T
opical
Medicine,
22
(1),
13-25,
Ma ch, 1980
13
S udies
on
he
Pa asi e
Fauna
o Thailand
3.
Mi es associa ed
wi h Thai mammals
Kimi o UCHIKAWA
Depa men
o
Pa asi ology
,
Facul y
o
Medicine ,
Shinshu Uni e si y,
Ma sumo o,
390 Japan
and
Hi oshi SUZUKI
Depa men
o
Vi ology,
Ins i u e
o
T opical Medicine,
Nagasaki
Uni e si y,
Nagasaki,
852 Japan
Abs ac
:
Fo y
i e
species
o
mi es
pa asi ic
o
pho e ic
on
Thai
small
mammals we e
eco ded.
De ailed
mo phology
and
igu es
o
an
anonymous
Laelaps,
he
male
o Longo-
laelaps
wha oni
D umond
e
Bake
and
bo h
he
sexes
o he
mi e
iden i ied
as Hi s ionyssus
indosinensis
B ege o a
e
G oho skaja
we e p esen ed.
INTRODUCTION
Du ing
he
su ey
o
Thai
mammals
ca ied
ou
in
1978
and
1979,
he
junio au ho
collec ed
aca i
ound
on
all
he
ca ches.
Pa asi ic
aca i
ha e
so
a
been
s udied ex ensi ely
in
Indochina
egion,
and,
in
some
gene a
o
e en
in
some
amilies,
i is
a he di icul
o ge
new
eco ds
in
his
egion.
Collec ion
eco ds
o
any
scale
ha
always
yield some
aluable
in o ma ions
should,
howe e ,
be
p ope ly
p esen ed.
On
his
iew,
all he
pa as ic
aca i,
exclusi e
o
icks
(Ixodides)
and
ombiculids
(T ombidi o mes)
ha will
be
deal
wi h
in
o he
pape s,
collec ed in
he
su ey
a e
eco ded below.
I
is
usual
o
ind
non-pa asi ic
aca i
on
he
u
o
mammals.
Some
species a e
egula
pho e ic
mi es,
and
some
o he s
become
associa ed
wi h
mammals
qui e acciden a-
ly.
Bo h
he
g oups
o
mi es
a e
expec ed
o
be
indica i e
o
some
ex en
o en i onmen al
condi ions
o
anges
o
ca ie
mammals.
In o ma ions
o
hese
non-pa asi ic
mi es as
well
as
nidicolous
mi es
should
be
compiled
he ea e
sys ema ically,
imp o ing any
occasion.
Thus,
all
he
non-pa asi ic
mi es
will
be
also
eco ded
in
his epo .
Con ibu ion
No.
997
om
he
Ins i u e o
T opical
Medicine,
Nagasaki Uni e si y.
Recie ed
o
Publica ion,
Feb ua y
13, 1980
14
HOSTS
AND LOCALITIES
The
mammals
ha
yielded mi es
comp ized
8
species
o
4
o de s as
shown
in
Table
1.
These
hos s we e
caugh
a
Mae
Hongson,
Nako n Na-
yok
and
The
Ri e
Kuwai, Thailand,
in
Sep embe
o
Augus ,
1978, and
a
Nako n
Nayok
and
Doi In hanon,
Thailand,
in
Feb ua y, 1979.
Table
1.
Hos s
ha
yielded mi es
Chi op e a
Rouse us leschenaul i
Myo is siligo ensis
Insec i o a
Anou oso ex squamipes
Roden ia
Eo henomys melanogas e
Muspaha i
Ra us a us
Ra us sabanus
Ra us ni i en e
Ra us la escens
Ra us su i e
Mene es be dmo ei
P ima es
Tupaia glis
MITES RECORDS
The
di ec
mi e
sampling
me hod
employed
du ing
su ey
esul ed
in he collec ion
o
a
o al o
45
species
ep esen ing
25
gene a,
inclusi e
o
some
unde e mined
ones, o
19
amilies
as
p esen ed
in
Table
2.
Among
hese
mi es,
he
alidi y
o
Rad o dia (Rad-
o dia)
sp.
n .
ensi e a
(Poppe)
(Myobiidae)
is
no con i med.
Pa asi ic
Laelapidae
we e mos
abundan
on
an
insec i o e
and
oden s. Haemo-
gamasus
suncus
All ed
new
o
Thailand
was
o iginally
desc ibed
as
he pa asi e
o Suncus
(Pachyu d)
sp.
and
So iculus
spp.
on
Hymalayan
high
land,
Nepal
(All ed,
1969). The
p esen eco d
on
Doi
In hanon
sugges s
a con inuous
dis ibu ion
o
H.
suncus om he
no he n
high
land o
Hymalaya,
associa ing
wi h
a ious
insec i o es.
The
Thai hos ,
Anou oso ex
squamipes,
was
also
in es ed
wi h
he
2
mi es,
And olaelaps (Haemolaelaps)
so cinus
(Jameson)
and Placomyobia
wilsoni
Jameson,
bo h
o
which
occu
on
he same
hos
on
he
high
al i ude
o
Taiwan
(Jameson,
1966, 1970).
The
mi es
o
he
genus
Longolaelaps
a e
endemic
o
he
O ien al
egion.
The h ee
species
o
he
genus ha e
been
desc ibed
based
on
he
emales.
In
he
case
o L.
wha oni
D ummond
e Bake ,
he
o iginal
desc ip ion
was
made
on
he
8
emale speci-
mens
aken
om
Ra us
ajah
( ype
hos )
and
R.
a us
a gen i en e
in Malaya(D ummond
e
Bake ,
1960).
Then,
Dom ow
e
Nadcha am
(1963)
eco ded
162
emales
om 3
hos s
oge he
wi h
206
emales
o
L.
longula
Vi z hum
om Malaya.
Despi e
o abundan
occu ence
o
L.
wha oni
emales,
he
pa ne
male
has
no
so
a
been
ound
ou . This
is
also
case
o
L.
longula, hough
he
hi d
species,
L.
aubi
D ummond
e
Bake , is
15
k
now
om
only
a
small
numbe ,
ha
is
9,
o
he
emale.
In
he
p esen
su ey,
he 2
male
and
7
emale
specimens
o
L.
wha oni
we e
o una ely
ound
on
he
2 oden s.
The
de ailed
mo phology
o
he
i s
Longolaelaps
male
will
be
p esen ed la e .
The
mi es
o he
genus
Laelaps
we e
mos
equen ly
collec ed
om Mu idae, and
he
subgenus
Laelaps
p edomina ed
o e
he
subgenus
Echinolaelaps
in
numbe
o species
and
ice
e sa
in
numbe
o
indi iduals
as
shown
in
Table
2.
This
indica es
ha he
mode
o hos
p e e ence
is
di e ence
is
di e en
in
bo h
he subgene a.
Wi h
espec
o
he
mi es
o
he
subgenus
Laelaps,
L.
(L.)
de h ionomydis Lange
new
o
Indochina
is
sligh ly
de ia ed
mo phologically
om
ha
desc ibed by Russian
aca ologis s
in
ha ing
la ge
do sal
shield
(ca.
0.6mm),
s ou
legs
and
se ae,
and adanal
se ae
ha
a e
s ong
and
somewha
spini o m
and
ex ending
o
he basal
le el
o he
pos anal
se a.
Al hough
u he
s udy
o
bo h
he
sexes
is
necessa y,
i
is
hough
o be
easonable
o elega e he
mi e
o
L.
(L.)
cle h ionomydis.
An
anonymous
species aken
om
Mene es
be dmo ei
was
ega ded
as
Laelaps
(L.)
B
in
All ed
(1970),
which was
eco ded
om
Ra us
sp.
in
Vie
Nam.
This
mi e
seems
o
be
iden ical
wi h Laelaps
(L.)
sp.
n .
hamnomys
Tau lieb,
1954,
in
Hadi
e
al.
(1977)
ha
was
aken
om M.
be dmo ei
in
Sou h
Vie
Nam.
All ed
(1970)
no ed
as
his
"This
mi e
ep esen s an
undesc ibed
species".
The
p esen
au ho s
suppo
D .
All ed's
iew.
As
his desc ip ion
wi hou
igu e
is
b ie ,
supplimen al
mo phological
accoun s
will
be
p esen ed la e ,
expec ing
an
accomplishmen
o
D .
All ed's wo k.
O he 4
species
o he
subgenus
Echinolaelaps,
L.
(£)
aubi
Dom ow
was en-
coun e ed
only
on
he
hailand,
Doi
In hanon.
This
mi e
was
o iginally
desc ibed on
specimens
aken
on
Came on
Highlands,
Malaya
(Dom ow,
1962),
and
S and man and
Mi chell
(1963)
added
he
eco ds
om
high
al i udes in
Vie
Nam
and Thailand.
A
p e y
la ge
numbe
o
he
mi e
o
he
genus
Hi s ionyssus*
we e
aken om
Ra us
sabanus.
This
mi e
was
en a i ely
iden i ied
as
H.
indosinensis
B ege o a
e G o-
ho skaja
despi e
o
some
ema kable
di e ences
as
compa ed
wi h
he
o iginal desc ip ion
(B ege o a
e
G oho skaja,
1961).
B ege o a
and
G oho skaja
(1961)
desc ibed
H. indo-
chinensis
on
he
specimens
ound
on
R.
sabanus
ssp.?,
R.
a us
la ipec us, Suncus
mu inus
om
No h
Vie
Nam
and
on
Callosciu us
macclellandi
and C.
swinhoei o m
sou he n
China.
They
also
p esen ed
he
closely
allied
species,
H.
callosciu i B ege o a
e
G oho skaja,
as
he
pa asi e
o
Callosciu us
e y h eus
subspp.
om
he
same locali ies
in
he
same
pape .
The
mi es
o
he
genus
Hi s ionyssus
a e
no
so
igidly
hos species
speci ic,
bu
phylogene ically
and
ecologically
di e en
hos
species
a e
usually associa ed
wi h
di e en
Hi s ionyssus.
In
his
espec ,
he
hos
eco ds
p esen ed
in
he o igina]
desc ip ion
o
H.
indochinensis
as
ci ed
abo e
a e
a he
excep ional.
Al hough
he p esen
au ho
we e
no
able
o
examine
he
ype
specimens
o
H.
indochinensis,
he Hi s ionyssus
The
synonymy
o
he
geuus
Hi s ionyssus
Fonseca,
1948,
wi h he
genus Echinonyssus
Hi s ,
1925,
was
ecen ly
sugges ed
by
Teno io
e
Rado sky (1979)
16
Table
2.
Mi e
Reco ds
-
(1)
Sub-o de Mesos igma a
Family Species
Pa asi idae
Laelap idae
Gen. sp.
Hypoaspis
(Cosmolaelaps) sp.
Haemogamasus
suncus All ed
Longolaelaps
wha oni
D ummond
e Bake
And olaelaps (Haemolaelaps)
so cinus (Jameson)
Laelaps
(Laelaps)
alge icus Hi s
Laelaps
(Laelaps)
cle h ionomydis Lange
Laelaps
(Laelaps)
libe iensis Hi s
Laelaps
(Laelaps)
nu alli Hi s
Laelaps
(Laelaps)
u kes unicus Lange
Laelaps
(Laelaps)
sp.
B All ed
Laelaps
(Laelaps) sp.
Laelaps
(Echinolaps)
aingwo hae
S and mann
e Mi chell
Laelaps
(Echinolaelaps)
echidninus Be lese
Laelaps
(Echinolaelaps)
sanguisugus Vi z hum
Laelaps
(Echinolaelaps)
aubi Dom ow
Mac ochelidae
U opodidae
Spin u nicidae
Mac onyssidae
Hi s ionyssus
indosinensis
B ege o a
e G oho skaja
Mac ocheles
muscaedomes icae (Scopoli)
Gen.sp.
Me is aspis
la e alis (Kolena i)
Ancys opus
eonyc e is
Del inado
e Bake
Ancys opus
zelebo ii Kolena i
Ancys opus sp.
S ea onyssus sp.
n .
a e
Rado sky
e Y unke
17
Locali y
:
A-Doi
In hanon,
B-Nako n
Nayok
;C-Mae Hongson
Numbe
Hos
Locali y
and Da e
3J)
N
Anou oso ex
squamipes
A,
1979
à"
II
à" 22-23
19
A.
squamipes
A, 1979à"IIà"21
146
£35?
923DN1PN
A.
squamipes
A, 1979à"IIà"21-24
2a
61$
Ra us
su i e
B,
1978 à"VII.23
69
9
Mene es be dmo ei
C,
1978
à"VII
à"7
129
94DN1PN
A.
squamipes
A, 1979à"IIà"21-23
1
£
Ra us a us
3
9
9
Eo henomys
melanogas e
A,
1979
à"
II
à" 22
2$
$
Muspaha i
A, 1979à"IIà"20
1PN
*.
a ws
C,
1978
à"VIII
à"1
1
$3
9
$
a ws lu escens
B,
1978
à"VII
à"23
18
M.
be dmo ei
C, 1978-VIII-7
8
5
$
a ws lu escens
A,
1979
-
ll
- 23
35
5
A .
be dmo ei
C, 1978-VIIà"4
!
$
#am«
ni i en e
A,
1979
-
ll
- 22
7a
a3PN
R.
su i e
B, 1978-VII-23
4$
$69
$
M.
be dmo ei
C, 1978-VIIà"7
10S
S10$?
R.
su i e
B, 1979-II-10
26
£35
$
K.
l us
B, 1979-ll-10
Ia9$
9
U.
5« (/e
A, 1979à"IIà"24
6$
9
R.
a us
B, 1979à"II-10
29 9
M.
be dmo ei
C, 1978-VIII-7
2662$$1DN
K-
*««>
B, 1978-VII-23
499
R-
á" us
B' ^79-H-lO
28
$2?
9
A .
be dmo ei
B, 1979à"IIà"10
36 68$ 9
*.
wi i/ e
B> 1979*11-10
3aS1699
R.
wi e
A, 1979-ll-24
139
$
Ra us ni i en e
A,
1979
-
II
- 21-22
89
91DN
R.
la escens
A,
1979à"II à"23
1$
E.
melanogas e A,
1979
-
II
' 23
118
S39
§
K.
5 l&a»«5
B, 1978-VII-23
29
9
A.
sgwaim/HW
A, 1979-ll-22-23
1DN
£.
melanogas e
A,
1979
à"
II
à" 21
186
624$
$21DN
Rouse us
leschenaul i
C,
1978
à"
VII
à"31
19
R.
leschenaul i
C,
1978
-VII
- 31
23
$2DN
#,
leschenaul i
C,
1978 à"VII-31
!9
£.
leschenaul i
C,
1978
-VII
- 31
1
9
7PN
A yo w
siliga ensis
B
,
1978
à"
VIII
à"4
18
(Con inued)
mi e
Reco ds
-
(2)
Sub-o de T ombidi o mes
Locali y
:
A-Doi
In hanon
;
B-Nako n
Nayok
;
C-Mae Hongson
;
D-The
Ri e Kuwai
Family
Species
Numbe
Hos
Locali y
and Da e
Cheyle idae
Cheyle us
e udi us
(Sch ank)
1 $
M.
be dmo ei
C,
1978
à"VIII
- 7
Cheyle iellidae
Nihelia
quin a
Dom ow
e
Bake
59?
Tupiaglis
Q, 1978à"VIIà"19
Myobiidae
Placomyobia
wilsoni
Jameson
26
62$
$
A.
squamipes
A, 1979-ll-24
Rad o dia
(Rad o dia}
ensi e a
1$2$
$
R.
a us
C, 1978à"VIII-2
Rad o dia:
(Rad o dia)
sp.
n .
1
$
J?.
5a&aw«5
B,
1978
-
VII
- 23
ensi e a (Poppe)
Pygmepho idae
Pygmepho us
sp.
n .
e cipa us
1
$
4.
squamipes
A,
1979
-
II
- 23
Willmann
Rack a
sp.
1$
A.
squamipes
A, 1979à"II*23
Bake dania
sp.
1
19
A.
squamipes
A, 1979à"IIà"23
Bake dania
sp.
2
1$
4.
squamipes
A, 1979à"IIà"24
(Con inued)
Mi e
Reco ds
-
(3)
Sub-o de
Sa cop i o mes
(no enume a ed)
Locali y
:
A-D0i In hanon
;
B-Nako n
Nayok
;
C-Mae
Hongson
;
D-The
Ri e Kuwai
Family
Speci
Hos
Locali y
and Da e
Sa cop idae
Nyc e idocop es
asia icus
Fain
R.
leschenaul i
C,
1978
à"
VII
- 31
A opomelidae
Lis opho oides
(Me aquesania)
cuculla us
R.
a us
B
1979
à"
XI
à"10
(T ouessa )
' u
Lis opho idae
A olis opho us
sp.
n .
maculo us
a us
Fain
R. sabanus
B,
1978
à"
VII
à"23
A olis opho us
sp.
(nymph)
R.
a us
B
,
1979
à"XI
à"10
A olis opho us
sp.
n .
mediolinea us
Fain
M.
be dmo ei
C,
1978
à"VII
à"4
Sciu ochi us
ailandiae
Fain
£.
ni i en e
A,
1979
à"
II
à" 22
( ans e ?)
Lynxaca us
upaiae
Fain
T.
#/«
B,
1978
à"
VII
à"23
Aca idae
Psylloglyphus
ie namensis
Fain
e
Beaucou n R.
a us
B,
1979
à"
II
à" 10
R.
su i e
B, 1979-ll-10
M.
be dmo ei
C, 1978à"VIIà"4
A.
squamipes A, 1979à"IIà"23
Glycyphagidae
De maca us
sp.
n .
no aequineae
Pain
#.
ni i en e
A,
1979
à"
II
à"22
Alabidopus
bipili e
Fain
e
Uchikawa
4.
squamipes
A,
1979
à"
II
à"23
Tupaiopus
hailandicus
Fain
e
Uchkawa
£.
sabanus
B
,
1978
à"
VII
à"23
?"à"
£/M
B, 1978à"VIIà"23
.4.
squamipes
A, 1979à"IIà"23
( ans e ?)
(O iba endae
Suc obella
sp'
*à"
^*W
C,
1978
à"VIII
à"1
19
mi e
collec ed
in
he
p esen
su ey
was
iden i ied
as
H.
indochinensis
on mo phological
simila i y
and
hos
eco d.
A
de ailed
mo phology
o
he
p sen
mi e
will
be desc ibed
and
igu ed below.
Mac ocheles
muscaedomes icae
(Scopoli)
is
a
common
p edacious
mi e
pho e ic on
he
lies.
I
was
in e es ing
o
ind
he
mi e
on
he
insec i o e,
which
is
easy
o ge
composed
and
o
become
a ac an
o
lies
sho ly
a e dea h.
The
mos
ex ensi e
s udy
o
Spin u nicidae
pa asi ic
on
Thai
ba s
was
made by
D .
N.
Wilson
in
Hill
and
McNeely
(1975).
He
eco ded
he
ou
species,
inclusi e o
an
anonymous
one,
o
Ancys opus
om
Rouse us
leschenaul i.
The
p esen Ancys opus
ha
is
close o
A.
eonyc e is De inado
e
Bake
and
D .
Wilson's
anonymous Ancys opus
a e
possibly conspeci ic.
A
S ea onyssus mi e
was
newly
ound
on
Myo is
siliga ensis.
The mi e
was
e y close
o
S.
a e
Rado sky
e
Yunke ,
bu
4
pai s
o
minu e
pos e io
se ae
on
he spis hosomal
shield
o
he
emale
di e
in
size in
he
p esen
mi e
and 5.
a e .
The
mi e
is hough
o
belong
o
a
dis inc i e
axon,
bu
he
desc ip ion
on
bo h
he
sexes
is disi able.
Nine
species
o
T ombidi o mes
we e
aken.
Nihelia
quin a
Dom ow
e Bake ,
which has
been
eco ded
om
T.
glis
on
Malaya
(Dom ow
and
Nadcha am,
1963) in
succession
o
he
o iginal
desc ip ion
(Dom ow
and
Bake ,
1963),
seemed
o
be common
on
he
hos ,
bu
male
is
s ill
unknown.
Al hough
Smiley
(1970)
elega ed
he
mi e in
he
genus
C ieke on
Volgin,
u he
s udies
o
he
genus
o
he
mi e
a e
necessa y o
com
i m
his
p oposal.
Placomyobia
wilsoni
Jameson
was
o iginally
desc ibed
om Taiwan
as
he
pa asi e
o
Anou oso ex
(Jameson,
1970).
Bo h
he
myobiid
genus, Placomyobia,
and
i s
hos
genus,
Anou oso ex,
a e
mono ypic,
sugges ing
a
phylogene ic dis inc i eness
o
he
sh ew-mole.
As
o
a
single,
damaged
emale
o
Rad o dia
aken
om
R. sabanus,
iden i ica ion
was
es ained
because
o
i s
la ge
size
and
sligh ly
di e en
se al na u e,
hough
he
mi e
was
e y
close
o
Rad o dia
(JR.)
ensi e a (Poppe).
Wi h
espec
o
he
12
mi es
o
he
sub-o de
Sa cop i o mes,
only Nyc e idocop es
asia icus
Fain
and
Tupaiopus
hailandicus
Fain
e
Uchikawa
a e
picked
up.
The o me
mi e
has
been
known
om
he
male
and
male
i onymph
(Fain,
1959).
Female
o his
mi e
is
s ill
unknown,
hough
some
males
and
a
conside able
numbe
o
imma u e s ages
we e
aken
in
he
p esen
su ey.
Tupaiopus
hailandicus
Fain
e
Uchikawa
was eco ded
om
T.
glis
and
A.
squamipes,
bu
he
la e
hos
eco d
seemed
o
be
dubious. A
single
mi e
was
p obably
mislabelled
by
he
senio au ho .
DETAILED MORPHOLOGY
1.
Longolaelaps
wha oni
D ummond
e
Bake ,
I960,
male
(Fig. 1.)
Idiosoma
420-415^
long by
230-225^
wide. Do sal
shield
sligh ly
conca e la e ally,
420-410^
long
by
200-200^
wide
a
na owes
poin
o e
coxa
IV,
s ia ed
dis inc ly and
20
bea ing
39
pai s
o
se ae;
17
pai s
o
elonga e
and
swollen se ae
on
cen al
pa
and he
o he
ma ginal
se ae,
bu
D$,
which
a e
conspicuous,
being
minu e
and
spini o m. Abou
7
pai s
o
minu e
se ae
on
do sal
so
in egumen .
Holo en al
shield
expanded mode a ely
behind
coxae
IV, bea ing
10
pai s
o
p ominen
se ae
and
minu e
adanal
and pos anal
se ae.
Abou
10
pai s
o
minu e
se ae
en ally
on
so
in egumen .
Pei eme ex ending
o e
midpoin
o
coxa
II.
Legs
as
illus a ed
in
Fig.
1-1
and
2
;
leg
se ae
a he sho .
Deu os e num
wi h
8
ows
o
3
ee h.
Chelae
as
igu ed;
ixed
chela
weak
and memb a-
nous,
p obably
wi h
e y
minu e
pilus
den ilis;
spe madac yl
50-50^
long; a h odial
ilamen s p ominen .
The
abo e
desc ip ion
was
based
on
he
wo
specimens
in
Table 2.
2.
Laelaps
sp.
B
All ed,
1970,
emale
(Fig. 2.)
Idiosoma
650//
long by
430j«
wide.
Do sal
shield
650jw
long
by
430^
wide, bea ing
39
hai s
o
sho
se ae
;
se ae
7*2
and Zs
being
longes
and
cen al
se ae
sho es . Abou
10
pai s
o
se ae
ma ginally
on
do sum
o
he
shield.
S e nal
shield
95-100/
long by
135-150/*
wide
a
le el
o
s z
s i
70-78/*
long.
Geni o- en al
shield
weakly expanded
Fig.
1.
Longolaelaps
wha oni
P umond
e
Bake ,
1960, male.
1-do sum; 2- en e .
21
Fig.
2.
Laelaps
(Laelaps)
sp.
B
All ed,
1970, emale.
1-do sal
shield
; 2- en e .
be ween
he
second
and
hi d
se ae,
h- h.
Dis ance
be ween
se ae
gi,
i s
pai
o se ae
on
geni o- en al
shield,
58-63ju
;
k- h,
o h
se ae
on
he
shield,
73-80^
Anal shield
conca e
an e io ly,
83-88jU
long
om
an e io
ma gin
o
base
o
pos anal
se a
and 103-103^
wide.
Me apodal
shield
elonga e.
Abou
10
pai s
o
se ae
on
en al
in egumen . Pe i eme
e mina ing
o e
an e o
one
hi d
o
coxa
I.
Legs
s ou
;
coxa
I
wi h
ine
ex e nal and
hickened
in e nal
se ae
;
an e io
se ae
on
coxae
II
and
III
hickened
;
pos e io
se a on
coxa
III
spini o m.
Fi e,
6
and
6
hickened
se ae
on
a si
II,
III
and
IV, espec i ely.
Chaelae
minu e,
abou
28 i long.
The
abo e
desc ip ion
was
made
on
a
whole
moun
and
2
dissec ed
specimens, all
o
which
we e
gi en
in
Table 2.
3.
Hi s ionyssus
indochinensis
B ege o a
e
G oho skaja, 1961
Female
(Fig.
3).
Idiosoma
480-530//
long
by
330-380/;
wide.
Do sal
shield 425-450
long
245-255^
wide,
weakly
s ia ed
ma ginally
and
g anula ed,
be ing
26
pai s
o minu e
se ae.
Se en een
pai s
o
se ae
ma ginally
on
so
in egumen . T i os e num
wi h well
spinose
laciniae.
P es e nal
sculp u e
weak.
S e nal
shield
45-50^ long
and
110-115/ wide
