo.
The gene al colo o body, abo e and below, is awny och aceous. A
na ow s eak om occipu , o e nape o be ween shoulde s, and
sides o neck, including on o shoulde s, a e a ligh e and b igh e
awny och aceous; head, including ea , whi ish o c eam colo ; ail,
be ween clay colo and och aceous, wi h he e minal hi d bu y.
Measu emen s. — Head and body, 366 mm.; ail, 222; hind oo , 58.
The single Pu o ius was sho as i was ying o en e a ca e. — W. L. A.
MUSTELA HENRICII Wes e man.
1848-1854. Mus ela (Ma es) hen icii Wes e man, Bijd . Die k, ol. 1, p. 13, and
unnumbe ed pla e. Type-locali y, Padang, Suma a. (See Jen ink, Mus.
d'His . Na . Pays-Bas, ol. 9, Ca . Os eol. Mamm. Leiden, 1887, p. 112.)
1901. Mus ela la igula hen icii, Bonho e, Ann. Mag. Na . His ., se . 7, ol. 7,
p. 346, Ap il, 1901.
Doc o Abbo collec ed wo specimens o he yellow- h oa ed
ma en in sou hwes e n Bo neo and one in sou heas e n Bo neo.
By a singula coincidence he h ee Bo nean specimens a e all emales,
and wo om Suma a a e bo h males. The e a e no essen ial
di e ences be ween he skins om he di e en islands. One o he
Bo nean specimens, howe e , Ca . No. 145579, om Gunong Palung,
is gene ally da ke han any o he o he s, and he an e io po ions
o he uppe pa s a e p ac ically as da k as a e he pos e io po -
ions and ail. The o he ou a e dis inc ly bicolo abo e. The
skulls and ee h o he Suma a ma ens, he males, a e dis inc ly
i See P oc. U. S. Na . Mus., ol. 36, pi. 39, June 1, 1909.
120 PROCEEDINGS OF THE NATIONAL MUSEUM. ol. 40.
la ge han hose in he Bo nean ma en, he emales. The di e -
ence in size is abou he same as ha usually ound be ween he
wo sexes in he Mus elidae.
(Fo measu emen s see able below.)
Measu emen s o specimens o Mus ela hen icii.
1 Collec o 's measu emen s.
2 Collec o 's measu emen s in pounds and qua e s compu ed o kilog ams.
HELARCTOS EURYSPILUS Ho s ield.
1826. Uela c os eu yspilus Ho s ield, Zool. Jou n., ol. 2, pp. 221-234, pi. 7.
1907. Ilela cios eu yspilus, Lyon, P oc. U. S. Na . Mus., ol. 33, p. 561, Decem-
be 24, 1907.
Since 1905 Doc o Abbo secu ed se en addi ional specimens o
he Bo nean sun bea , six skulls ob ained om na i es in sou h-
wes e n Bo neo, and a skin and skull collec ed a Pamukang Bay.
The p esen se ies o skulls shows ha he di e ences be ween he
Bo nean and Suma an sun bea s a e no as g ea as I was led o
belie e in 1907, bu he di e ences poin ed ou hen, hold in he
main, as may be seen on e e ence o he able o measu emen s on
page 121. The e is no doub as o he dis inc ness o he wo o ms.
The Suma an specimens ha e smalle colla s and he ligh a ea o
he nose is mo e conspicuous han in he single Bo nean skin.
Bea s seem o be p e y common whe e e he e is a plan a ion o cocoanu s. Some
o he ees a e said o be dead o dying as a esul o hei dep eda ions. The bea s
climb up and ea he "hea " ou o he palm. In a email g o e o abou nine y
ees, nea Tanjong Pamukang, he bea s had des oyed abou hal he ees. I
was ull moon du ing my s ay he e, so he owne s said i was no use o wa ch o
bea s, as hey only came ou on da k nigh s. The Du ch au ho i ies ook away all
he guns abou a yea ago, so now he animals ha e i all hei own way. — W. L. A.
no. 1809. MAMMALS FROM BORNEO AND VICINITY— LYON. 121
Measu emen s o Bo nean and Suma an sun bea s.
1 Collec o 's measu emen s.
TUPAIA SPECIOSA (Wagne ).
1840. Cl[adoba es] speciosa Wagne , Sch ebe 's Sauge hie e, Supplbd. 2, 1840,
p. 43.
1907. Tupaia speciosa, Lyon, P oc. U. S. Na . Mus., ol. 33, p. 562, Decembe
24, 1907.
Nine specimens o his ee-sh ew ha e been aken by Doc o
Abbo in Bo neo, wo in he Kapuas Ri e egion, h ee in sou h-
wes e n Bo neo, and ou in sou heas e n Bo neo. (Fo exac
locali ies, see able o measu emen s, p. 123.) The only adul specimen
om he Kapuas egion di e s in i s much la ge size, c anially and
ex e nally, om Doc o Abbo 's o he specimens as well as om
specimens om no he n Bo neo. In colo and o m, howe e , i
esembles he o he s in all essen ial espec s. I is no olde han
some o he smalle indi iduals. Wi hou mo e ma e ial i does no
seem ad isable o ecognize i as a dis inc o m.
(Fo measu emen s see able, p. 123.)
TUPAIA DORSALIS Schlegel.
1857. Tupaia do salis Schlegel, Handl. beoe . Die k., p . 1, p. 59, pi. 3, ig. 31,
1857.
1907. Tupaia do salis, Lyon, P oc. IT. S. Na . Mus., ol. 33, p. 562, Decembe 24,
1907.
One specimen om he Kapuas Ri e , collec ed in 1905.
(Fo measu emen s see able, p: 123.)
122 PROCEEDINGS OF THE NATIONAL MUSEUM. ol.40.
TUPAIA SPLENDIDULA G ay.
18G5. Tupaia splendidula G ay, P oc. Zool. Soc. London, 1865, p. 322. Type-
locali y, Bo neo, p obably no he n.
Th ee specimens, wo om he Sa a ok Ri e , Pamukang Bay, and
one om he Kendawangan Ri e .
M . Old ield Thomas, o he B i ish Museum, has kindly compa ed
one o he specimens wi h G ay's ype. He ema ks:
Compa ing 151884 wi h he ype o splendidula, i is he bes ma ch o ha ouble-
some species which I ha e ye seen, he colo being iden ical (allowing o a sligh
bleaching om black o b own in he old specimen), and he hind ee p ac ically he
same leng h. The skull o splendidula is, howe e , ce ainly a he la ge , as you say,
bu i is a e y old specimen. The e is no di e ence o he han size ha I can see.
Condylo-basal leng h o 151884, 42.5; o splendidula, 44.2.
The la ges o he h ee specimens collec ed by Doc o Abbo was
sen o M . Thomas, bu he di e ence in size be ween he ype o
splendidula and he h ee specimens om sou he n Bo neo is so
sligh and he ma e ial so limi ed ha i can be a ibu ed o indi-
idual a ia ion only.
(Fo measu emen s see able, p. 123.)
TUPAIA LONGIPES (Thomas).
1893. Tupaia e uginea longipes Thomas, Ann. Mag. Na . His ., se . 6, ol. 11, p.
343, May, 1893.
Fi e specimens, wo om sou hwes e n and one om sou heas e n
Bo neo. Tupaia discolo Lyon,1 om Banka, is e y close o his
o m, and in he o iginal desc ip ion should ha e been compa ed wi h
i . A specimen o he p esen se ies was sen o M . Old ield Thomas.
He ema ks as ollows:
Wi h ega d o longipes he di e ences you no e a e qui e ue as a as he ype is
conce ned, bu a la e specimen om he same egion has p ac ically he same wa m
colo on o e back, he same Jiypoch ysa-like ches , and he same g ayish ail. The
skulls a e iden ical, and he size he same.
(Fo measu emen s see able, p. 123.)
TUPAIA CARIMATjE Mille .
1906. Tupaia ca ima se Mille , P oc. U. S. Na , Mus., ol. 31, p. 61, July 23,
1906.
In addi ion o he o iginal se ies o Tupaia ca ima se. collec ed in
1904, Doc o Abbo ook one o he specimen in 1908.
(Fo measu emen s see able, p. 123.)
TUPAIA INFLATA Lyon.
1906. Tupaia in ia a Lyon, P oc. U. S. Na . Mus., ol. 31, p. 600, Decembe 18,
1906.
A specimen o a small Tupaia aken on Ka ima a Island in 1908
does no di e essen ially om T. in ia a o Banka and Billi on
' P oc. U.S. Na . Mus., ol. 31. p. 602, Decembe 18, 1906.
no. 1809. MAMMALS FROM BORNEO AND VICINITY— LYON. 123
Islands. I is a young adul emale wi h a somewha impe ec skull
no sui able o compa ison wi h skulls o he o iginal se ies. The
skin closely esembles he da ke indi iduals o he Banka-Billi on
specimens.
(Fo measu emen s see able below.)
TUPAIA GRACILIS Thomas.
1893. Tupaia g acilis Thomas, Ann. Mag. Na . His ., se . 6, ol. 12, p. 53, July,
1893.
One example o Tupaia g acilis was aken along he Kendawangan
Ri e .
(Fo measu emen s see able below.)
TUPAIA MINOR Giin he .
1876. Tupaia mind Gun he , P oc. Zool. Soc. London, 1876, p. 426.
Th ee small Tupaias a e e e able o Tupaia mino . Two a e
om he mainland o Bo neo, Kendawangan Ri e , and mou h o
Sempang Ri e , and hi d is om Pulo Lau . The la e is yellowe
unde nea h han a e he mainland specimens, his pe haps due o
s aining. I is also somewha da ke abo e, especially he ail.
The skull o he Pulo Lau specimen is oo badly b oken o compa e
wi h hose o he mainland animals.
(Fo measu emen s see able below.)
Measu emen s o ee sh ews.
Name.
T paia do salis.
Tupaia si*cciosa.
Do
Do
Do
Do
Do
Do
Tupaia splendi-
dula.
Do
Do
Tnpaialongip«s...
Do...:
Do.
Do
Do
Tupaia ca ima x.
Tupaia in la a. . .
Tupaia g acilis. ..
Tupaia mino . . .
Do
Do
Locali y.
Kapuas Ri e
Sungei Ma an
do
do
Sa a ok Ri e
Pasi Ri e
Balik Papau Bay.
Pamukang Bay. ..
....do
.do.
Kendawangan Ri e .
Ba u Ju ong
....do
Kendawangan Ri e .
....do,
Pangkallqhan Ri e
Ka ima a Island
....do
Kendawangan Ri e
Mon h o Sempang Ri e
Kendawangan Ri e
Pulo Lau .
Ca a-
logue
No.
142247
145573
142247
145574
151885
154339
154340
154341
157883
151884
153856
153852
153853
153854
153855
151882
153860
153859
153N5S
145575
153857
151881
Sex and age. jR
+3 ^S
Female, adul ..
Male, adul
do
Female, adul . .
do
Male, adul
Female, adul . .
Male, adul
do
do
Female, adul . .
Male, adul
Male, young
adul .
Female, nea ly
adul .
Male, adul
do
do
Female, adul ...
Male, adul
Female, adul . . .
Male, adul
....do
mm.
17.1
20.4
23.2
20.5
20.0
19.6
19.9
18.7
17.2
10.5
16.4
19.1
18.2
19.4
19.0
20.3
16.2
13.3
13.2
12.2
12.0
12.0
' Collec o 's measu emen s.
124 PROCEEDINGS OF THE NATIONAL MUSEUM. ol.40.
GYMNURA ALBA Giebel.
1909. Gymnu a alba, Lyon, P oc. U. S. Na . Mus., ol. 36, p. 453, May 27, 1909.
Six specimens om he Sempang Ri e , sou hwes e n Bo neo.
(Fo a ull accoun o and measu emen s o hese specimens see
pape e e ed o abo e.)
In one o hem a nea ly ma u e emb yo in u e o. — W. L. A.
GALEOPTERUS BORNEANUS, new species.
Type.— Skin and skull o adul emale, Ca . No. 151888, U.S.N.M.,
collec ed a Tjan ung, sou heas e n Bo neo, Janua y 30, 1908, by
D . W. L. Abbo . O iginal numbe 5775.
Diagnos ic cha ac e s. — A medium-sized membe o he genus
closely esembling he lying lemu om he Malay Peninsula1 and
he one om Bungu an o he Na una Islands,2 bu skull sligh ly
smalle han ha o G. peninsulse, and wi h dis inc ly sho e max-
illa y oo h ow and smalle audi al bullae; skull and maxilla y oo h
ow abou he same gene al size as in G. na unse, bu b aincase and
in e o bi al egion dis inc ly b oade and nasals mo e pinched up
in o idge.
Colo . — Type: Uppe su ace o head, neck, body, ail, ee , and
ou e su aces o limbs a ying shades o g ay, p oduced by i egula
mix u es o di y whi e and blackish, no ably abou he shoulde s
and hips he whi ish colo da kens o a bu yellow. The usual ligh
spo ing is ound on he ee , and a a he conspicuous whi e spo is
ound on each side o he base o he ail. The ou e bo de o he
memb ane abo e is dull ches nu . Gene al colo o he en i e unde
pa s excep chin and h oa , a ying mix u es and shades o dull
awny och aceous and clay colo ; chin simila o uppe pa s and
h oa , a g adual blending o he colo s o he chin wi h hose o he
unde pa s.
Skull and ee h. — The skull is o medium size, abou equal o ha
o Galeop e us na unse, bu he in e o bi al cons ic ion is wide , he
b ain-case decidedly wide , he nasals mo e pinched up in o a idge,
and he os um deepe ; audi al bullae o abou he same size, he
pala e, pos e io na es, and in e p e ygoid space decidedly wide in
he Bo nean o m. Compa ed wi h a skull om he Malay Penin-
sula, ha o he Bo nean animal is sligh ly smalle , wi h smalle
bullae, less in la ed mas oids. The ee h ha e abou he same size
ha he} do in he Malay o m, bu he oo h ow is sho e . The
indi idual ee h appea sligh ly la ge , bu he oo h ows ha e abou
he same leng hs in he Bo nean and Na unan o ms.
Measu emen s. — See able, page 126.
i Galeop e us peninsulx Thomas, Ann. Mag. Na . His ., se . 8, ol. 2, p. 303, Sep embe , 1908.
Selango -Pahang bounda y, Malay Peninsula.
2 Galeopi hecus na unx Mille , Smi hs. Misc. Coll., ol. 45, p. 50, No embe G, 1903, Bungu an Island,
No h Na unas.
no. 1809. MAMMALS FROM BORNEO AND VICINITY— LYON. 125
Specimens examined. — Two, he ype and nea ly adul male om
he Sempang Ri e .
Rema ks. — I qui e ag ee wi h Thomas * ha he e is a esemblance
be ween he lying lemu s o Bo neo, he Na unas, and he Malay
Peninsula. I is no plausible o suppose ha hey ep esen a single
species, no does he a he limi ed ma e ial a hand show his o be
he case. E en Thomas himsel has ecen ly named he Penin-
sula o m. Galeop e us .bo neanus can no be conside ed a s ik-
ingly dis inc species. I is ce ainly e y dis inc om G. ao is
Mille . I can see e y li le wisdom in Thomas and W ough on's
combining he Ao , Rhio A chipelago, Na una, and Bo nean Gal-
eop e us unde he name ao is.2 The only eal way o se le he case
is o ge oge he good se ies, bu meanwhile i seems bes o ecog-
nize he desc ibed o ms. Flying lemu s ough o be subjec o as
much geog aphic and insula a ia ions as a e o he g oups o mam-
mals.
GALEOPTERUS LAUTENSIS, new species.
Type.— Skin and skull o adul emale, Ca . No. 151886, U.S.N.M.,
collec ed on Pulo Lau o sou heas e n Bo neo, Decembe 23, 1907.
O iginal numbe , 5679.
Diagnos ic cha ac e s. — A medium-sized membe o he genus
closely ela ed o he Bo nean animal, bu sligh ly la ge , wi h a dis-
inc ly longe oo h ow and wi h he mas oids conside ably sh unken.
Colo . — The colo is essen ially like ha o he Bo nean animal
desc ibed abo e, bu he whi e spo ing occu s equen ly in la ge
splo ches, and he e is mo e och aceous in he colo o he back.
Skull and ee h. — The skull and ee h o Galeop e us lau ensis closely
esemble hose o he nea by Bo nean animal. The skull is sligh ly
la ge , mo e angula , wi h a na owe b ain-case, and less p ominen
mas oids. The oo h ow is conside ably longe .
Measu emen s. — See able, page 126.
Specimens examined. — Th ee; he ype, a young one aken wi h
he , and a male om he island o Sebuku.
Rema ks. — The male om he island o Sebuku is only p o isionally
e e ed o G. lau ensis. I has abou he size and gene al appea -
ance as i i migh be he male o ha species. Un o una ely he e
is no ma e ial wi h which o make a sa is ac o y compa ison.
Rega ding he ype-specimen, Doc o Abbo w i es:
This animal has a somewha in e es ing his o y. I was seized by a small o es
eagle (Spiza'e us limna'e us) and he wo ell o he g ound and we e seized by some
Malays who obse ed he occu ence. The place was in a small clea ing on he eas
side o Pulo Lau . The e we e some gigan ic ees s anding on he edge o he clea ing.
The " kubong " was on one o hese ees when he hawk g abbed he . She was ca y-
ing a newbo n young one. The e was an old wound ull o maggo s upon one shoulde ,
i Ann. Mag. Na . His ., se . 8, ol. 1, p. 254, Ma ch, 1908.
2 Jou n. Fed. Malay S a es Museum, ol. 4, p. Ill, 1909.
132 PROCEEDINGS OF THE NATIONAL MUSEUM. ol, 40.
Measu emen s o Rhinolophus.
EMBALLONURA MONTICOLA Temminck.
1838. Emballonu a mon icola Temminck, Tijdech . Na uu . Gesch. Physiol.
Leiden, ol. 5, p. 25. Type-locali y, Ja a.
Se en specimens om he Ma an Ri e , en om he Sa a ok
Ri e , Klumpang Bay.
(Fo measu emen s see able, p. 133.)
B ough by a Malay boy who said he go hem unde an o e hanging ock. * * *
B ough by a Malay boy who said he go hem in a ca e. W. L. A.
EMBALLONURA ANAMBENSIS Mille .
1900. Emballonu a anambensis Mille , P oc. Wash. Acad. Sci., ol. 2, p. 236,
Augus 20, 1900. Type-locali y, Anamba Islands.
1906. Emballonu a anambensis Mille , P oc. U. S. Na . Mus., ol. 31, p. 63,
July 23, 1906.
In addi ion o he wen y-six ba s o his species collec ed on
Ka ina a Island in 1904 Doc o Abbo secu ed nine een o he s in
1908. Fi e specimens om Pulo Panebangan may be e e ed o
he same species.
(Fo measu emen s o he Panebangan specimens, see able, p. 133.)
EMBALLONURA PUSILLA, new species.
Type.— Adul emale, p ese ed in alcohol, and skull emo ed,
Ca . No. 153940, collec ed a Mankol, Kendawangan Ri e , sou h-
wes e n Bo neo, Sep embe 29, 1908, by D . W. L. Abbo . O iginal
numbe , 6223.
Diagnos ic cha ac e s. — Like he Bo nean o m o Emballonu a
mon icola, bu wi h a dis inc ly smalle skull.
no. 1809. MAMMALS FROM BORNEO AND VICINITY— LYON. 133
Colo . -Type: D ied ou o alcohol empo a ily, hai o uppe pa s
be ween seal and mummy b own, becoming much ligh e a base
like a pale sepia; unde pa s simila o a a he da k B occoli b own'
wi h he hai s o he h oa and chin inclining owa d usse - ea s'
memb anes, e c., blackish.
Skull and ee c.— Essen ially like hose o he Bo nean Emballonu a
monhcola, bu dis inc ly smalle and weake ; space be ween he
canine and he la ge p emola dis inc ly less in he smalle ba
Measu emen s.— Type: Head and body, 36 mm.; ail, 9; o ea m
44; humb, 6; i s inge , 35; second inge , 60; hi d inge , 44- ou h
inge , 43; ibia, 17; oo , 8; ea om c own, 9; g ea es leng h o
skull, 13.3; zygoma ic wid h, 8.3; g ea es wid h o b ain case 6 5-
in e o bi al cons ic ion, 2.8; g ea es supe io wid h o os um' 5 3-
maxilla y oo h ow, including canine, 5.; mandibula oo h' ow'
including canine, 5.2. Also see able below.
Specimens examined.— Fou , wo om he Kendawangan Ri e and
wo om he Pangkallahan Ri e .
Hanging benea h a ock in he o es .— W. L. A.
Measu emen s o Emballonu a.
Name. Ca a-
logue
No.
Emballonu a mon i-
cola.
Do
1)0
Do
Do
Do
do ;
Emballonu a anam-
bensis.
Do
Do
Emballonu a pusilla
Do
Do
Locali y.
145658
145659
145660
145664
152099
152100
152102
145665
145666
145669
153939
153940"
152091
Ma an Ri e .
do
....do
....do ;;
Sa a ok Ri e ...
....do
....do
Pulo Panebangau
....do
...do
Kendawangan
Ri e .
....do
Pangkallahan
Ri e .
Female
..do....
.do....
Male
Female .
..do
..do
..do
..do....
..do....
..do....
.do.,
.do..
Ill III .
43
46
43
43
47
42
44
■12
43
•14
41
36
■Hi
mm
11
11
13
13
11
12
13
11
11
13
12
9
13
mm. mm
18 mm.
13.8
14.4
14.0
14.3
14.2
13.8
14.3
13.7
13.4
13.5
12.5
12.4
12.5
.a
>/3
mm.
5.9
6.1
5.8
6.0
6.0
5.7
6.0
5.6
5.7
5.7
5.2
5.0
5.0
1 Type.
MYOT1S CARIMAT.S Mille .
1906. Myo is canma se Mille , P oc. U. S. Na . Mus., ol 31 n 6? Jul 23
1906. '
Two specimens collec ed on Ka ima a in 1904 and h ee om he
Kendawangan Ri e in 1908. The mainland specimens do no ap-
pea o be di e en om he island o m.
(Fo measu emen s see able, p. 136.)
134 PROCEEDINGS OF THE NATIONAL MUSEUM. ol.40.
MYOTIS MURICOLA (G ay).
1841. Vespe ilio mu icola Hodgson, Jou n. Asia ic Soc. Bengal, ol. 10, p. 908
( Nomen nudum ) .
1846. Vespe ilio mu icola G ay, Ca . Spec. D aw. Mamm. Bi ds Nepal and
Thibe , p esen ed by B. H. Hodgson o B i ish Museum, p. 4.
1907. Myo is mu icola, Lyon, P oc. U. S. Na . Mus., ol. 33, p. 563, Decembe
24, 1907.
Thi y-se en specimens om Mankol, Kendawangan Ri e , nine
om he Sempang Ri e , and o y-one om Pulo Lau , all p ese ed
in alcohol.
(Fo measu emen s see able, p. 136.)
Caugh oos ing in plan ain lea es.
Roos ing in cu led-up young banana lea es.
Roos ing in he olds o he mainsail, ou nea he end o he main boom. — W. L. A.
GLISCHROPUS TYLOPUS (Dobson).
1875. Vespe ugo (Glisch opus) ylopus Dobson, P oc. Zool. Soc. London, p. 473.
1907. Glisch opus ylopus, Lyon, P oc. U. S. Na . Mus., ol. 33, p. 563, Decembe
24, 1907.
Fi y-six specimens om he Kapuas Ri e egion, collec ed in 1905.
KERIVOULA HARDWICKII (Ho s ield).
1825. Vespe ilio ha dwickii Ho s ield, Zool. Resea ches in Ja a.
1907. Ke i oula ha dwickii, Lyon, P oc. U. S. Na . Mus., ol. 33, p. 563, Decembe
24, 1907.
One specimen om he Kapuas Ri e in 1905.
KERIVOULA BOMBIFRONS, new species.
Type. — P ese ed in alcohol, skull emo ed, adul male, Ca . No.
145616, U.S.N.M., collec ed along he Ma an Ri e , wes e n Bo neo,
Augus 15, 1907, by D . W. L. Abbo . O iginal numbe 5497.
Diagnos ic cha ac e s. — A small membe o he genus, ligh in colo ,
wi h la ge ea s, appa en ly a he closely ela ed o Ke i oula pellu-
cida Wa e house om he Philippines, bu middle inciso s no unusu-
ally slende , and memb anes and legs mo e hai y.
Colo . — Type ; empo a ily d ied om alcohol : Uppe pa s, a ligh
wood b own wi h a sligh in o cinnamon ; unde pa s, a ligh ec u
d ab, bu a he dull; wing memb anes, b ownish some hing be ween
P ou 's and hai b own; bases o hai s ligh e in colo han ips; u o-
pa agium simila in colo o u o uppe pa s.
Memb anes, e c. — U opa agium mode a ely well u ed, hai s a
ee edge no longe o mo e nume ous han elsewhe e; legs and
do sum o oo qui e well u ed; o ea m, and ca pus and bases o
inge s wi h a he scan , sho hai s; edge (wid h o 4 mm.) o mem-
b ane om oo o i h digi , hai y o abou same ex en as u o-
pa agium, memb ane be ween elbow and oo mode a ely hai ed.
no. 1809. MAMMALS FROM BORNEO AND VICINITY— LYON. 135
Ea s. — La ge, ex ending abou 3 mm. beyond end o muzzle when
laid o wa d; inne ma gin uni o mly con ex; ou e ma gin wi h a
e y ob use angle abou opposi e ip o agus; agus, long (8.5 mm.)
slende (1.7 mm. wide a base), ape ing g adually o a poin , a
lowe ou e angle, a small ob use lobe, p ojec ing abou 0.5 mm.
Skull. — B ain-case a he na owed om side o side bu conside -
ably enla ged and swollen om abo e downwa d, an e io ^; in e -
o bi al egion na ow (3 mm.); a small conca e dep ession a base o
os um, in on o in e o bi al cons ic ion; as he skull es s on i s
na u al base he acial po ion is much il ed upwa d ; o wi h he
skull es ing on he maxilla y oo h ow, he c anial po ion is much
il ed upwa d; in compa ison wi h K. ha dwickii and K. pic a, he
po ion o he pala e pos e io o he oo h ow is e j^ na ow (abou
1.5 mm. as o 2.0 mm.)
Tee h. — Ve y simila o hose in he Kapuas Ri e specimen o
Ke i oula ha dwickii, excep ha m1 is mo e comp essed an e o-
pos e io ly, pm3 has a sligh ly g ea e c own su ace and less e ical
dep h as compa ed wi h he o he uppe p emola s; he lowe p e-
mola s in K. bombi ons a e ela i ely sligh ly la ge and wi h g ea e
an e o-pos e io diame e han hey a e in K. ha dwickii, especially
he middle p emola s.
Measu emen s. — Head and body, 39 mm.; ail, 40; o ea m, 30;
second digi , 30; hi d inge , 60; ou h inge , 47; i h inge , 41;
ibia, 14; oo , 9; ea om c own, 12; ea , ip o ip (wi hou s e ch-
ing), 25; g ea es b ead h o ea , 11; condylo-basal leng h o skull,
13; maxilla y oo h- ow, including canine, 5.6; mandibula oo h-
ow, on o canine o back o las mola , 6.2; zygoma ic wid h, 7.8;
wid h o b ain-case, 6.8.
Specimens examined. — One, he ype.
Rema ks. — The only o he species o Ke i oula hi he o eco ded
om Bo neo a e ha dwickii,1 pusilla,2 and papillosa.3 F om I . ha d-
wickii, K. bombi ons di e s conspicuously in i s na owe skull,
an e io ly in la ed b ain-case, na owe in e o bi al cons ic ion, and
by i s ligh e colo ; om K. pusilla, by i s gene ally la ge size and
dis inc ly la ge ea s; and om K. papillosa by i s much smalle size,
ela i ely la ge ea s, and ela i ely la ge inciso s.
Sho while hanging om a lea in dense o es . — W. L. A.
1 See page 134.
2 Thomas, Ann. Mag. Na . His ., se . 6, ol. 14, p. 461, Decembe , 1894.
a Hose, Mammals o Bo neo, p. 40, 1893.
136 PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. 40.
Measu emen s o ba s.
1 Type.
TARSIUS BORNEANUS Ellio .
1907. Ta sius a sie , Lyon, P oc.U. S. Na . Mus., ol.33, p. 565, Decembe 24, 1907.
1910. Ta sius bo neanus Ellio , Bull. Ame . Mus Na . His ., ol. 28, p. 153, May
27, 1910.
Fou specimens collec ed by Doc o Abbo in wes e n Bo neo in
1905.
NYCTICEBUS BORNEANUS Lyon.
1906. Nyc icebus bo neanus Lyon, P oc. U. S. Na . Mus., ol. 31, p. 535, No em-
be 9, 1906.
1907. Nyc icebus bo neanus, Lyon, P oc. U. S. Na . Mus., ol. 33, p. 565, Decembe
24, 1907.
Nine specimens om wes e n Bo neo, collec ed by Doc o Abbo
in 1905.
PITHECUS NEMESTRINUS (Linnaeus).
1766. [Simia] nemes ina Linn/Eus, Sys . Na ., ol. 1, 12 h ed., p. 35. Type-
locali y, Suma a.
1907. Macaca nemes ina, Lyon, P oc. U. S. Na . Mus., ol. 33, p. 565, Decembe
24, 1907. (Landak and Sakaiam i e s specimens.)
no. 1809. MAMMALS FROM BORNEO AND VICINITY— LYON. 137
Th ee specimens, a skin and skull om Sukadana, an odd wea he ed
skull om he Kendawangan Ri e , and a skin and skull om Pamu-
kang Bay. The wo skins ha e hai s ha a e dis inc ly annula ed,
and he skulls a e o he na ow ype, qui e unlike he skin and skull
o he ype o P. h oca Mille * om no he n Bo neo.
(Fo measu emen s see able, p. 138.)
PITHECUS MANDIBULARS Ellio .
1907. Macaca ascicula ^, Lyon, P oc. U. S. Na . Mus., ol. 33, p. 565, Decem-
be 24, 1907.
1910. Pi hecus mandibula is Ellio , P oc. U. S. Na . Mus., ol. 38, p. 347,
Augus 6, 1910.
Only one specimen o he usually common Malay long- ailed
macaque was ob ained by Doc o Abbo in Bo neo, on he Sungei
Sama, nea Pon ianak, in 1905.
(Fo measu emen s see able, p. 138.)
PITHECUS CARIMATjE Ellio .
1906. Macaca ascicula is Mille , P oc. U. S. Na . Mus., ol. 31, p. 65, July 23,
1906.
1910. Pi hecus ca i ia se Ellio , P oc. U. S. Na . Mus., ol. 38, p. 346, Augus 6,
1910.
Two skins and skulls omTelok Pai, Ka ima a Island, collec ed by
Doc o Abbo in 1904.
(Fo measu emen s see able, p. 138.)
PITHECUS CUPIDUS Ellio .
1910. Pi hecus cupidus Ellio , P oc. U. S. Na , Mus., ol. 38, p. 348, Augus 6,
1910.
Skin and skull and an odd wea he ed skull om Pulo Ma a Si i.
(Fo measu emen s see able, p. 138.)
PITHECUS BAWEANUS Ellio .
1910. Pi hecus baweanus Ellio , P oc. U. S. Na , Mus., ol. 38, p. 347, Augus 6,
1910.
Two skins and skulls om Bawean Island.
(Fo measu emen s see able, p. 138).
J P oc. U. S. Na . Mus., ol. 29, p. 558, Feb. 3, 1906.
138 PROCEEDINGS OF THE NATIONAL MUSEUM. ol. 40.
Measu emen s o Pilhecus.
PYGATHRIX RUBICUNDA RUBICUNDA (Mulle ).
1838-39. Se anopi hecus ubicundus Mulle , Tijdsch . Na uu . Gesch. Physiol.,
ol. 5, p. 137.
1839-1844. Semnopi hecus ubicundus, Mulle and Schlegel, Ve b. Na . Gescb.
Nede l. Bezi , pp. 61 and 69, pi. 9, igs. 1, 2, 3, and 1; pi. 11, ig. 1.
Type-locali y, sou heas e n Bo neo.
Doc o Abbo secu ed h ee specimens o Pyga h ix ubicunda
ubicunda om Balik Papan Bay and h ee om he Se a ok Ri e .
These six specimens a e p ac ically opo ypes o he species, he
o iginal specimens ha ing been collec ed a Moun Sakumbang and
Tana Lawu . (The la e place does no appea on any o he a ail-
able maps. Moun Sakumbang is shown on he map on page 57.)
They a e o a gene ally da ke ed colo han any o he o he
Bo nean specimens o his species in he Na ional Museum, and he
hands and ee ha e a mo e dis inc endency o be su used wi h
blackish.
The e seem o be h ee dis inc o ms o his species on Bo neo,
P. ubicunda ubicunda om sou heas e n Bo neo, P. ubicunda igni a 5
om he no he n pa s o he island, a new o m desc ibed below as
P. ubicunda ubida om wes e n Bo neo, sou h o he Kapuas Ri e .
A ou h subspecies, P. ubicunda ca ima se, om Ka ima a Island is
e iden ly de i ed om he Bo nean auna.
The di e en ial cha ac e s o he ou o ms a e se ou h in he
able below.
(Fo measu emen s, see able, p. 141.)
i Collec o 's measu emen s.
2 Collec o 's measu emen s compu ed o kilog ams.
3 Type.
1 Wea he ed skull.
6 Dollman, Ann. Mag. Na . His ., se . 8, ol. 4, p. 204, Sep embe , 1909.
no. 1809. MAMMALS FROM BORNEO AND VICINITY— LYON. 139
PYGATHRIX RUBICUNDA RUBIDA, new subspecies.
Type.— Skin and skull o adul emale, Ca . No. 153790, U.S.N.M.,
collec ed a Ba u Ju ong, sou hwes e n Bo neo, June 17, 1908, by
D . W. L. Abbo . O iginal numbe , 5979.
Diagnos ic cha ac e s. — Closely ela ed o Pyga h ix ubicunda ubi-
cunda, di e ing in ha ing a somewha ligh e colo , no conspicuous
su usion o blackish colo on he hands and ee , less p ominen ou e
edge o he mas oid, and a dis inc space be ween ha edge and he
edge o he pos e io oo o he zygoma.
Colo . — Type: Gene al colo simila o Ridgway's hazel, bu a he
b igh e , he hai s gene ally da ke a he ends and ligh e a bases,
so ha on he back whe e he hai s a e longes , he gene al colo is
da k hazel; a conside able numbe o blackish hai s on he do sal
su ace o he inge s and oes.
SJcull and ee h. — The skull and ee h o Pyga h ix ubicunda ubida
a e in gene al as hey a e in P. . ubicunda; he ou e edge o he
mas oid, howe e , is mo e inconspicuous han i is in he ypical o m
and is usually sepa a ed om he ou e edge o he pos e io oo o
he zygoma by a mo e o less well-de ined g oo e o dis inc space.
Measu emen s. — See able, page 141.
Specimens examined. — Twel e, om a ious locali ies in sou h-
wes e n Bo neo. (See able o measu emen s o exac locali ies.)
PYGATHRIX RUBICUNDA CARIMAT^: Mille .
1906. P esby is ca ima x Mille , P oc. U.S. Na . Mus., ol. 31, p. 65, July 23,1906.
In addi ion o he o iginal se ies o se en specimens collec ed in
1904, wo addi ional specimens we e collec ed on Ka ima a Island a
Telok Eda by D . W. L. Abbo in 1908.
(Fo ex e nal and c anial measu emen s see able, p. 141.)
Diagnos ic cha ac e s o he o ms o Pyga h ix ubicunda.
P. ubicunda ca ima x.
Ligh e colo ed, wi hou
well-ma ked blackish
su usion on hands and
ee .
Pos -glenoid p ocess long
and hea y.
Fossa be ween p e ygoids
ela i ely shallow.
Ou e edge o mas oid
ela i ely less conspicu-
ous.
Ou e edge o pos e io
zygoma ic oo as a ule
no sepa a ed om ou -
e mas oid edge by a
dis inc space o g oo e,
bu he wo edges soon
con luen .
Angle o jaw and condyle
la ge and hea y.
P. ubicunda ubida.
Ligh e colo ed, wi hou
well-ma ked blackish
su usion on hands
and ee .
Pos -glenoid p ocess no
conspicuously de el-
oped.
Fossa be ween p e y-
goids ela i ely shal-
low.
Ou e edge o mas oid
ela i ely less conspic-
uous.
Ou e edge o pos e io
zygoma ic oo sepa-
a ed om ou e mas-
oid edge by a mo e o
less well-de ined
g oo e o dis inc
space.
Angle o jaw and con-
dyle no so la ge and
hea y.
P. ubicunda ubicunda.
Da ke colo ed, wi h
blackish su usion on
hands and ee .
Pos -glenoid p ocess no
conspicuously de el-
oped.
Fossa be ween p e y-
goids in e media e in
cha ac e .
Ou e edge o mas oid
a he conspicuous.
Ou e edge o pos e io
zygoma ic oo as a
ule no sepa a ed
om ou e mas oid
edge by a dis inc
space o g oo e, bu
he wo edges soon
con luen .
Angle o jaw and con-
dyle no so la ge and
hea y.
P. ubicunda igni a.
Ligh e colo ed, wi hou
blackish su usion on
hands and ee .
Pos -glenoid p ocess no
conspicuously de el-
oped.
Fossa be ween p e y-
goids ela i ely deep.
Less conspicuous.
Ou e edge o pos e io
zygoma ic oo as a
ule no sepa a ed om
ou e mas oid edge by
a dis inc space o
g oo e, bu he wo
edges soon con luen .
Angle o jaw and condyle
no so la ge and hea y.
140 PROCEEDINGS OF THE NATIONAL MUSEUM. ol.40.
PYGATHRIX FRONTATA (Miille ).
1838-39. Semnopi hecus on a us Mulle , Tijdsch . Na uu . Gesch. Physiol.,
ol. 5, p. 136.
1839-44. Semnopi hecus on a us Mulle and Schlegel, Ve h. Na . Gesh.
Nede l. Bezi ., pp. 62 and 78, pi. 8, igs. 1-4.
Ten specimens o his a e monkey we e collec ed by Doc o Abbo
in sou heas e n Bo neo, eigh a Klumpang Bay, and wo a Balik
Papan Bay.
(Fo measu emen s see able, p. 141.)
Gene ally common. — W. L. A.
PYGATHRIX CRISTATA (Ra les).
1822. Simia c is a a Ra les, T ans. Linn. Soc. London, ol. 13, p. 244. Type-
locali y, Suma a.
1907. P esby is c is a a, Lyon, P oc. U. S. Na . Mus., ol. 33, p. 568, Decembe 27,
1907. Kapuas Ri e specimens.
In addi ion o wo specimens o P esby is c is a a aken by Doc o
Abbo in he Kapuas Ri e egion, an adul emale and he young
we e aken a he Pasi Ri e , sou heas e n Bo neo.
(Fo measu emen s o he specimens see able, p. 141.)
In poin o colo a ion he young specimen bea s no esemblance
wha e e o i s pa en . The hai s a e e e ywhe e och aceous in
colo , wi h he excep ion o a inge o long blackish hai s o e he
o ehead. A good illus a ion o he young may be seen in Ve han-
delingen o e de Na uu lijke Geschiedenis de Nede landsche o e -
zeesche Bezi ingen, pla e 12, ig. 1.
PYGATHRIX CHRYSOMELAS (Miille ).
1838-39. Semnopi hecus ch ysomelas Mulle , Tijdsch . Na uu l. Gesch. Physiol,
ol. 5, p. 138. Type-locali y, Pon ianak, wes e n Bo neo.
1907. P esby is ch ysomelas, Lyon, P oc. U. S. Na . Mus., ol. 33, p. 567, Decem-
be 27, 1907.
Nine specimens om Pon ianak, and he lowe Landak and Kapuas
i e s. None we e collec ed by Doc o Abbo in sou hwes e n o
sou heas e n Bo neo.
MAMMALS FROM BORNEO AND VICINITY— LYON. 141
Measu emen s o Pyga h ix.
1 Collec o 's measu emen s.
2 Collec o 's measu emen s in pounds and qua e s compu ed in kilog ams.
8 Type.
NASALIS LARVATUS (Wu mb).
1907. Nasalis la a us, Lyon, P oc. U. S. Na . Mus., ol. 33, p. 569, Decembe
24, 1907.
Ten specimens om he Kapuas Ri e egion, ou om sou hwes -
e n and i e om sou heas e n Bo neo.
(Fo exac locali ies and measu emen s see able, p. 142.
In sou heas e n Bo neo Nasalis la a us was common in he swampy lowlands a he
edges (sca cely banks) o i e s and c eeks. They unde s and he a o "si ing
igh ." I ha e o en gone wi hin a ew ee o hem in he Nipa Palms wi hou
no icing hem. — W. L. A.
