Doñana, Ac a
Ve eb a a,
2 (2):
145-160,
1975
!he
s ue u e o a
Medi e aaean
U.a d
eommuni y
Cl)
J.
MELLADO, F. AMORES,
F. F.
PARREÑO
AND
F. HIRALDO
145
The
s uc u e
o liza d communi ies
has
ecen ly been
he
objec
oí
nu-
me ou$ s udies in which cha ac e is ic
p ope ies
o
animal
communi ies,
such
as
sha ing
o esou ces, niche
b ead h
and
o e lap, '
as
well
as
s a e-
gies u ilized by he di e en species
o
educe compe i ion, ha e be
en
ana-
lyzed.
Mos o hese s udies (Rand,
1964;
Schoene ,
1968;
Schoene
and
Go
man
,
1968;
Hillman,
1969;
Pianka,
196~a;
Schoene ,
1970;
Huey
e
al.,
1974:)
ha o
been
made
on
g oups o
sympa ic
oongene ic speci'es, whe eas
Pianka
(1967;
1969b;
1971;
1973)
and
mo e
ecen ly Sage
(1974)
analyzed en i e communi-
ías o liza ds.
He e
We
analyze
he
s uc u e
o a liza d communi y ep esen a i e o
ha
Eu opean
Medi e anean
zone,
an
a ea
as
ye
uns udied
excep
o
a
p &-
cu so y
in es iga ion
ca ied
ou
by
Val e de
(967).
We also
compa e
his
communi y
o
hose desc ibed
by
Hu ubia
and
Di
Cas i
(1973)
and
Saga
(1973)
o
he
Medi e anean zones
o
he
New Wo ld.
Ma e ial
and
Me hods
610 specimens belonging ' o
eigh
specíes o
liza ds
we e
collec ed
a
i egu-
la
in e als
hl'ougholl a
pe iod
oi
35
mon hs
(Oc obe1'
1971
o
Augus
1974).
These
samples
we e
cap u ed
p incipally
by
shoo lng
01'
by
hand
and
p ese cd
in
70
pe
cen
Alcohol,
excep
iol'
mos
Lace- a. lepid4
whicb
we e
dona ed
by
he
Ownel'S
o
p í a e
es a es
whe e
hey
a e
usually
killed, being conside ed
as
ha m-
ul
o
hun ing;
hese
samples
we e
p ese ed
in
10
pe
cen
o ma
Un.
Tlle
buck
o
all
ma e ial
used he1'e
has
been
dona ed
o
he
Biological
S a-
ion o Doñana, Se illa, Spain.
(1):
This papel' was o iginally deli e ed
in
a Symposillm
on
he
«Ecosys em
s uc u e
and
s abili y"
in
No embe
1974
a
Se illa, Spain.
146
J.
MELLADO,
F.
AMORES,
F. F.
PARREÑO
AND
F.
HIRALDO
Time
be ween
cap u e
and
analysis
a ied
g ea ly,
om
immedia ely
a e -
wa ds
ill
se e al
mon hs
la e o
Samples
collec ed
p o ided
he
ollowing
mo phological
da a
(measu ed
on
he
p ese ed
animaD
:
SVL
(snou
en
leng h)
:
Measu ed
om
he
end
o
he
snou
o
he
dis-
al
edge
o
he
anal
pla eo
ea
.
HL
(
Head
lengh
):
F om
he
end
o
he
snou
o
he
an e io
bo de
o
he
HW
(Head
wid hl:
Maximu n
wid h
o
he
head
.
W (Weigh ):
Only
in
esh
specimens.
Al!
neasu emen s
a e
in
millime e s,
weigh s
in
g ams
.
Only
s o
m
ac
h
con en s
we e
conside ed
in
he
s udy
o
ood.
P ey
i ems
we e
coun
e
d Ol"
each
s om
a
ch
and
clasi ied
in o
21
ood c
a eg
o
i
es
lis ed
in
able
II. The l
eng
h
o
in ac
p ey
was me
asu ed
o
he
ne
a
es
.1
o a
milli-
me
e
l·.
Fi
e
ld
ob
se
a
íon
s
we e
m
ade
a
i egula
in
e
als,
a
leas
once
a mon h
(exc
ep
Sep e
mh
e ),
a
nd
du l
ng
a
minimum
p
e l
od o
i e
days
ea
ch m
on
h,
a
di
e e
n
ime
s o day
an
d nigh
.
W
hen
poss
ibl
e, he
ollo
win
g d
a a
we e
no
ed
in
ie
ld obs
e
a i
o
ns
:
species
,
ap o
x
íma e
size a
nd
age
,
ime
,
locall y
and
cx
ac
loca ion
as we
ll
as
ac i i y
wh
en
i s
sigh ed.
Sludy
A es
Field
obse a ionsand
ma e ial
employed
in
his
s udy
come
om
an
ex ensi e
a ea
si ua ed
a
he
mos
wes e n
pa
o
he
Sie a
Mo ena
moun ain
ange
in
he
P o ince
o
Huel a
(S.
W.
Spa n,
ap oxima ely
38°
N
by
7°
W).
This
zone
comp ises
lines
o
moun ainous
coun y
o
oeunded
p o-
ile,
a
low
al i ude
(a e age ele a
ion
o
500
me e s
abo e
sea
le eD, 'which
al e na es
wi h
wide
alleys
and
peniplanes
( o
a
de ailed
desc ip ion
o
his
zone, see
Lau ensach,
1967).
The
clima e
is o
he
ype
«Mesomedi e--
mnéen
accen ué»
as
desc ibed
by
Embe ge
e
al. (1963)
wi h
75
o
100
' bio-
logi'cally
d y
days. Annual
p ecipa ion
a ies
om
abou
800
o
1,300
mm.
o
ain
which
alls
mainly
om
No embe
o
Ma ch
.
Tempe a u es
below ze o,
a e
in equen ,
and
he
o al
numbe
o
hou s
pe
yea
bel
ow ze o
does
no
e
a
ch
he
h ee
pe
cen
indica ed
by
Asch-
mann
(1973) as a
limi
o
Medi e
I'
anean
clima es.
F os
ls
sca ce
and
snow
excep ional
on
his
land,
as
any
snow
all
usually
mel s
immedia ely.
WHh
ega d
o
ege a ion,
h
is o iginally co
nsis ed
in
a
dense
e e g een
scle ophylous
o es
o
Que cus
ilex
and/o
Q.
sube
wi h
so ne
isola ed
s ems
o
Q.
lusi anica
in
dampe
a eas
.
This
Q. es
possessed
a
dense
unde
.
b ush
made
up
chie ly
o
Ulex spp.
and
se e al
species
o
Cis us
as
well
as
A bu us
unedo.
In
he
b ooks
and
go ges
galle ies
o
Alnus
glu inosa
and
F axinus
sp.
occu
wi h
hick
bushes
o
Rubus
and
Ne ium
oleande .
This
ege a ion
has
been
adically
ans o med
by
he
hand
o
man
in
wide
ZQ_
Dañan
:>.,
Ac a
Ve eb a a,
2 (2),
1975
The
$ uc u e
01
a
Medi e anean
liza d
communi y
147
nes,
and
due
o
degene a ion o
he
o .es , wo
main
ypes o
habi a
ha e
appea ed:
Chapa al.-This
consis s basically
o
Cís us
ladanije us,
E ica
aus al s
and
E.
umbella a,
oge be wi b
Genis a
sp., A.
unedo
and
E.
a bo ea
appea-
ing
in
damp
a eas. This
unde b ush
o ms
a e y dense con inuous blan·
ke
oí
up
o
ou
me e s
bigh
in
some places, and gene ally occupie·s high
a eas
o
moun ain
anges.
E e g een
oak
wood.-A
hin
o es
o
Q.
ilex
and/o
Q.
sube
in
which
be
soíl
has
been comple ely clea ed o
unde b ush.
This
habi a
occupies
he
alleys
and
plains. Ex ensi e
pa k~
oí
Cas anea
sa i a
oí
simila
s uc·
u e
a-ppea
in
he
coole
and
dampe
a eas.
T o hese
mus
be
added
a e y peculia o ma ion consis ing
oí
ou ·
c ops
o
g ani ic
ocks de e mining ue so·called «Plu onic bu onsll (see
Lau ensach,
1967).
These
a- e
ex emely ocky
a e
as
wi h some
s ems
o ooks ,
and
unde b ush
o
Pis ada
sp
.,
Mi us
sp
.
and
Cis us
.
ResuU.
A
maximum
o eigh liza d species
can
co·exis
in
he
egian
s udied.
These
a e
(2):
Tm
Ta en ola
m.
mau i aniea
Be
Blanus
e.
cine eus
Lh
Lace a
hispanica
auche i
Ll
Lace a
l.
lepida
Pa
Psammod omus
a.
algi us
Ph
Psammod omus
h.
hispanicus
Cb Chalcides
bed iagai
pis aciae
Ce Chalcides chalcides
s ia us
Gekkonidae
Amphisbenidae
Lace ida.e
))
))
))
))
))
))
Scincidae
» »
In
he
es
o
his pape ,
he
symbols p eceding each liza d
name
lis ed
aboye will be used.
Table 1 shows
he
main
biome ic
cha ac e is ics
o
he
di e en spe-
cies,
and
igu e 1
p esen s
he
dis ibu ion
o
snou ' en
leng hs (SVL). Th oo
pai s
o congene ic speci'es occu
in
sympa y; hese di e
om
one ano
he
wio hin
each
pai ,
in
aH
measu emen s
aken
in o
conside a ion.
(2):
Following
Me e ns
and
We mu h,
1960,
excep
o
Chalcides
bed iagai
(see
Val e de, 1966).
Doñana,
A.
c a
Ve eb a a,
2 (2),
1975
148
J.
MELLADO, F.
AMORES,
F. F.
PARREÑO
AND F.
HIRALDO
All
hese
di e ences
a e
s a is ica11y
signi ican
(P<
.05; - es ) excep
be ween
he
wo
species
o
he
genus
Chalcides
whe e
none
o
he
di e en-
ces
we e
s aUs ica11y signi ican .
Table 1
Mo phological s a is cs used
in
bis s udy
SVL W
HW
HL
m
c
n m
c
n m
c
n m
c
n
Tm
54·04
14.1
21
6.6
4.4
15
9.66
2.4
18
15.0
3.6
18
Be
160.05
30.3
37
4.6
1·7
15
4.82
1.9
28
7.8
Vi
28
Lh
46.U3
4.7
131
2.0
1.0
45
5.87
1.0
56
10.3
1.8
.'l9
Ll
138.46
53.7
167
71.7
8.4
35
15.9
9.8
73
32.8
4.3
74
Pa
58.88
14.6
141
7.7
2.4
65
10.6
1.9
39
15.9
2.2
39
Ph
40.05
5.S
57
1.1
1.1
42
5.54
4.0
34
9.8
1.4
35
Cb
66.14
10.6
21
4.4
1.2
11
6.71
1.8
21
8·0
.9
21
Ce
95.08
31.0
22
4.9
3·8
21
5.90
1.2
22
8.5
1.7
19
SVL
= Snou - en leng h;
HW
= Head wid h; HL = Head leng h;
W = Weigh .
Food
A
o al
o
511
s omachs
we e
analyzed.
Table
2
summa izes
he
main
ea u es
o
he
die
o
he
eigh
liza d species
in eg a ed
in
he
communi y.
Mos
liza d
species
a e
ecclec ic
p eda o s,
eeding
on
a
wide
~ange
o
in e eb a e
p ey
(see e.g. Pianka, 1973,
and
included
e e ences).
In
e-
quency
o
appea ance
o
p ey
üems
in
s omachs
analyzed
and
equency
oí
each
p ey
i em
in
he
o al,
he
mos
abW1dan
g oups
in
he
die
we e bee-
Ies,
spide s
and
.insec
18i ae,amonga
wide
a ie y
o
g· oups
o
in e e-
b a es.
Seeds
8ind
e eb a e
p ey
a e
also included
in
he
die
o
se e al
species.
Table 3
shows
he
p ey
size
dis ibu ion
and
he
mean
p ey
size
o
each
liza d
species. Liza d size
i:;
gene ally
co ela ed
wi h
ha
o
he
p ey
inges ed.
In
his
case,
mean
p ey
size
was
col1 ela ed
bo h
wi h
snou - en
leng h
and
he8id dimensions o
he
liza d
o
8111
species.
O
hese
compa í-
sons,
only
mean
p ey
size s.
mean
head
wid h
was
s a is ically
signi ican
( :
.720;
P<
.05;
n:
8).
Doiíana,
Ae a
Ve eb a a, 2 (2),
1975
The
s uc u e
01
a Medi eTTanean
liza a
communl 'll
1.9
•
Be
Ll
Ce
-
Cb
~
Pa
Tm
Lh
40
Ph
o
20
100
,
.o
.
180
220
mm.
60
Fig.
1.
F equency
dis ibu ions
o
snou - en
leng hs
(millime e s)
lo
he
eigh
liza d
species comp'lsing
he
communi y
.
Doñana, Ac a
Ve eb a a,
2 (2),
1975

150
J.
MELLADO,
F.
AMORES,
F. F.
PARREÑO
AND
F.
HIRALDO
T ophic
di e si y
was
calcula ed
by
Pielou's
me hod
(1966),
using
he
B iUouin
o mula:
H=(1/N)
(logIa Ni! -I
loglO
Ni!)
This
me hod
was
o me ly
applied
o
he
analysis
o
die
o
p eda o s
by
Hu ubia
(1973).
Plo ing
accumula ed
ophic
di
'
e si y
(Hk)
s.
o al
numbe
o
s omachs
(k),
di e si y
cu es
a e
ob ained
(Figu e
2).
These
cu o
es
a e
highe
o
la ge
species,
in
pai s
o
congene s.
Duo
o
sample
he e ogenei y,
including
s omachs
om
e y
di e en
pe-
iods,
locali ies
and
habi a s,
i
is
di icul
o
de e mine
he
s abiliza ion
poin
( )
o
he
cu es,
and
so
he
di e si y
o
a11
he
s omachs
(Hz)
o
each
Table
2
Pe cen age
compm:i ion,
by
numbe s,
o
he
die
o
Uza d
species
composing
he
communi y
The
numbe
oC
s omachs
and
p ey
i ems
on
which
hese
pe cell ages
a e
based,
as
well
as
he
a e age
ophic
di e si y
(Hz),
a e
al
so
shown.
Oligoehae a
Isopoda
Gas e opoda
Miliapoda
A ane
ae
halangidae
Seo pionidae
O hop el'a
De map e a
'BJa la ia
Hemyp e a
Fo micidae
O he
hymenop e a
Lepidop c a
Dip e a
Coleople a
Lepidop e a
Ia ae
O he
Ia ae
Seeds
Ve eb a e
i ems
lTniden i ied
P eys
i ems
S omaehs
T ophic
di e si y
(Hz)
Tm
2.85
2.
85
20.0
1.42
10.0
7.14
1.42
8.57
1.42
10.0
17.13
8.57
4.28
2.85
70
18
.975
Be
4.9
.98
2.94
12.74
.98
Lh
3.24
.36
1.79
24.46
1.08
2·16
.72
1.08
2.16
Ll
.47
1.12
.42
1.26
0.5
·08
1.77
.28
.56
16.66 9.35 1.85
3.95 2,4
.03
7.
19
.
98
9.80 20.87 68.42
l7.64
9.71
5.0
28.40 6.47 9.36
2.94
103
23
.760
1.5
.36 .25
5.09 .90
268
2919
9S
158
1.059 1.087
Doñana,
Ac a
Ve eb a a,
2 (2), 1975
Pa
Ph
Cb
l.19 9.4 22.5
.39
2.5
1.
38
2.3 5.0
7,14 24.8 20.0
.39
8.53
.19
.19
1.98
12.60
10.11
2.
97
38.07
8.92
3.76
1.1
.5
15.3
1.1
2.3
23.9
5.9
8.8
7.5
35.0
7.5
Ce
10.3
2·06
18.55
6.18
8.25
3.10
1.03
4.12
24.73
10.jO
7.21
3.10
.19
1.78
504
3.5
169
48
40
79
132
1.225 .852
15
22
.707 .960
The
s uc u e
01
a
Medi e anean
liza d
communi y
151
species,
is
used
as
indica o
o
ophic
di e si y; alues
ob ained
appea
in
able
II.
A
s a is ically
signi ican cOI1 ela ion exis s
be ween
ophic
di e -
si y
and
head
wid h
o
he
liza d
species
( s:
.643;
P<.05;
n:
8.
Spea man
ank
co ela ion
coe .icien . Siegel,
1956).
Finally,
wi h
espec
o
hun ing
s a egy,
Pianka
0966;
see
also
1973)
Table
3
T ophic
esou ce
ma ix.
Rows
ep esen s
p ey
size
i ems
a
5
mm.
in e alo
To al
numbe
o
p ey
measu ed
(N),
and
mean
p ey
size
(MPS)
a e
also
shown.
Tm
Be
Lh
Ll
Pa
Ph
Cb Ce
O-
S 9
15
64
10
39
60
11
16
6-
10
23
47
51
660
63
54
15
28
11-
15
7 2
25
389
34 9 2
15
16-
20 3 4 7
255
33
2 4
21-
25
1 4
55
7 3
26-
30
~
5 1 66
11
31-
35
5
23
36-
40
1 10 4
41-
45
1 1 3
46-
50 4
51-
55
1
56
· 60 1
61-
65
3
66-
70 2
71-
75
1
76-
80
1
81-
85
86-
90
91·
95
1
96-100
2
N
46
76
158
1486
197
125
30
68
MPS
10.06 11.35 8.72 13.72 12.9 6.0 7.16 9.72
-de ines
wo
basic
ones:
«si -and-wai »
and
«ac i e o aging».
In
ou
case
only
one
species
(Tm)
is a «si -and-wai )), while
he
emainde
seem
o
be
<mc i e o aging»
hun e s.
..
¡
Doñana,
Ae a
Ve eb a a,
2
(2),
1975
152
J.
MELLADO,
F.
AMORES, F.
F.
PARREÑO
AND
F.
HIRALDO
Ac i i y
The
p esence
oí
a well-de ined wi . e
la eney
has
been
a ibu ed
as
a
undamen al
cha ae e is ie
o
he
auna
o
ep iles
o
he
Tempe a e
zone
(Sain -Gi ons, 1956). Sain -Gi ons (1953)
a ibu es
wo
diapauses
o
he
Me'
di e anean
zones,
he
one
in
win e
and
a
summe
la ency,
due
o
he
ex·
Hk
11
.9
7 ,
,
.5
..-
.
I
-'
/
..
....
'"
........
',
.....
Ce
Cb
Ll
Pa
Ph
.3
.1
o 5
10
15
5
10
15
5
15
20 K
Fig.
2.
Accumula e
ophic
di e si y
(Hk)
cu es
s
.
o al
numb
o
s omachs
(k).
Compa ison
be ween
congene ic
pai s
oi
species.
eesi e
d yness.
Da a
o e ed
by
ou sel es
in
his
espee ,
al hough
ineo n-
pIe e
o
so ne species,
do
howe e ,
pe ni
us
o
d aw
a en ion
o
wo
ypes
oí
beha iou .
Tablc
4
Spa ial esou ce ma ix. Symbols heading he able Indica es mic ohabUa ypea
d awn in de ail in he
ex o
A B e
2 3 1 2 3
Tm
o
22
o o o o o
Be
o
20
o o
17
o
Lh ()
40
o o 6 o o
Ll
27
24
o o 3 o o
Pa o o o
141
3 o o
Ph
32
o o o o o
(j
Cb
o o o 1 o
21
o
Co
o o o o o o
22
Doñana, Ac a
Ve eb a a,
2 (2),
1975
The
s uc u e
01
a
Medi e anean
liza d
communi ll
153
A
leas
wo
species
(Ll
and
Pa)
show
a
ma ked
diapause,
wi h
an
inac-
i e
pe iod
o
some
ou
mon hs
(No embe
o
Feb ua y),
al hough
he
smaH-
e
membe s
o
hese
species
a e
ac i e
h oughou
he
Whole yea .
In
o he
species
(Tm
and
Lh),
bo h
adul s
and
young
a e
ac i e
aH
yea .
As
a
as
dai1y
ac i i y
is
concemed,
only
one
species
(Tm)
is
noc u -
nal, while
he
emainde
a e
essen ially
diu nal.
Howe e ,
his
pa e n
oí
ac i i y
is
subjec
o
app eciable
changes
h oughou
he
yea ,
wi h
he
noc-
u Il8l
species
becoming
diu nal
du ing
win e
mon hs
,
whe eas
some
diu -
na!
species
(Ll
and
Pa)
ha e
o en
been
obse ed
a
nigh
du ing
ho e
mon hs
(Mellado
and
Hi aldo,
unpublished).
Mie ohabi a
p ele enee
AH
place
loca ions
obseJ' ed
o
liza ds
when
i s
sigh ed
we e
classi ied
in o
se en
weH-di e en ia ed
ca ego ies
o
de ine
he
spa ial
eS<Ju ce sha.-
ing
o
he
communi y.
We
conside
only
obse a ions
e e ing
o
eeding
beha iou
o
he mo egula o y
ac i i ies.
These
ca ego ies
include:
A
Zones
o
sca ce
o
no
plan
co e .
B
Zones
o
dense
plan
co e
.
Wi hin
each
o
he
abo e-inen ioned
ypes
he
oHowing
di isions
a e
made:
1
On
he
g ound.
2
On
ocks,
ees
and
buildings.
3
Unde
s ones,
ee
unks,
e c.
In
adi ion
o
hese
ca ego ies,
ano he
one
mus
be
added
.
I
is:
e
In
small
hick
pas u es
in
alleys
and
pJains.
One
o
he
species
(Cc)
is
es ic ed
o
C;
mo phologically
i
is
a snake-
like
liza d
which
U e ally «swimsll
in
he
pas u es
i
inhabi s
.
The
es
ol
he
species
a e
limi ed
o
a
special
kind
o
subs a um:
h ee
species
(Ll,
Pa
and
Ph)
li e
on
he
g ound
su ace,
one
o
hem
(Pa)
in
places
we e
he e
íS
a
hick
unde b ush,
whe eas
he
o he
wo
(Ll
and
Ph)
a e
ound
on
open
g ound
(Ll
needs
aes
and
ocks
o
p o ec ion).
Be
is
a
ypical
sub e a-
nean
species,
mo phologicallya
wo m-like
liza d.
Cb
li es
unde
s ones
in
hls
egion. Finally,
Lh
and
Tm
li e
on
ocks
and
l ee
unks.
Doñana,
Ac a
Ve eb a a,
2 (2),
1975
160
J.
MELLADO,
F.
AMORES,
F.
F.
PARREÑO
AND
F.
HIRALDO
I.&VINS,
R.
(1968):
E olu ion
in
changing
en iTonmen s.
P ince on
Uni .
P ess:
120
pp.
MERTENS,
R,
and
WERMUTII,
H. (1960):
Die
Amphibien
uoo
Rep ilien
EUTOpa&
Ve lag
Waldema
K ame ,
F ank u
a.
M.
PIANKA, E. R (1966):
Con exi y,
dese
liza os
and
spa ial
he e ogenei y.
Ecolog¡¡
47: 1.055-1.059.
- - (1967):
On
liza d
species
di e si y:
No h
Ame ican
la land
dese s.
Eco·
logy
48:
333-351.
- - (1969a):
Habi a
speci ici y,
specia ion
and
species
densi y
in
Aus alian
de-
se
liza ds.
Ecology
50:
498-502.
~
-(1969b):
Sympa
y
o
dese
liza ds
(C eno us)
in
Wes e n
Aus alia.
Eco-
logy
50:
1.012-1.020.
- - (1971):
Liza d
spacies
densi y
in
he
Kalaha i
dese o
Ecology
52: 1.024-1.029.
- - (1973):
The
s uc u e
o
liza d
co nmuni ies.
Ann.
Re . Ecol. Sys .
4:
53-74-
- - (1974):
Niche
o e lap
and
di use
compe i ion.
P oc.
Na .
Acad.
Sci.
USA
71: 2.141-2.145.
PIELOU,
E.
C.
(1966):
The
measu emen
01
di e si y
in
di e en
ypes
o
biolo-
gical
collec ions.
J.
Theo e .
Biol.
13: 131-144-
RAND,
A.
S.
(1964): Elcological
dis ibu ion
in
anoline
liza ds
o
Pue o
Rico.
Eco-
logy
45: 745-752.
SAGE,
R D. (1973):
Ecological
con e gence
01
he
liza d
aunas
o
he
chapa al
communi ies
in
Chile
and
Cali o nia,
p.
339-348.
In:
Medi e an!lan
ype
ecosys em.
F.
Di
Cas i
and
H. A.
Mooney
eds.,
Sp inge -Ve lag,
Be lin-
Heildelbe g-New
Yo k.
- - (1974):
The
es uc u e
o
liza d
1auna :
compa a i e
biologies
o
liza ds
in
wo
A gen ina
dese s.
Ph.
D.
hesis,
Uni .
o
Texas
a
Aus in.
Diss.
Abs .
35,1.
SAINT-GIRONS,
H. (1953):
No e
su
les
pe iodes
de
la ence
des
ep iles
au
Ma oc.
Bull.
Soco
Zool.
F ance
78: 377-381.
- - (1954):
Cycle
d'ac i i é
e
he mo egula ion
chez
les
ep iles
<leza ds
e
se pen s).
Vie
e
MiHeu
7:
133-226.
SCHOENER,
T. W. (1968):
The
Anolis
liza d
o
Bimini:
esou ce
pa i ioning
in
a
complex
auna.
Ecology
49:
704-726.
- -
aud
GORMAN,
G. C. (1968):
So ne
nicha
di e ences
among
h ee
species
o
Lesse
An illeau
anoles.
Ecology
49: 819-830.
SCHOENER,
T. W.,
and
GoRMAN,
G.
C.
(1970):
Nonsynch onous
spa ial
o e lag
o
li-
za ds
in
pa chy
habi a s.
Ecology
51:
408-4118.
- -
and
SCHOENER,
A. (1971a):
S uc u al
habi a s
o
Wes
Indian
Anolis
li-
za ds
I.
Lowland
Jamaica.
B e io a
368: 1-53.
- - - - (1971b):
S uc u al
habi a s
o
Wes
Indian
Anolis
liza ds
n.
Pue o
Rican
uplands.
Ibid.
375: 1-39.
- - (1974):
Resou ce
pa i ioning
in
ecological
communi ies.
Science 185: 27-
39.
SIEGEL,
S. (1956):
Nonpa ame ic
s a is ics
OT
he
beha io al
sciences.
McG aw
Hill
Book
Company,
Inc.,
New
Yo k-To on o-London.
VALVERDE,
J.
A. (1966):
Sob e
las
sub
especies
de
Chalcides
bed iagai
(Basca).
Bol.
R.
Soco
Española
His .
Na .
(BioU,
64: 169-170.
- - (1967):
Es uc u a
de
una
comunidad
de
e eb ados
e es es.
C.
S.
.
C.
Mad id.
J.
MELLADO,
F.
AMoRES,
F.
PARRKÑO
and
F.
HIRALDO
Cen o
Biológico
del
Su
Reina
Me cedes,
17.
Se illa-12
ESPAÑA
(SPAIN).
:Doiíana, Ac a
Ve eb a a,
2 (2),
1975