The bats of Argentina

SPECIAL P'UBLIC,ATIONS
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Museum o Texas Tech Uni e si y
NUMBER
42 15
JUNE 1999
in collabo a ion wi h he Sam Nobie Oklahoma Museum o Na u ai His o y
,
.THE
BATS
OF
ARGENTINA
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Huben-M, Ba quez
Michael A. Ma es
and Jane K. B aun
,"
We dedica e his publica ion wi h g ea espec and a ec ion o
DR. KARLF. KOOPMAN,
whose dea h lea es a oid in he ield o mamma logy in gene al,
and in he biology o ba s in pa icula .
His in ellec p o ided a shining ligh on he sys ema ics o ba s
ha illumina ed he way o a gene a ion o in es iga o s.
He will be g ea ly missed.
Fo now, he e a e no mo e ques ions.
SPECIAL PUBLICATIONS
Museum
o
Texas Tech Uni e si y
NUMBER 42
THE BATS OF ARGENTINA
RUBEN M. BARQUEZ, MICHAEL A. MARES,
AND JANET
K.
BRAUN
Layou and Design:
Co e Design:
Jane K. B aun
Pa ick Fishe
Copy igh 1999, Museum o Texas Tech Uni e si y
All igh s ese ed. No po ion o his book may be ep oduced in any o m o by any means,
including elec onic s o age and e ie al sys ems, excep by explici , p io w i en pe mission
o he publishe .
This book was se in Times New Roman and p in ed on acid- ee pape ha mee s he guidelines
o pe manence and du abili y o he Commi ee on P oduc ion Guidelines o Book Longe i y o
he Council on Lib a y Resou ces.
P in ed: 15 June 1999
Lib a y o Cong ess Ca aloging-in-Publica ion Da a
Special Publica ions o he Museum o Texas Tech Uni e si y, Numbe 42
Se ies Edi o : Robe J. Bake
THE BATS OF ARGENTINA
Ruben M. Ba quez, Michael A. Ma es, and Jane K. B aun
Illus a ions by No be o Giannini
ISSN 0169-0237
ISBN 0-9640188-9-6
Museum o Texas Tech Uni e si y
Lubbock, TX 79409-3 9 USA
(806)742-2442

CONTENTS
INTRODUCTION _ 1
MATERlALS AND METHODS _ 3
Collec ion and P epa a ion o Specimens 3
Measu emen s 3
Specimens Examined 5
Locali ies and Maps 6
Figu es 6
Taxonomic Keys o he Ba s o A gen ina 8
Tex 8
Desc ip ion o Species 9
ACCOUNTS OF SPECIES 11
Key o he Families o A gen ine Ba s 11
Family Noc ilionidae _ 11
Key o he Species 11
Genus Noc ilio 11
Noc ilio albi en is " , , ,.........
12
Noc ilio lepo inus _., ,.....................................
15
Family Phyllos omidae 19
Key o he Sub amilies 19
Sub amily Phyllos ominae 20
Key o he Species 20
Genus Mac ophyllum , """"" 20
Mac ophyllum mac ophyllum . "" " , , '., 20
Genus Tona ia , " " ',.,.." ,..,,.,., , " " 25
Tona ia
bidens " " , , _ '" ,.
25
Genus Ch o op e us ,.... "" , , " , " 29
Ch o op e us au i us
29
Sub amily Glossophaginae 33
Key o he Species 33
Genus Glossophaga 33
Glossophaga so icina 33
Genus Anou a 38
Anou a caudi e 38
Sub amily Ca olliinae
41
Genus Ca ollia
41
Ca ollia pe spicilla a
41
Sub amily S enode na inae 45
Key o he Species 45
Genus S u ni a 46
S u ni a e y h omos 46
S u ni a lilium 50
S u n i a opo aphilum 57
Genus Pla y hinus 59
Pla y hinus linea us 59
Genus Vampy essa 62
Vampy essa pus ilia 62
Genus A ibeus 63
A ibeus imb ia us 66
A ibeus li u a us 67
A ibeus plani os is 72
Genus Pygode ma 75
Pygode ma bilabia um 75
Sub ami ly Desmodon inae 78
Key o he Species 78
Genus Desmodus 78
Desmodus o undus 78
Genus Diaemus 84
Diaemus youngi 84
Family Vespe ilionidae 86
Key o he Species 87
Genus
Myo is
89
Myo is
aelleni 89
Myo is
albeseens
90
Myo is chiloensis 92
Myo is
keaysi 99
Myo is
le is
99
Myo is
nig icans
110
Myo is ipa ius 144
Myo is ube 113
Myo is simus
115
Genus Ep esicus 117
Ep esicus b asiliensis 117
Ep esicus diminu us
120
Ep esicus
u inalis
123
Genus His io us . 128
His io us mac o us 129
His io us magellanicus 133
His io us mon anus
136
His io us ela us 141
Genus Dasyp e us 142
Dasyp e us ega 143
Genus Lasiu us 145
Lasiu us blosse illii 146
Lasiu us cine eus . 152
Lasiu us a ius
157
Family Molossidae 160
Key o he Species 160
Genus Molossops 162
Molossops neglec us 162
Molossops
emminckii
163
Genus Cynomops 168
Cynomops ab asus 169
Cynomops pa anus
173
Cynomops plani os is 174
Genus Tada ida 178
Tada ida b asiliensis 178
Genus Nyc inomops 184
Nyc inomops la icauda us 184
Nyc inomops mac o is 186
Genus Eumops 190
Eumops au ipendulus 191
Eumops bona iensis
193
Eumops dabbenei 197
Eumops glaucinus 199
Eumops pa agonicus 203
Eumops pe o is 207
Genus P omops 211
P omops cen alis 211
P omops nasu us 212
Genus Molossus 217
Molossus a e , ,.. 218
Molossus molossus , 22]
DISCUSSION 227
BARQUEZ ET AL.-THE BATS OF ARGENTINA
a
An i agus
Nose lea
Figu e 2. Loca ion o he: (a) agus and an i agus in
molossids and (b) he nose lea in phyllos omids.
SPECIMENS EXAMINED
Ac onyms o he museums and ins i u ions om
which specimens we e ob ained ollows Ha ne e al.
(1997) o collec ions in he Wes e n Hemisphe e and
Genoways and Schli e (1981) o all o he s; o p i-
a e collec ions, ac onyms a e he owne 's ini ials: AA,
~----GSL
CBL
5
C-C
ZB
LMdT
Figu e 3. C anial measu emen s as desc ibed in he
Ma e ials and Me hods.
collec ion o Analia Au ino, Tucu nan, A gen ina;
AMNH, Ame ican Museum o Na u al His o y, New
Yo k, New Yo k, Uni ed S a es; BMNH, B i ish Mu-
seum (Na u al His o y), London, England; CEM, col-
lec ion o
Elio
Massoia and amily, Buenos Ai es, A -
gen ina; CM, Ca negie Museum o Na u al His o y,
Pi sbu gh, Pennsyl ania, Uni ed S a es; CML,

6SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
Coleccion de Ma ni e os Lillo, Facul ad de Ciencias
Na u ales e Ins i u o Miguel Lillo, Tucuman, A gen ina;
EBD, Es acion Biologica Do iana, Se iJJe, Spain; EJ,
collec ion o Ennque Jus o, La Pampa, A gen ina; FCM,
Coleccion Mami e os, Facul ad de Ciencias Exac as y
Na u ales, Uni e sidad Nacional de Buenos Ai es,
Buenos Ai es, A gen ina; FMNH, The Field Museum,
Chicago, Illinois, Uni ed S a es; IADlZA-CM, Ins i u o
A gen ino de In es igaciones de las Zonas A idas,
Mendoza, A gen ina; JRC, collec ion o Julio R.
Con e as, CAPRlM, Co ien es, A gen ina; KU, Uni-
e si y o Kansas, Museum o Na u al His o y,
Law ence, Kansas, Uni ed S a es; MACN, Museo
A gen ino de Ciencias Na u ales "Be na dino
Ri ada ia," Buenos Ai es, A gen ina; MCZ, Museum
o Compa a i e Zoology, Ha a d Uni e si y, Cam-
b idge, Massachuse s, Uni ed S a es; MD, collec ion
o Monica Diaz, Tucu nan, A gen ina; MFA, Museo
Flo en ino Ameghino, San a Fe, A gen ina; MLP,
Coleccion de Ma ni e os, Museo de La Pla a, La Pla a,
A gen ina; MJCM, Museo de Ciencias Na u ales y
An opologicas "P o . Juan C. Moyano," Mendoza, A -
gen ina; MPB, collec ion o Ma ia P. Be olini, Buenos
Ai es, A gen ina; MSB, Museum o Sou hwes e n Bi-
ology, Uni e si y o New Mexico, Albuque que, New
Mexico, Uni ed S a es; MSU, Michigan S a e Uni e -
si y Museum, Eas Lansing, Michigan, Uni ed S a es;
MVZ, Museum o Ve eb a e Zoology, Uni e si y o
Cali o nia, Be keley, Cali o nia, Uni ed S a es; OMNH,
Oklahoma Museum o Na u al His o y, Uni e si y o
Oklahoma, No man, Oklahoma, Uni ed S a es; PIDBA,
P og am a de In es igaciones de Biodi e sidad A gen-
ina, Facul ad de Ciencias Na u ales, Tucu nan, A gen-
ina; RAO, collec ion o Rica do A. Ojeda, IADlZA,
Mendoza, A gen ina; RMB, collec ion o Ruben M.
Ba quez, Tucu nan, A gen ina; ROM, Royal On a io
Museum, To on o, Canada; SIG, collec ion o Sil ia 1.
Gue e o, Fundacion Miguel Lillo, Tucu nan, A gen-
ina; SUVM, Ve eb a e Museum, Shippensbu g Uni-
e si y, Shippensbu g, Pennsyl ania, Uni ed S a es;
TCWC, Texas Coope a i e Wildli e Collec ion, Texas
A
&
M Uni e si y, College S a ion, Texas, Uni ed S a es;
TTU, Museum o Texas Tech Uni e si y, Lubbock,
Texas, Uni ed S a es; IBUNAM, Uni e sidad Nacional
Au ono na de Mexico, Ins i u o de Biologia, Mexico,
Dis i o Fede al, Mexico; CUNRC, Uni e sidad
Nacional de Rio Cua o, Coleccion de Mami e os,
Co doba, A gen ina; USNM, Uni ed S a es Na ional
Museum o Na u al His o y, Washing on, Dis ic o
Co-
lumbia, Uni ed S a es.
LOCALITIES AND MAPS
All locali ies lis ed in Appendix I we e o de ed
alphabe ically and numbe ed consecu i ely. No e ha
en ies beginning wi h "ch" ollow en ies beginning
wi h "c," in he Spanish s yle. Maps show he dis ibu-
ion o he numbe ed collec ing locali ies o each spe-
cies. In gene al, each locali y is gi en as i appea ed on
he label. P o ince, depa men , and geog aphic coo -
dina es a e lis ed; hese a e based on Keegan and Keegan
(1993), Anonymous (1968), and egio al maps ob ained
om he Ins i u o Geog a ico Mili a de A gen ina
(Anon., 1954, 1972). Locali ies no ound in hese e -
e ences o in maps a ailable o us appea in Appendix I
as "no loca ed." Locali ies ob ained om museum
specimens o om he li e a u e o en had insu icien
in o ma ion conce ning he poli ical depa men in
which he ba s we e collec ed; hese we e assigned o
an app op ia e depa men , aking in o accoun ha
many depa men s ha e changed hei bounda ies and
e en hei names o e he las 100 yea s. We ha e lis ed
he o iginal depa men whe e a specimen was collec ed
i his was epo ed by he collec o . We also Jis he
cu en depa men o all specimens, e en i hei de-
pa men o collec ion was no eco ded o iginally. The
p o inces o A gen ina a e shown in Figu e 4. Reco ds
o occu ence a e shown on he maps by do s. Because
some locali ies we e in close p oximi y, o we e gi en
in sligh ly di e en o m, a single do may indica e mo e
han one locali y. In such cases, indi idual locali ies can
be ob ained by examining he Specimens Examined,
Addi ional Reco ds, and Appendix 1.
FIGURES
D awings o each species we e made using li e
ba s as models o , in mos cases, om luid-p ese ed
specimens. Some species a e ep esen ed poo ly in col-
lec ions and d awings could no be made o hese. Fo
a ew species, d awings we e made using museum skins,
pho og aphs o li e animals, and li e a u e desc ip ions.
Pho og aphs o la e al, do sal, and en al iews o he
skull, and o he uppe oo h ow, we e aken om A -
gen ine specimens in mos cases. Fo hose species no
pho og aphed, a d awing o hei mos impo an cha -
ac e s was made wi h a came a lucida.
BARQUEZ ET AL.-THE BATS OF ARGENTINA 7
72'
68'
64'
60
56' 52'
Tucuman
24'
28'
32'
36'
40'
48'
52'
Tie a
del
Fuego
.pp
Mal inas Islands
o
I
200
I
400 km
!
igu e 4. P o inces o A gen ina.
8SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
a
Pa acone
Mesos yle
Pa as yle
Mesos yle
Hypocone
Pa acone
Me acone
Me as yle
b
En oconid
P o oconid
P o oconid
Me aconid
T igonid
Talonid
Pa aconid
Me aconid
Hypoconid
Figu e 5. Cusps o he ee h o he: (a) maxilla and he (b) mandible. Abb e ia ions: C, canine; P o p, p emola ;
M o m, mola . I, 2, and 3 indica e i s , second, and hi d, espec i ely.
TAXONOMIC KEYS TO THE BATS OF
ARGENTINA
Keys we e de eloped om an examina ion o mu-
seum specimens and o he da a ob ained in he ield.
Diagnos ic cha ac e s men ioned in he keys we e de-
e mined a e examining he a iabili y o he species
wi hin i s ange o dis ibu ion in A gen ina.
TEXT
We epo he i s lise o he scien i ic name o
each species o subspecies. Mo e ex ensi e synonymies
a e p o ided in cases whe e he axonomic his o y is
mo e complex.
The ype locali y is, in gene al, ha gi en in he
i s ci a ion. I is p esen ed in a sepa a e sec ion be-
cause in many cases u he discussion is necessa y. Dis-
ibu ion ou side A gen ina is based on he li e a u e;
dis ibu ion in A gen ina is based on ou esea ch, wi h
commen s on unce ain o doub ul eco ds.
Species desc ip ions a e based, whene e possible,
on specimens collec ed in A gen ina o , when specimens
om A gen ina we e sca ce, on specimens om neigh-
bo ing coun ies. All species o subspecies a e desc ibed.
Colo is a he subjec i e and may be a ec ed by
ac o s such as s a e o p ese a ion, ime in s o age,
pelage de e io a ion, ea men o skins wi h chemicals,
empe a u e, humidi y, and ligh , as well as such bio-
logical ac o s as indi idual and popula ion a iabili y.
In
some cases, colo is use ul in helping disc imina e
be ween simila axa and we use simple desc ip ions
o colo pa e ns whe e hese a e impo an in dis in-
guishing be ween simila species o subspecies.
Skull cha ac e s ha aid in iden i ica ion a e p e-
sen ed. Den al o mula and oo h desc ip ions (Fig. 5)
ollow Mille (1907) and Handley (1959). The sample
size, mean, ange, and s anda d de ia ion o ex e nal
and c anial measu emen s a e gi en in he ex o in
ables. The numbe o specimens a ailable o each spe-
cies, when g ouped by sex, age, o pa icula a ea, was
no su icien ly la ge o allow a mo e de ailed s a is i-
cal analysis (see Willig,
J
983, o he impo ance o
sample size in ba s udies).
The Commen s summa ize a ailable in o ma ion
on a ious aspec s o he biology o he species.
Specimens Examined lis s specimens ha we e
examined by us and iden i ied as o axon. Locali ies
BARQUEZ
ETAL.~THE
BATS OPAR.GENTINA
a e gi en o each specimen as hey we e w i en on he
ags, wi h ob ious e o s co ec ed.
Addi ional Reco ds lis s specimen eco ds ound
in he li e a u e ha we e unambiguously assignable o
a pa icula axon. Locali ies o museum specimens ha
we e no examined we e included only when he spe-
cies was easily iden i iable.
DESCRIPTION OF SPECIES
In
gene al, he a angemen o species ollows
Cab e a (1958), al hough ecen in o ma ion on he e-
la ionships o amilies, sub amilies, gene a, and sub-
gene a is included. The Sub amily S u ni inae is no
9
ecognized. Species o he Genus
S u ni a
ha e been
included in he Sub amily S enode ma inae o he Fam-
ily Phyllos omidae. The Family Desmodon idae is no
ecognized, bu is conside ed a sub amily o
Phyllos o nidae. The Genus
Dasyp e us
is ecognized
as a alid genus. The Genus
Cynomops
is conside ed
dis inc om
Molossops
(Thomas,
1920a),
and
Nyc i nomops
as dis inc om
Tada ida
(F eeman,
198
j
a).
O he di e ences wi h espec o Cab e a (1958)
a e discussed in he Desc ip ion sec ion o each species
accoun , and deal p ima ily wi h he alidi y o scien-
i ic names, cla i ica ion o he synonymy, spelling, and
dis ibu ion.

ACCOUNTS OF SPECIES
I.
KEY TO THE FAMILIES OF ARGENTINE BATS
Nose lea p esen , well de eloped, and iangula in shape, o educed and
ounded Phyllos o nidae (p. 19)
Nose lea absen . 2
I'.
App oxima ely one-hal o he ail included in he u opa agium and
one-hal is ee Molossidae (p. 160)
Tail long and almos comple ely included in he u opa agium, o sho
and appea ing on he do sal su ace o he u opa agium 3
2 (l ').
2'.
3 (2').
3'.
Tail e y long and almos comple ely included in he u opa agium Vespe ilionidae (p. 86)
Tail sho , included in he u opa agium, bu ip appea ing on he
do sal su ace Noc ilionidae (p. II)
FAMILY NOCTILIONIDAE
KEY TO THE SPECIES
I. Leng h o o ea m> 70 mm; leng h o hind oo > 25 mm; wingspan
abou 500 mm
Noc ilio lepo inus
I'. Leng h o o ea m
<
70 mm: leng h o hind oo
<
20 mm; wingspan
abou 400 mm
Noc ilio albi en is
Genus Noc ilio
Desc ip/ian.-
Size medium o la ge when com-
pa ed wi h o he Ame ican ba s, wi h
Noc ilio lepo inus
being la ge han
Noc ilio albi en is .
The os um is
ele a ed and he muzzle is poin ed; a nose lea is ab-
sen and he na es open down and o wa d. The uppe
lip has a p onounced e ical g oo e o old o skin,
con e ing a ha e-like (lepo ine) aspec . The ea s a e
ubula o unnel-shaped, la ge, and sepa a ed; he a-
gus is small and se a ed. The wings a e long and na -
ow, and a ach o he body jus below he knees. The
emu is sho e han he ibia, which is well de eloped,
pa icula ly in
N. lepo inus.
Claws a e long, sha p, and
hooklike. The calca is la ge and plays a unc ional ole
in he mo ion o he u opa agium when p ey a e seized.
The ail is sho , ex ending o he middle o he
u opa agiu n and p o udes h ough he do sal su ace.
Pelage colo a ies, bu always has ed, o ange, o yel-
low ones. A pale mid-do sal s ipe may be p esen .
The skull is simila in bo h species, excluding size
and a angemen o mola s and p emola s. In N.
lepo inus
spaces a e p esen be ween he mola s,
whe eas in
N. albi en is
hese gaps a e no eadily ap-
pa en . Skull shape is dis inc i e; he b aincase is high
and he sagi al c es is well de eloped, pa icula ly in
adul males. The os um is sho , app oxima ely one-
hal o he leng h o he b aincase. The pala e is con-
ca e and ex ends pos e io ly beyond he oo h ows. The
ympanic bullae a e small.
The den al o mula is I 2/1, C 1/1, P 1/2, M 3/3,
o al 28. In he uppe oo h ow, 12 is smalle han 11 and
lies behind i ; P I has well-de eloped cusps; M I and
M2 a e simila in size and, in
N. albi en is,
lack pos e-
io ema gina ions (pos e io ema gina ions a e p esen
in he p emola s o
N. lepo inus
so ha he e a e no
clea ly de ined spaces be ween hem). The lingual side
o C I is sligh ly conca e and has a medial idge, whe eas
he labial side is uni o mly con ex. Ano he di e ence
be ween species in his genus is ha in
N. albi en is
he hypocone o M I and M2 is ela i ely la ge han
ha o
N. lepo inus.
Addi ionally, MI and M2 ha e a
commissu e ha connec s he hypocone wi h he com-
missu e connec ing he p o ocone and me acone. The
occlusal su ace o M3 is one-hal he size o M2.
11
12 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
64°
60°
,,
,,,
L--------------
.458
!
:465
,
'
,
'
"
,,--'
,
"
: 627.:
, ,
--- :
~""----,-"
I,,: >
I~
:
/
Noc ilio albi en is Desma es
Figu e 6. Map o he locali ies o Noc ilio albi en is.
No c il io albi en is Desma es , Nou eau
dic ionnai e d' his oi e na u elle, appliquee aux a s,
p incipalmen
a
I'ag icul u e, e
a
I'economie u ale e
domes ique; pa une socie e de na u alis es. Nou elle
edi ion, p esqu' en ie emen e ondue e
conside ablemen augmen ee. Ch. De e ille, Pa is,
23:15,1818.
TypeLocali y.-
B azil, Bahia, Rio Sao F ancisco.
Dis ibu ion>:
Pa ana Ri e basin in sou he n
B azil, Pa aguay, and A gen ina no hwa d o sou he n
Mexico. Hood and Pi occhelli (1983) desc ibed he dis-
ibu ion o N. a. cab e ai as including wes e n U u-
guay, no heas e n A gen ina, Pa aguay, eas e n Boli ia,
and hose pa s o adjacen B azil. Howe e , Ande son
e al. (1982) and Ande son (1997) did no men ion speci-
mens o his subspecies om sou heas e n Boli ia.
Da is (1976) desc ibed N. a. cab e ai and di e en i-
a ed i om neighbo ing popula ions o
N.
a. a inis by
i s smalle size and pale colo . Mye s and We zel (1983)
no ed ha measu emen s o specimens om many o
he locali ies ci ed by Da is (1976), in addi ion o hei
own specimens, sugges ed a mo e complex si ua ion.
o
100 200 300 km
I
I I
I
We ha e summa ized hei esul s below. Specimens
om he Depa men o Beni, Boli ia, which acco d-
ing o Da is (1976) belong o
N.
a. a inis, a e small bu
sligh ly la ge han N. a. cab e ai, and s ikingly di e -
en han nea by samples o N. a. cab e ai om
Magdalena and San Joaquin, also loca ed in Beni. Speci-
mens om San a C uz a e signi ican ly la ge han hose
om Beni and a e simila in size o hose om he wes -
e n Amazon basin. Popula ions om cen al and sou h-
e n Pa aguay a e simila o hose om Beni and a e
sligh ly la ge han N. a. cab e ai, bu much smalle
han hose om San a C uz. Mye s and We zel (1983)
also poin ed ou ha he a e age size o
N.
a. a inis
(sensu Da is, 1976) seems o esul om a mixing o
he highly he e ogeneous popula ions om Beni and
San a C uz. We examined i e male specimens (CML)
om Beni. In almos all measu emen s, specimens om
Resis encia, Chaco P o ince, A gen ina, a e la ge han
he specimens om Beni. Based on he dis ibu ions
gi en by Da is (1976), he specimen om Chaco should
pe ain o N. a. cab e ai; howe e , i is mo pho ne i-
cally mo e allied wi h
N.
a. a inis. Ibanez Ula gui
(1981) epo ed ha he Llanos de Apu e in Venezuela
a e loca ed in an a ea suppo ing di e en subspecies
o N. albi en is and also no ed ha o ea m measu e-
BARQUEZ ET AL.-THE BATS OF ARGENTINA
Figu e 7.
Noc ilio albi en is:
(a) ace and (b) u opa agium,
men s o specimens om Romang, San a Fe P o ince,
A gen ina, gi en by C espo (1974), a e simila o N.
a.
a inis
om he uppe Amazon, a he han
N.
a.
cab e ai.
Clea ly, he axonomic s a us o he subspe-
cies is un esol ed.
Dis ibu ion in A gen ina
(Fig.
6).-
Bu meis e
(1879) epo ed
N. lepo inus
in Sal a P o ince; how-
e e , Cab e a (1938), based on Bu meis e 's desc ip-
ions, ea ed i as
N. albi en is
a he han
N. lepo inus.
N. albi en is
has no been collec ed in no hwes e n
A gen ina.
13
Desc ip ion
(Figs. 7 and
8).-
Size is much
smalle han
N. lepo inus.
Pelage colo anges among
a ious shades o b own, g ay, o ange, and yellow.
Sexual dimo phism in colo is common, emales being
b owne han males; a pale mid-do sal s ipe may be
dis inc , ha dly e iden , o absen . Males a e usually
sligh ly la ge han emales and ha e a mo e p onounced
sagi al c es , which may be absen o no eadily ap-
pa en in young specimens and emales; males gene -
ally a e hea ie han emales. Fee a e long and obus ,
bu less so han
N. lepo inus.
Measu emen s.-
Table 1.
Commen s.-
Mos specimens cap u ed by Mye s
and We zel (1983) in he chaco lew o e lakes o i -
e s; wo we e ound in oos s in "queb ac ho"
iAspidospe mum
sp.) ees, one oos ing in a hollow
ee wi h he en ance I m abo e he g ound and he
o he oos ing in a ho izon al b anch 20 e n in diame e
and 6 m abo e he g ound. Roos s also a e placed in
houses and buildings. Smi h and S a e (1979) no ed
he abili y o his species o ly h ough dense ege a-
ion; o he in es iga o s ha e commen ed on hei o -
aging o e wa e , some imes lying in g oups o e i -
e s and s eams.
Food habi s a e simila o hose o
N.
lepo inus,
excep o a g ea e p e e ence o aqua ic insec s.
Whi ake and Findley (1980) s udied ecal pelle s and
ound emains o Coleop e a (Dy iscidae, Ca abidae),
Hemip e a, Homop e a (Ce copidae), Lepidop e a, and
Dip e a; Dy iscidae was he mos abundan p ey (22.5%
o he o al olume). Howell and Bu ch (1974) obse ed
indi iduals eeding on mulbe y ui s (Mo aceae).
No hing is known abou he ep oduc i e biology
o his species in A gen ina. Mye s and We zel (1983)
cap u ed p egnan emales in he chaco o Pa aguay in
Augus and Oc obe , lac a ing emales in Sep embe ,
and a young indi idual in Janua y.
Ibanez
Ula gui (1981)
epo ed ha emales had well-de eloped e uses in Ven-
ezuela in la e Janua y, ga e bi h om mid-May o ea ly
June, and lac a ed un il la e July. Ma ing p obably oc-
cu ed om la e Decembe o Janua y, wi h a ges a ion
pe iod las ing abou i e mon hs. He sugges ed ha his
Neo opical species has a seasonal monoes ous sexual
cycle in he egion o "Ha o El F io." Gi en he di e -
ences in iming o ep oduc ion in Venezuela and Pa a-
guay, i is unlikely ha a consis en seasonali y cha ac-
e izes he ep oduc i e cycle o his species h ough-
ou i s ange.
14
SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
'
Figu e 8. Skull and ee h o Noc ilio albi en is. Adul male om Depa amen o Ben , Boli ia (CML
1408). Ba
= ]
em. Tee h no o scale.
Specimens Examined
(27).- CORRIENTES:
Co ien es, 1 (ROM); Mbu ucuya, 13 (MACN);
Posadas, 61 kmW,alongHwy 12,2(1 CML, IOMNH).
MISIONES: Posadas, I(MACN). SANTA FE Ro nang,
Isla EI Lau el, 10 (MACN).
Addi ional Reco ds.-
CHACO: Apipe G ande
(Ro na ia
and Abalos, 1950); Resis encia (Fo nes and
Massoia, 1968). CORRIENTES: Manan iales (Fo nes
and Massoia, 1967). FORMOSA: Clo inda (Da is,
1976; TCWC no examined); Pa que NacionaJ Rio
Pileomayo (S. Heinonen F. and R. M. Ba quez, in li .).
BARQUEZ ET AL.-THE BATS OF ARGENTINA
60°
32°
----------1
,,--,'
1,,--_,-,
,
,
//
'
o
I
300 km
I
Figu e 12. Map o he loca li ies o Mac ophyllum
nac ophyllum.
eas e n and wes e n B azil, Pa aguay, and no heas e n
A gen ina.
Dis ibu ion in A gen ina (Fig. 12).- Al hough
his species is widely dis ibu ed, i is uncommon o
di icul o ca ch, and he e a e ew known eco ds (see
Ha ison, 1975; Wilson and Gama a de Fox, 1991).
Desc ip ion (Figs. 13 and 14).- Size is small,
and he o ea m a ies om 34 o 39 mm; weigh a -
ies om 6 o 9 g. The ail, which is enclosed in he
u opa agium, almos eaches he pos e io edge, as in
espe ilionids. The nose lea
is
long and b oad, wi h a
conspicuous cen al keel, many c enula ions on he
lowe edge, and a sha p ip. Fou o i e well-de el-
oped glands a e si ua ed behind he nose lea on bo h
sides o he muzzle. A g oup o h ee wa s enci cled by
eigh smalle wa s is ound on he lowe lip. The muzzle
is sho and he eyes small. The wing memb anes a e
b oad, naked, and a ach a he middle o he ibia. The
ea s a e b oad and well de eloped and sha p a he ip;
he inne edge is dis inc ly con ex. The u opa agium is
wide and long, comple ely enclosing he ail. Se ies o
de mal den icles a e p esen along he dis al en al su -
ace o he u opa agium. The ee a e long and shaped
like Noc ilio; he ou e digi has a single phalanx,
whe eas he o he s ha e wo. Do sal colo a ion o he
pelage is da k b own, he hai s pale a he base; he
en e is pale han he do sum.
0
l og and Luce o (1981 )
21
desc ibed he colo a ion o his species as blackish
b own wi h a pale en e .
The skull is e y small and is cha ac e ized by
he o wa d displacemen o he maxillae and he e-
ac ion o he nasal openings o he second p emola s.
P opo ionally, he os um is less han one-hal he
leng h o he b aincase (measu ed as he dis ance om
he pos o bi al cons ic ion o he pos eno mos poin
o he skull). The pala e is long, ex ending pos e io ly
beyond he edge o he uppe oo h ow. The hamula
p ocesses o he p e ygoids a e pa allel an e io ly and
s ongly di e gen pos e io ly. The ympanic bulla is
small and encloses less han one-hal o he cochlea.
The sagi al c es is absen ; he zygoma ic a ch, al hough
agile, is comple e.
The den al o mula is I 2/2, C 1/1, P 2/3, M 3/3,
o al 34. The 11 is la ge and p ocumben ; 12 is educed
and only eaches he cingulum o 11. The canines a e
app oxima ely wice he size o he la ges inciso and
a e cu ed. The PI is small, much smalle han P2, and
is sepa a ed om i by a small gap; iewed la e ally,
he P2 is highe han he mola s. The M I is he la ges
oo h in he mola ow and, like M2, has well-de ined
W-shaped commissu es. Only wo commissu es and a
ace o a hi d can be seen on M3. In he lowe oo h ow,
iI and i2 a e small, wi h ilobula e edges. The an e io
ace o p I is in con ac wi h he canines; p2 is small and
displaced o he lingual side o he oo h ow, so ha pI
and p3 a e in con ac . The m I and m2 a e well de el-
oped and simila o one ano he ; m3 is much smalle
and he alonid basin is indis inguishable due o he
closeness o he hypoconid and en oconid.
Measu emen s.- Table 2.
Commen s.- S udies dealing wi h his species a e
sca ce and he e a e no da a on i s biology in A gen ina.
Ha ison (1975) e iewed i s biology. Dicke man e al.
(1981)
epo ed he i s eco d om Gua emala; hey
obse ed colonies o mo e han 70 indi iduals li ing in
sewe sys ems and no ed hey we e ha d o cap u e.
Whi ake and Findley (1980) ound Chi ono nidae, spi-
de s, and Coleop e a in ecal pelle s o specimens om
Cos a Rica and Panama, whe e ba s we e cap u ed a
ew cen ime e s o e he wa e 's su ace. Ga dne
(1977a)
summa ized da a on ood habi s in Panama and
men ioned he occu ence o Hemip e a (Gue idae) in
s omachs. Da is e al. (1964) collec ed his species a

22 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
c
Figu e 13. Mac ophyllum mac ophyllum: (a) and ( ) ace; (b) u opa agium; (c) agus; (d) hind oo ; and (e) a icula ion
o he o ea m and digi s.
BARQUEZ ET AL.-THE BATS OF ARGENTINA 23
• 1
/
... .....
.'
, I
.
,
Figu e 14. Skull and ee h o Mac ophyllum mac ophyllum Adul male om Cue a Ma ia An onia,
Misiones P o ince (MACN 17026). Ba
=
1 em. Tee h no o scale.
he bo om o ne s placed close o he wa e which, o- wa e low; i was absen om sewe sys ems wi h no
ge he wi h ea u es such as long ee and a la ge wa e low. They no ed ha males we e mo e common
u opa agium, sugges ed o he au ho s ha his species han emales, and epo ed g oups o med o males only.
ed on aqua ic Insec s o small ish, al hough he s om- B asse and Cha les-Dominique (1990) epo ed ha i
achs we e emp y. Seymou and Dicke man (1982) ob- oos s in cul e s, ca es, and sub e anean ca i ies in
se ed he p esence o his species in Gua emala In- F ench Guiana. Wilson and Ga na a de Fox
(l99l)
col-
side cul e s
I
m wide and 10 m long and wi h s able lec ed se e al indi iduals in he mon h o Feb ua y om
24 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
Table
2.
Measu emen s
o Mac ophyllum mac ophyllum, Tona ia bidens,
and
Ch o op e us au i us.
Fo a desc ip ion o he measu emen s, see he Ma e ials and Me hods. Mean, s anda d de ia ion, and sample
size a e ollowed by ange.
Va iable
Mac ophyllum mac ophyllum Tonalia bidens Ch o op e us au i us
To al leng h
81.0, 1 96.1 i 5.13,12 106 1i9.42,7
85.6 - 1050 95.0 - 1250
Tail leng h
35.0, I 16.5 i 2.36,12 11.0, 1
12.0-21.0
Hind oo leng h
10.0, I 15.3 i 2.30, 12 21.5 i 0.86,7
10.0 - 20.0 20.0 - 22.5
Ea leng h
16.0, I 25.2 i 2.25, 12 398 ±4.68, 7
22.0 - 295 32.0 - 44.7
Fo ea m leng h
36.0, I 57.6iI.16,12 820 i 1.05, 8
55.0 - 59.0 80.0 - 83.0
Weigh
32.7 i 9.63,4 85.0 i 7.07,2
26.0 - 47.0 800 - 90.0
Condylobasal leng h
14.3, I 24.5 ± 0.32, 10 31.2 i 0.28, 8
24.1-25.1 309-31.8
Leas in e o bi al b ead h
7.2 i 0.40,5 8.1 i 0.3 1,7
6.8 - 7.8 7.7 - 8.7
Zygoma ic b ead h
13.6iO.22,10 19IiO.37,8
13.4-14./ 18.6-19.8
G ea es leng h o sku II
6.3, 1 27.9 i 0.46, 10 35.4 i 0.44, 8
27.3 -
n.7
34.6 - 36.2
Pos o bi al cons ic ion
3.2, I 5.7iO.12,10 62iO.II,8
5.7 - 6.1 6.1 - 6.4
B ead h o b aincase
7.9, I 11.2±0.16,10 13.9 i 0.29, 8
11.1 - 11.5 13.3 - 143
Leng h o maxilla y oo h ow
5.5, 1 10.OiO.31,10 12.9 i 022,8
97 - 10.8 126 - 13.3
Pala al leng h
11.9 i 0.22,10 156 i 0.54, 8
11.6-12.3 14.9 - 16.6
Mas oidal b ead h
8.4, I 13.3 i 0.13, 7 7.4 i 0.35, 8
13.2 - 13.6 16.9 - 17.9
Leng h o mandibula oo h ow
11.3iO.21,6 12.6 i 0.55,8
11.1-11.6 12.1 - 13.9
Leng h o mandible
18.0 i 0.26, 9 24.3 i 0.21, 8
17.7-18.5 24.1 -24.7
C-C (wid h ac oss canines)
3.4, I 5.9 i 0.28, 10 7.2 i 0.27,8
5.5 - 6.3 69 - 7.7
M-M (wid h ac oss mola s)
8.9 i 0.20, 10 11.7 i 022, 8
8.7 - 9.3 11.3 - 11.9
a ca e in Pa aguay, whe e hey occu ed wi h
Ca ollia
pe spicilla a;
one emale
M mac ophyllum
showed e i-
dence o ecen lac a ion.
a wide one in which wo specimens o
Mac ophyllum
and one
Ca ollia
we e cap u ed.
Fo nes e al. (1969) p o ided he only da a o his
species in A gen ina. They eco ded emains o chi in
on he inne side o he u opa agia o specimens col-
lec ed in Cue a Ma ia An onia, a ew me e s om he
Pa ana Ri e . The ca e had wo en ances, a na ow one
used by a colony o abou 150
Desmodus o undus,
and
Ha ison (1975) epo ed ha a 17.5 mm e us al-
eady had he dis inc i e nodules on he u opa agium
p esen . Wilson (1979) cap u ed p egnan emales in Oc-
obe in El Sal ado , in Ma ch, May, and Augus in Cos a
Rica, and in Oc obe and No embe in F ench Guiana.
Specimens Examined
(I ).-MISIONES: Cue a
Ma ia An onia, 1 (MACN).
BARQUEZ ET AL.-THE BATS OF ARGENTINA
60°
25
,
,
-----------I
,
,
,
,
Figu e 15. Map o he locali ies o Tona ia bidens
Genus Tona ia
Tona ia bidens (Spix)
Vampy us b id ens Sp
ix ,
Si
m
i
a u n e
Vespe ilionu n B asiliensium species no ae ou
h s oi e
na u elle des especies nou elles de singes e de chau e-
sou is obse ees e ecueillies pendan Ie oyage dans
lin e ieu du B esil execu e pa o d e de S. M. le Roi
de Ba ie e dans les annees 1817, 1818, 1819, 1820.
F ancisci Se aphici Hubsch nanni, Monachii, p. 65,
1823.
Tona ia bidens: G ay, A synopsis o he species o
he Class Mammalia, as a anged wi h e e ence o hei
o ganiza ion, by Cu ie , and o he na u alis s, wi h spe-
ci ic cha ac e s, synony na, &c &c, ol. 5,
in
The a u-
mal kingdom a anged in con o mi y wi h i s o ganiza-
ion, by he Ba on Cu ie , wi h addi ional desc ip ions
o all he species hi he o named, and o many no be-
o e no iced. (E. G i i h, C. H. Smi h, and E. Pidgeon,
eds.). G. B. Whi ake , London, p. 69,1827.
Type Locali y. - B azil, Bahia, Rio San F ancisco.
Dis ibu ion.- Williams e al. (1995) sepa a ed
T bidens, which hey desc ibed as a mono ypic spe-
cies, om T sau ophila, which was sepa a ed
in o
h ee
subspecies. The ange o T bidens ex ends om no h-
o
200 400 km
I I I
eas e n B azil h ough Pa aguay, eas e n Boli ia, and
no he n A gen ina.
Dis ibu ion in A gen ina (Fig. 15).- Tona ia
sil ico!a was ci ed p e iously o Palma Sola, Jujuy
P o ince (Fo nes e al., 1967) and Ca a a as del Iguazu,
Misiones P o ince (Villa-R. and Villa-C., (971). How-
e e , hese specimens we e T bidens and we e he i s
epo ed o A gen ina (Ba quez e al., 1993).
Desc ip ion (Figs. 16a, 16b, 16c, and 17).-
In
gene al, his species esembles T sil icola (Figs. 16d
and 18). Diagnos ic cha ac e is ics a e: he ea s a e com-
ple ely sepa a ed; he size o he skull is gene ally la ge
han
T
sil ico!a (smalle han
T
sau ophilay;
he b ead h
o he pos o bi al cons ic ion is much la ge han in
T
sil icola;
he a io o he g ea es leng h o he skull
and he pos o bi al cons ic ion is less han 5.5 in T
bidens and g ea e han 5.9 in T sil icola. The ail p o-
udes do sally om he middle o he u opa agiu n. The
gene al colo pa e n o he pelage is g ayish b own,
wi h pale ashy g ay on he sides o he neck, on he
shoulde s, and on he en e . The hai o he en e is
usually da ke basally, al hough his is di icul o see
because o he pale ips. The p oximal one-hal o he
o ea m is u ed do sally and en ally, di e en ia ing
his species om specimens o T sil icola examined
26 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
d
Figu e 16. Tona ia bidens (a, b, and c) and Tona ia sil icola (d): (a) ace; (b) o ea m; (c) u opa agium (do sal iew);
and (d) ace.
om Boli ia, which we e less u ed on he o ea m.
The memb anes gene ally a e naked excep o a small
s ip a he ou e edge o he plagiopa agium. The ea s
a e la ge and ounded, al hough smalle han in
T
sil icola, and do no ex end beyond he nose when laid
o wa d. The
inne
edges o he p oximal one-hal o
he ea s a e immed wi h whi e o pale g ay hai s. The
nose lea is b oad a he base and na ows ab up ly om
he middle o he ip; he ip is no sha ply poin ed. The
uppe lip is smoo h and lacks wa s; he lowe lip has a
cen al wa su ounded by smalle ones. The agus is
well de eloped; he an i agus is no iceably smalle han
in T sil icola. The u opa agiu n is b oad and suppo ed
by calca s, which a e smalle and weake han in
T
sil icola.

BARQUEZ ET AL.- THE BATS OF ARGENTINA
,
,I
27
Figu e 17. Skull o
Tona ia bidens.
Adul male om Laguna La B ea, Jujuy P o ince (OMNH
18788). Ba
=
1 e n.
The b aincase is b oad and he pos o bi al con-
s ic ion sub le in compa ison wi h
T sil icola.
The zy-
goma ic a ches a e slende , al hough hey appea b oad
1D
la e al iew and show a medial cons ic ion. The
lambdoid and sagi al c es s a e poo ly de eloped. Tym-
panic bullae a e small, ba ely co e ing he middle o
he cochlea. The mas oid p ocesses a e well de eloped.
In la e al iew, he os um shows a p onounced slope
om he b aincase o he op o he nasal opening.
The den al o mula is 12/1, C 1/1, P 2/3, M 3/3,
o al 32. The inne inciso s a e well de eloped, in con-
ac wi h he each o he , and sligh ly p ocumben and
conca e an e io ly, appea ing g oo ed. The 12 is iny
and placed wi hin he cingulum o he canine. The up-
pe canines a e sligh ly ecu ed. The PI is small and
ouching he canine; i is iangula in shape in la e al
iew; he an e io edge i s comple ely in he canine
cingulum. The P2 is he highes oo h in he ow. The
M1 and M2 a e squa e-shaped, he cusps a e well de-
eloped, and all ha e he W-shape cha ac e is ic o he
sub amily. The M3
IS
educed and only wo com nis-
su es a e dis inguishable wi h he pa as yle be e de-
28
SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
Figu e 18. Skull and ee h o Tona ia sil ico!a. Adul emale om San a C uz de la
Sie a, Boli ia (CML 1476). Ba
=
I em. Tee h no o scale.
eloped han in he o he mola i o m ee h. In he lowe
oo h ow, iI is well de eloped, highe han i is wide,
and has smoo h edges. The canines a e in con ac
wi heach o he behind he inciso s. Bo h p 1 and p3 a e
la e ally comp essed and unicuspida e; p2 is minuscule
and usually displaced o he labial side excep
10
one
specimen in which i was displaced o he lingual side.
The ou h commissu e o m3 is educed
10
compa i-
son wi h he o he mola s.
Measu emen s.s-« Table 2.
Commen s.s-:
Ve y li le is known abou he eed-
ing habi s o his species. Acco ding o Ga dne (1977a),
ui s and insec s a e i s main ood. Humph ey e al.
BARQUEZ ET AL.-THE BATS OF ARGENTINA
(1983) epo ed a die o insec s, mainly Coleop e a and
Homop e a, a lesse p opo ion o A aneidae and Lepi-
dop e a; he also ound a ace (0.7%) o e eb a es.
Mye s and We zel (1983) ound insec chi in and ui
pulp in he s omachs o Pa aguayan specimens. Obse -
a ions o ca ni o ous eeding in his species ha e been
a e and i is conside ed o be p ima ily a lea -insec
eede . Howe e , he s omach o a specimen caugh in
Laguna La B ea, Jujuy P o ince, was illed wi h down
ea he s, indica ing he consump ion o a young bi d.
The ep oduc i e cycle is poo ly known. B eed-
ing may occu mo e han once a yea (Wilson, 1979),
since p egnan emales we e eco ded in Janua y (Cos a
Rica), Feb ua y (Gua emala), May (T inidad), July
(Pe u), and Augus (Hondu as). Young we e epo ed
in Augus in Hondu as and Cos a Rica. A specimen om
Laguna La B ea, a male wi h well-de eloped sc o al
es es, was collec ed in mid-June (win e ). Ga dne
(1976) epo ed p egnan emales om Pe u a he end
o July, bu no ep oduc i e ac i i y in Ap il.
Specimens Examined
(13).-
JUJUY: Laguna La
B ea, 8 (6 MACN, 1 OMNH, I ROM); Palma Sola,
Mina de Azu e, 4 (3 ROM, 1 TCWC). MISIONES:
Ca a a as del Iguazu, 125 m, 1 (lBUNAM).
Genus
Ch o op e us
Ch o op e us au i us (Pe e s)
Vampy us au i us Pe e s, Mona sbe ich e de
Koniglich P eussischen Akademie de Wissenscha en
zu Be lin, p. 415, 1856.
Ch o op e us au i us: Pe e s, Mona sbe ich e de
Koniglich P eussischen Akademie de Wissenscha en
zu Be lin, p. 505, 1865.
Type Locali y.- Mexico (Oaxaca o Ve ac uz, see
Medellin, 1989, and Ca e and Dolan, 1978, o dis-
cussion o ype locali y).
Dis ibu ion.- Th ee subspecies a e ecognized:
C. a. au i us , dis ibu ed om Mexico o no he nmos
Sou h Ame ica; C. a. guaianae om Venezuela, he
Guianas, and no he n B azil; and C. a. aus alis, om
eas -cen al Pe u, sou he n B azil, Pa aguay, Boli ia,
and no he n A gen ina.
Thomas (1905) epo ed specimens om Guyana
and sou he n B azil, dis inguishing be ween wo sub-
species, guaianae and aus alis, bu some au ho s ha e
29
ques ioned he exis ence o geog aphic aces (Handley,
1966). We ound no subspeci ic di e ences in speci-
mens collec ed h oughou he ange; howe e , a la ge
sample size is equi ed o sol e his p oblem. Un il a
e ision is conduc ed, we e ain Thomas' (1905) lis ing
o he A gen ine subspecies.
Ch o op e us au i us aus alis Thomas
Ch o op e us au i us aus alis Thomas, Annals
and Magazine o Na u al His o y, se . 7,16:308,1905.
Type Locali y.- Pa aguay, Concepcion, 300 m.
Dis ibu ion in A gen ina.- Figu e 19.
Desc ip ion (Figs. 20 and 21).- This is one o
he la ges ba s in A gen ina, wi h he o ea m measu -
ing om 80 o 83 nun in he specimens examined (o he
specimens ha e been epo ed o ange om 78 o 84
mm). The ea s a e la ge and sepa a ed om each o he .
The nose lea is long and hick, i s basal ma gin ee o
he muzzle. The agus is small compa ed o he ea s, is
Ianceo la e in shape and has a poin ed ip; he an i agus
is educed. The uppe lip is smoo h. The lowe lip has a
wa in he middle, a smalle one on each side, and a
small g oo e on i s an e io su ace. The wing mem-
b anes a e b oad and well de eloped; he
plagiopa agium is a ached o he base o he oo a he
p oximal end o he oes. The u opa agium is long,
eaching he base o he ee and is suppo ed by well-
de eloped calca s. The ail is iny and enclosed in he
u opa agium. The u is long and ine, and ex ends o e
he wing memb anes o he middle o he o ea m, bo h
do sally and en ally; a b ush o hai is p esen a he
base o he humb. Colo is blackish b own abo e and
g ayish below. The memb anes a e da k and he wing
ips a e whi e; he p opa agium is somewha anspa -
en , and is pale han he o he memb anes.
The b aincase is obus and ele a ed, wi h a well-
de eloped sagi al c es . A lambdoidal c es is less de-
eloped bu clea ly isible. The pos o bi al cons ic ion
is appa en , ma king a sha p di ision be ween he b ain-
case and he b oad os um. The zygoma ic a ches a e
hin (al hough b oad in la e al iew); iewed la e ally
hey expand do sally, and especially pos e io ly. The
mas oid p ocesses a e hea y. The basila pi s a e shal-
low and he ympanic bullae a e small.
30 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
60
0
',.
{
198 .·';
Figu e 19. Map o he locali ies o Ch o op e us au i us.
The den al o mula is I 2/ 1, C 111, P 2/3, M 3/3,
o al 32. The II is la ge han he 12, he la e is
educed in size and esembles Tona ia bidens. The PI
is small, sligh ly displaced o he labial side, low and,
al hough sca cely in con ac wi h he canine, ills
he gap be ween he canines and he p emola s. The
mola s a e simila o hose o Tona ia, bu no iceably
mo e obus . The cusps a e well de eloped, al hough
he p o ocone is ela i ely low in compa ison wi h he
o he cusps. The iI is small; i is displaced an e io ly
in some specimens, hus pe mi ing he canines o be
close o each o he . The p I is sligh ly iangula and
lowe han p3. The p2 is iny and lies be ween pl and
p3, displaced somewha o he lingual side and no is-
ible om he labial side o he mandible because pI
and p3 a e close oge he . The lowe mola s a e na -
owe and ha e no iceably highe cusps han he uppe
ones.
Measu emen s.- Table 2.
Commen s.- In A gen ina he species occu s p i-
ma ily in mois o es s up o 1700 n. In o es ed a eas
abu ing sa annas, i will no be caugh in he open sa-
anna (Del pie o e a1., 1992). I is di icul o cap u e
because o i s abili y o a oid ne s (see also Delpie o e
o
I
200
I
400 km
I
al., 1992), and has o en been conside ed o be a e (e.g.,
B osse and Cha les-Dominique, 1990), al hough i has
been obse ed o ly in small, appa en ly cohesi e,
locks o males and emales (Delpie o e al., 1992).
We o en ha e ound bi ds and ba s (especially S u ni a
liliumy hanging in ba ne s wi h hei heads ea en, al-
hough his species was no obse ed o eed on ne ed
ba s in Co ien es P o ince (Delpie o e al., 1992). The
heigh abo e he g ound a which hey we e en angled
sugges s p eda ion by
Ch o op e us.
Pa k gua ds om
Iguazu Na ional Pa k epo ha Ch o op e us is com-
mon in he a ea and hey obse ed one holding a mouse
(Oligo yzomys) in i s mou h; he ba had bi en he skull
o he oden . Ol og (1973) examined s omachs o
Ch o op e us collec ed in Sal a and Jujuy and ound
lesh, bones, ea he s, and emains o a Ma mosa
(=Thylamys). Yilla-R. and Yil1a-C. (1971) collec ed i e
indi iduals in Sal a P o ince 40 m in o a mine, whe e
hai and skele al emains o mammals, p obably
C enomys, we e sca e ed on he loo . Ba quez and
Gue e o (1985) epo ed he emains o Cu e eb idae
la ae (a common ec opa asi e o mammals) in he gu
o a specimen om Sal a. Ga dne (I 977a) epo ed he
species eeding on insec s and ui s, bu Sazima (1978)
BARQUEZ ET AL.-THE BATS OF ARGENTINA 37
Table
3.
Measu emen s
o Glossophaga soncina, Anou a caudi e ,
and
Ca ollia pe spicilla a.
Fo a
desc ip ion o he measu emen s, see he Ma e ials and Me hods. Mean, s anda d de ia ion, and sample size
a e ollowed by ange.
Va iable
Glossopl aga so/kina Anou a caudije Ca ollia pe spici la a
To al leng h 775 ± 10.54,2
590
± 2.57,5
65.0,
I
700 - 849 560 - 62.4
Tail leng h 70±0.14,2 4.7 ± 035,2
9.0,1
69 - 7.1 45 - 5.0
Hind oo leng h 9.4, I 9.2 ± 080,5 9.0, l
80 - 10.0
Ea leng h 15.9, I 129±2.19,5 15.0, I
110 - 155
Fo ea m leng h 37.4 ± 0.17,3 381 124,5 42.6 ± 115,3
372-375 360 - 39.0 42.0 - 440
Weigh 10.3 ± 076.3 10.4 l.72,3
9.5 - 110 8.5-11.8
Condylobasallcng h 203, I 22.1±051.5 190, I
217 - 230
Leas in e o bi al b ead h 46, I 5.4 ± 0.14,2
53 - 55
Zygoma ic b ead h 90,1 9.9 ± 0.42, 2
9.6 - 10.2
G ea es leng h o skull 21.5, I 22.9 ± 052,5 21.3 084, 2
22.7 - 239 20.7-219
Pos o bi al cons ic ion 46 ± 0.21,5 5.2 ± 025, 3
4.4 - 4.9 50 - 5.5
B ead h o b aincase 90 ± 0.27,5 9.3 ± 028,2
88 - 9.4 9.1 - 95
Leng h o maxilla y oo h ow 73, I 8.3±016,5 5.4±0.14,2
8.2 - 8.5 5.3 - 5.5
Pala al leng h 11.3, I 11.8 ± 0.35, 5
115 - 12.4
Mas oidal b ead h 9.2±0.14,2 106 ± 0.49,2
9.1 - 9.3 10.3 - 110
Leng h o mandibula oo h ow 89±0.21,2 6.9 ± 0.28, 2
88 - 91 6.7 - 71
Leng h o mandible 14.3, I 167±0.49,4 14.1 ± 0.63, 2
16.2 - 173 13.7 - 14.6
C-C (wid h ac oss canines) 4.1 ± 0.19,5 4.7±0.21,2
38 - 43 4.6 - 49
M-M (wid h ac oss mola s) 5.8±019,5 7.6±0.21,2
5 7 - 6.2 7.5 - 7.8
he B azilian caa inga. This species is known o be
p eyed upon by S ygian owls
(Asio s ygius)
in he
ce ado sa anna o B azil (Mo a Junio and Taddei,
J
992)
Addi ional Reco ds.-
BUENOS AIRES: Capi-
al Fede al (Cab e a, 1930). CHACO: Isla del Ce i o
(Fo nes and Massoia, 1967); Resis encia (Pod iaguin,
1944). MISIONES: Misiones (Cab e a, J930); TeyLI
Cua e (Vacca o and Massoia,
1988a).
Specimens Examined
(4 ).- BUENOS AJRES: La
Pla a, J (MLP). JUJUY: Plan a Caimanci o, Pa que
Nacional CaJilegua, 2 (CML). SALTA: Ab a G ande,
JO
kIn
N
O an,
I (CML).

38 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
,,
~"
I -.
,/// " <.
,--~--(--
--------,
,
'
,
,
'
,
_/i'~/~--'-;~-1/
l
"
,
'" I
::,/ j
"
,
o,,' :
'(
,
:
:
'
..•
_,....,.'
:_-----_.
68°
. -;
,
'
,
'
,
'
,
'
'
.•.
_./
Figu e 25. Map o he locali ies o Anou a caudi e .
Genus Anou a
Cab e a (1958) ea ed A. caudi e and A. geo oyi
as congene ic, emphasizing ha he di e ence be ween
hem is he p esence o a ail in caudi e . This cha ac-
e , howe e , could no be used in diagnosing he genus
since i is so a iable ha i can be comple ely absen .
Husson (1962) e ained caudi e in Lonchoglossa and
said ha he zygoma ic a ches a e slende , bu de ini ely
ossi ied. La e , Husson (1978) assigned caudi e o
Anou a. Tamsi and Valdi ieso (1966) also ea ed his
species as Anou a, men ioning he p esence o he zy-
goma ic a ches.
Anou a
caudi e
(E.
Geo oy S .- Hilai e)
Glossophaga caudi e
E.
Geo oy S .-Hilai e,
Me no es du Museum d'His oi e Na u elle, Pa is,
4:418,1818.
Anou a caudi e : Cab e a, Re is a del Museo
A gen ino de Ciencias Na u ales "Be na dino
Ri ada ia," e Ins i u o Nacional de In es igaci6n de las
Ciencias Na u ales, Ciencias Zcologicas, 4:74, 1958.
Sanbo n (1933) conside ed his species o be in
he Genus Lonchoglossa, di e en ia ing i om Anou a
p ima ily by he p esence o a ail. He indica ed ha in
Lonchoglossa he zygoma ic a ch
IS
always p esen ,
whe eas in Anou a i is gene ally lacking. Viei a (1942)
60°
o
I
200 400km
I I
also ecognized Lonchoglossa as a genus in which he
zygoma ic a ches, al hough weak, a e p esen .
In
he
only specimens known om A gen ina, he a ches a e
comple e, bu weak, and in one o hem, om Piqui enda
Viejo, hey we e ound o be comple e, bu upon being
cleaned wi h de mes id bee les, hey disappea ed. This
also happens wi h he ail, which is clea ly isible in
li ing specimens, bu absen in mos museum skins.
Type Locali y.- B azil, Rio de Janei o.
Dis ibu ion.- Two subspecies a e ecognized:
A. c. caudi e om Colombia, Venezuela, he Guianas,
B azil, Boli ia, and A gen ina; and A. c. aequa o is om
Pe u and Ecuado .
Dis ibu ion in A gen ina (Fig.
25).-
Specimens
om Sal a P o ince we e misiden i ied as A. geoJJ oyi
by Fo nes
(1972a),
Ol og and Ba quez (1979), and
Ba quez
(1984a).
Ba quez and Ol og (1985) included
A. caudi e in A gen ina, bu conside ed i he second
species o he genus o he coun y since hey we e
unawa e ha Fo nes
(1972a)
had misiden i ied speci-
mens. Ma es e al. (1989) co ec ly lis ed A. caudi e
o Sal a P o ince. Only A. caudi e is known o he
coun y (Ba quez e aI., 1993).
Desc ip ion (Figs. 26 and 27).- The muzzle is
elonga ed, longe han G.
so icina,
and he uppe bo -
de a he heigh o he eyes IS no ably con ex. The
BARQUEZ ET AL.-THE BATS OF ARGENTINA
39
c
b
Figu e 26. Anou a caudi e . (a) ace; (b) agus; and (c) u opa agium. Anou a geo J oyi: (d)
u opa agium.
ongue is long and p o ac ile. The nose lea is small
bu well de ined (mo e educed han in Glossophaga),
alle han wide, and a ached o he uppe lip. The up-
pe lip is smoo h, lacking w inkles o papillae. The lowe
lip is di ided medially by a deep u ow, wi h wo hick
callosi ies on each side. The ea s a e sho , sepa a ed,
and ounded; he an i agus is absen and he agus is
no mal, sho , and wi hou c enula ions. Pelage is dense
and silky, ex ending along bo h he do sal and en al
sides, o e pa o he plagiopa agium, p opa agium, and
one-hal o he o ea m. The u opa agium is semici cu-
la and bo de ed by a inge o spa se hai s (in
A.
geo oyi i is absen ). The calca is small, sligh ly sho e
han he leng h o he oo . The humb is sho and hin,
and i s nail is sho . The ail is small, included in he
u opa agium, and gene ally ex ends o i s bo de . Colo
is gene ally da k b own; basally he do sal hai s a e g ay
and he en al hai s a e unicolo ed. A eas behind he
ea s, he back o he neck, and an e io one-hal o he
back, a e pale o , in some specimens, eddish. Mem-
b anes a e da k b own o black.
The skull is elonga ed, bu he os um is less han
he leng h o he b aincase and is widened in he a ea o
he canine. Pos o bi al cons ic ion is insigni ican . Zy-
goma ic a ches a e hin and he middle po ion is e-
quen ly ca ilaginous in young indi iduals; in ma u e
specimens he a ches a e ossi ied and comple e. A sag-
i al c es is absen ; a sligh lambdoidal c es is p esen .
Tympanic bullae a e small. Basisphenoidal pi s a e
p esen bu shallow.
The den al o mula is I 2/0, C III, P 3/3, M 3/3,
o al 32. The 1I and 12 a e small, pai ed, and sepa a ed
by a wide space; I I is smalle han 12. P emola s a e
la e ally comp essed and inc ease in size om he i s
o he hi d; PI is educed and sepa a ed om he ca-
nine and P2 by a small space; P2 is sepa a ed om PI
and P3; and P3 is in con ac a i s pos e io bo de wi h
40 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
.
(
..
~
Figu e 27. Skull and ee h o Anou a caudi e . Adul emale om Rio Pescado, Sal a P o ince
(OMNH 18683). Ba
=
1 em. Tee h no o scale.
M I. All p emola s a e iangula in la e al iew excep
o PI; he cen al cusp is ele a ed and sha p, and he
an e io and pos e io cusps a e smalle . The mola s ha e
a s ong cen al dep ession simila o Glossophaga, al-
hough mo e la e ally comp essed. The absence o lowe
inciso s, a diagnos ic den al ai , p o ides a space
h ough which he ongue ex ends du ing eeding. Lowe
p emola s a e hin; pi is almos in con ac wi h he ca-
nine, bu sepa a ed om p2 by a small space. No gaps
sepa a e he emaining p emola s and mola s. The
pa aconid
IS
much smalle han he es o he cusps.
The co onoid p ocess o he mandible is no well de-
BARQUEZ ET AL.-THE BATS OF ARGENTINA
eloped, and is abou he same heigh as he mandibu-
la condyle.
Measu emen s.-
Table 3.
Commen s.-
This species has been sugges ed o
ha e an asynch onous ep oduc i e cycle (Wilson,
1979). A ju enile cap u ed in I au, Sal a P o ince in
Oc obe had deciduous ee h and was jus beginning o
ly. A second indi idual om nea by Piqui enda Viejo
was sligh ly mo e de eloped in mid-No embe , bu he
las mola s had no e up ed comple ely. A abou he
same ime, in Rio Pescado, a p egnan emale was ound
wi h a well-de eloped e us. This species may ha e a
long ep oduc i e pe iod in A gen ina, wi h bi hs oc-
cu ing om Sep embe h ough No embe .
In
F ench
Guiana, B osse and Cha les-Dominique (1990) ound
colonies ha a ied om a ew o 100 indi iduals, many
being nu sing colonies, o be s being ha ems, and all in
associa ion wi h
Ca ol lia
p
e s
p
icil la a
and
Mic onyc e is mic o is .
Li le in o ma ion on die is a ailable; specimens
we cap u ed had emp y s omachs. Ga dne (1977
a)
e-
po ed his species eeding on ui s, nec a , pollen, and
insec s. Sazi na (1976) cap u ed his species (and
A.
41
geo oyis
Visi ing lowe s o
Bauhinia u a
and ound
ha he s omachs con ained pollen, and agmen s o
Insec s o he o de s Thysanop e a, Hymenop e a, Co-
leop e a, and Lepidop e a.
Only ca es, unnels, and ee holes a e known shel-
e s; in ca es hey li e in associa ion wi h o he species
such as
A. geo oyi, A. cul a a, Ca ollia pe spicil la a,
Ca ollia
sp., and
P e ono us pa ne/lii
(Lemke and
Ta nsi , 1979). B osse and Cha les-Dominique (1990)
epo ed i oos ing in cul e s in F ench Guiana in as-
socia ion wi h
Ca ollia pe spicilla a.
One specimen
om Rio Pescado was cap u ed almos a g ound le el
in a ne loca ed a he side o he Rio Pescado in an
open a ea along wi h se e al specimens o
S u ni a
lilium;
he specimen om 6 k n W Piqui enda Viejo
was ob ained o e a s eam wi h calm wa e along one
end o a cul e loca ed unde a oad in he o es .
Specimens Examined
(8).- JUJUY: A oyo
Sauzali o, Pa que Nacional Calilegua,
I
(CML).
SALTA, Agua Blanca, 5 km W,
I
(MACN); hau (Campo
La go), Sie as de Ta agal, 1600 m,
I
(CML); Pa que
Nacional Ban u, nacien e A oyo San elmi a, 900 m,
I
(CML); Piqui enda Viejo, 6 k n W,
I
(CML); Rio
Pescado,3 (2 MACN,
I
OMNH).
SUBFAMILY CAROLLIINAE
Fou species a e known, bu only one occu s in
A gen ina,
Ca ollia pe spicilla a.
This species is cha -
ac e ized by he lack o a zygoma ic a ch, uppe mola s
Genus Ca ollia
Ca ollia pe spicilla a
(Linnaeus)
Vespe ilio pe spici//a us
Linnaeus, Sys ema
Na u ae pe egna ia na u ae, secundum classis,
o dines, gene a, species cum cha ac e ibus, di e en iis,
synony nis, locis. Ten h ed. Lau en ii Sal ii, S ockholm,
1:31, 1758.
Ca ollia pe spicilla a:
Mille , Bulle in o he
Uni ed S a es Na ional Museum, 128:53, 1942.
Type Locali y.-
O iginally, Linnaeus (1758) in-
dica ed "Ame ica" as he ype locali y. Pine (1972) de-
ailed he synonymy, men ioning ha in 1911, Thomas
was in o med by Lonnbe g ha he ype specimen o
Linne om he Museo Adolphi F ide ici had no ex-
is ed o a long ime. Consequen ly, Thomas (191
I)
sug-
wi hou W-shaped
co nmissu es,
and a sho ened, a he
han elonga ed, muzzle.
ges ed ba "Su inam" be gi en as he es ic ed ype
locali y, since he specimen collec ed by Seba p obably
came om ha colony. This ype locali y is now com-
monly accep ed. Ca e and Dolan (1978) also discussed
he designa ion o he holo ype by Thomas.
Dis ibu ion.-
Widely dis ibu ed om sou he n
Mexico o no he n A gen ina. Two subspecies a e cu -
en ly ecognized, C.
p. pe spici//a a,
o e mos o he
ange o he species, and C.
p.
az eca
in Cen al Ame ica
and adjacen no he n Sou h Ame ica. Al hough Pine
(1972) e ised he Genus
Ca o/lia,
subspecies desig-
na ions ha e no been cla i ied. Mos o he' cha ac e s
used o di e en ia e species in his genus ela e o colo
and banding pa e ns o he hai , he deg ee o which
he mandibula inciso s a e hidden by he canines, and
he dis ance be ween P
I
and P2. Such ai s a e highly
42 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
60°
,
,
,
--------------,
, ,
,
.
,
'
, ,
,
.'
/
('
~ -- -- -, J
I
Y ,
" ,'~---~--:
,
,
,
,
,
.
302
')
o
I
200 400km
I I
Figu e 28. Map o he locali ies o Ca ollia pe spicilla a.
a iable h oughou he ange o his species. Specimens
om A gen ina and Pa aguay a e di icul o iden i y,
and i is easy o con use C. pe spicilla a and C.
b e icauda. Pine (1972) summa ized he di e ences be-
ween he wo species. In C.
pe spicilla a,
he quan i y
o hai s on he o ea m and phalanges gene ally is less
han in C. b e icauda. Howe e , in he sou he n po -
ion o i s ange, especially in he d ainage o he Pa ana
Ri e , he hai in C. pe spicilla a is long, s ongly i-
colo ed, and silky, and he phalanges and o ea m a e
qui e hai y. Excep o size
(c.
pe spicilla a is gene -
ally la ge han C. b e icauda), C.
pe spicilla a
is no
eadily di e en iable om C. b e icauda. Specimens
om Pa aguay and no heas e n A gen ina key o C.
b e icauda using he cha ac e s gi en by Pine (1972).
In hese specimens, he cingula o he mandibula ca-
nines do no o ally hide he ou e inciso s, he e is an
e iden space be ween he uppe p emola s, and he
uppe oo h ow is no pe ec ly s aigh . Se e al speci-
mens om Boli ia also ha e a mix u e o hese cha ac-
e s. Ba quez (1977) iden i ied specimens he collec ed
in Boli ia as C.
pe spicilla a,
bu Ande son e al. (1982)
and Ande son (1997) la e indica ed ha 11 we e C.
b e icauda. Mye s and We zel (1983) epo ed speci-
mens om Pa aguay nea A gen ine locali ies ci ed he e,
and p oposed ha he subspecies should be ecognized
as C. p. icolo . Thus he e is some con usion wi h e-
spec o he axonomy o Ca ollia a he sou he n limi s
o he ange o he genus; a axonomic e ision is e-
qui ed o cla i y he axon in sou he n Sou h Ame ica.
Dis ibu ion in A gen ina (Fig.
28).-
Pod iaguin
(1944) men ioned Hemide ma pe spicilla um o he lo-
cali ies o Pue o Be mejo and Resis encia (Chaco) and
Clo inda (Fo mosa), bu did no men ion he au ho , a
p ecise sou ce o he in o ma ion, a ca alog numbe ,
o he museum whe e he specimens we e housed. Con-
sequen ly, he species is conside ed p obable in he p o -
inces o Chaco and Fo mosa.
Desc ip ion (Figs. 29 and
30).-
Medium size;
leng h o o ea m anges om 38 o 46 mm (42-44 mm
in A gen ine specimens). Pelage is so and dense. Hai s
o he do sum a e icolo ed, wi h he base o he hai s
ha ing a wide, da k b own band, ollowed by a ligh e
band, gene ally c eam colo ed (ashy in specimens om
Pa aguay) and wi h he ips ha ing a na ow band simi-
la in colo o he basal one. The gene al colo a ion is
like ha o he ips, bu he o e all aspec is s eaked.
The en e is bicolo ed, a ying in shades o ligh b own
and wi h hai s ha ing a da k basal band o hal hei
leng h. In some Pa aguayan specimens, hai s o he en-
e a e icolo ed wi h an addi ional e y small whi e
band, whe eas in Boli ian specimens he hai s a e
unicolo ed, ligh b own o g ayish b own. The pelage

BARQUEZ ET AL.-THE BATS OF ARGENTINA
Figu e 29. Ca ollia pe spicilla a. (a) ace; (b) agus; (c)
u opa agium; and (d) a icula ion o he o ea m and digi s.
is gene ally es ic ed o he body and ba ely ex ends
o e he do sal and en al side o he plagiopa agium,
o e he p oximal one-hal o he o ea m, and o e he
do sal dis al one-hal o he ea s. The u opa agium is
well de eloped and he ail small, abou 4 mm in leng h
and included in he u opa agium (in a specimen om
I uzaing6 he ail measu es 9 mm); he calca is sho ,
less han he leng h o he oo . Wing memb anes a e
well de eloped, he plagiopa agium a aching a he
43
heigh o he knees. The nose lea is long (abou 10 mm)
and na ow, he muzzle sho , he uppe lip is smoo h,
and lowe IIp has a la ge cen al wa su ounded by
i e o six smalle ones on each side a anged
in
a i-
angle. The ea s a e sepa a ed, longe han wide, and
become na owe owa d he ips, which a e ounded.
The inne su ace o he ea has a se ies o pa allel u -
ows and he an e io ma gin is keeled in some speci-
mens. The agus is well de eloped, na ow a he ip,
and wi h a la e al ema gina ion a he base.
The b aincase is ounded and well de eloped. The
zygoma ic a ches a e incomple e and he sagi al c es
is absen o sligh ly de eloped. The os um is sho bu
in la ed a he heigh o he o bi s, p oducing a p o-
nounced sepa a ion be ween he b aincase and he os-
um. The pala e is wide, ex ends pos e io ly like a ube;
he p e ygoids a e sho and he hamula p ocesses
nea ly each he glenoid ossae. The ympanic bullae
a e small and he basisphenoidal pi s a e deep.
The den al o mula is 2/2, Ill, 2/2, 3/3, o al 32.
The II is well de eloped and sligh ly p ocumben . The
12 is small and a iable in shape, gene ally no com-
ple ely illing he space be ween he canines. In some
specimens, he uppe inciso s a e ounded and small,
while in o he s hey inc ease in size, bu a ely each
he heigh o he cingulum o he canines. The canines
a e e y wide a he base and poin ed a he ip wi h a
well-ma ked conca i y on he in e nal ace. PI is in
con ac wi h he pos e ola e al bo de o he canine, is
unicuspida e, and gene ally in con ac wi h P2. In a
numbe o specimens examined, a space is clea ly e i-
den ; his space is one o he cha ac e s used by Pine
(1972) o di e en ia e C.
b e icauda
om o he
Ca ol/ia. The "W" pa e n o he mola s is no p o-
nounced and he cusps a e di icul o iden i y, al hough
he pa acone, me acone, and p o ocone a e isible in
M1 and M2, bu no in M3, which is educed and a -
anged pe pendicula ly o he oo h ow.
The iI and i2 a e e y small and mayo may no
be sepa a ed; i2 is abou one-hal he size o iI and gen-
e ally hidden by he cingulum o he canines in do sal
iew. In ju eniles, he inciso s a e bilobed, al hough
his condi ion
IS
los wi h wea . P emola s a e o simi-
la size, p I is in con ac wi h he pos e io bo de o he
cingulum o he canine. The m I and m2 a e simila in
size and m3 is smalle . The cusps a e li le di e en i-
a ed, al hough he p o oconid and me aconid a e in close
p oximi y.
44
SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
l
j
FIgu e 30. Skull and ee h o Ca o//w pe spici//a a. Adul emale om I uzaing6, Co ien es
P o ince (MACN 16994). Ba
=
I cm. Tee h no o scale.
Measu el71en s.~ Table 3.
Commen s.- This species may be abundan o e
much o i s ange, especially m dis u bed a eas (B osse
and Cha les-Dominique, 1990; Cha les-Domi nque,
991), bu A gen ine specimens a e sca ce. I was i s
epo ed in A gen ina by Fo nes and Massoia (1969).
Roos s include b idges, cul e s, ee holes, oo s o u-
al dwellings, ca es, and c e ices (e.g., Ma ques, 1985).
Willig (1983) ound i in buildings wi h G/ossophaga
so icina.
I
is colonial o soli a y, and o ages mainly
on ui , bu some au ho s ha e epo ed insec s. Sazima
(1976) ound indi iduals eeding on nec a o Pass i lo a
muc ona a and on he in lo escences o banana, Musa
acumina a, Rep oduc ion is summa ized by Wilson
(1979), who epo ed a bimodal polyes ous pa e n. In
BARQUEZ ET AL.-THE BATS OF ARGENTINA
specimens collec ed in Boli ia, ou emales had open
aginas and one was p egnan (one emb yo, c own- ump
leng h
=
5 nun) a he end o July, al hough no all ba s
collec ed we e b eeding (R, M, Ba quez, pe sonal ob-
se a ion),
Specimens Examined
(8),- CORRIENTES:
I uzaing6, 1 (MACN). MISIONES: Gobe nado
45
Lanusse, 1 (CML); Pa que Nacional Iguazu, I (MACN);
San Ignacio,S (MACN).
Addi ional Reco ds,-
CHACO: Pue o Be mejo
(Pod iaguin,
J
944); Resis encia (Pod iaguin, 1944),
FORMOSA: Clo inda (Pod iaguin,
J
944). MISIONES:
A oyo U ugua-i (Massoia e
al
1987); Cue a Ma ia
An onia (Fo nes e al., 1969),
SUBFAMILY STENODERMATINAE
Some au ho s di ide his sub amily in o
S u ni inae and S enode ma inae, he o me including
only he Genus
S u ni a;
we ollow Jones and Ca e
(1976) in using he single Sub amily S enode na inae
(see also Owen,
J
987), All species in his sub amily
ha e a sho , wide muzzle, lack an ex e nal ail, ha e a
poo ly de eloped o missing
(Slu ni a)
u opa agium,
1.
I',
2 (l ').
2'.
] (2).
]',
4 (2'),
4'.
5 (4).
5'.
6 (4'),
and ha e a iable colo pa e ns.
Pla y hinus
has a pale
mid-do sal s ipe;
Pygode ma
has conspicuous whi e
spo s on he shoulde s and a well-de eloped
u opa agiu n:
S u ni a
has och aceous shoulde pa ches
and lacks a u opa agium;
A ibeus
and
Vampy essa
ha e
acial s ipes, A small bu e iden calca (excep in
S u ni ai,
dis inguishes his sub amily.
KEY TO THE SPECIES
Whi e mid-do sal s ipe p esen .." .."", " .., """ , ", ..""
PIa y h inus linea us
Whi e mid-do sal s ipe absen .", ..", " "", "" """ "" .., ", "" , 2
Facial s ipes p ominen """""" "" .., "" , """ "", .."" " " " , 3
Facial s ipes obscu e o absen ' 4
Size small, leng h o o ea m
<
37 m n; agus and
ma gins o ea yellowish
Vampy essa pusilla
Size la ge, leng h o o ea m usually> 70 m n; agus and ma gins o
ea no yellowish , "" "" .." """ ,..", ", "",
A ibeus li u a us
Shoulde s usually wi h yellow, b own, o ange, o whi e pa ches;
u opa agium absen bu , i p esen , co e ed wi h hai s on he do sal
su ace; leng h o o ea m
<
50 mm , 5
Pa ches on shoulde s absen ; u opa agiu n naked; leng h o o ea m
> 50
l11J11,
bu
<
70
l11J11 "", , " , '",' " .. , ' ",
6
Whi e pa ch on each shoulde ; do sal su ace o u opa agium densely u ed;
ex e nal ma gin o he ea s, agus, and base o he nose lea yellowish;
a well-de ined old o skin ex ends om he en al ma gin o he nose lea
o he angle o he mou h, appea ing like a "double uppe lip"; uppe oo h ows
bowed and almos semici cula ; 28 ee h
Pygode ma bilabia um
Shoulde s wi h cinnamon pa ches, some imes no easily isible; u opa agiu n
absen ; inne ma gin o legs and ump densely u ed; uppe oo h ow somewha
bowed bu no o ming a semici cle; 32 ee h , 7
Leng h o o ea m 62-69 m n; condylobasal leng h> 25 mm; acial s ipes
weak, M3 small bu p esen ; M2 abou 3.6 m n wide; a p esen known only
o he p o inces o he no hwes o he coun y """ '"., ", , ',.
A ibeus plani os is
46 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
6'. Leng h o o ea m 64-67 mm; condylobasalleng h > 27.9 nun; acial s ipes
weak o absen ; M3 absen (p esen in a low pe cen age); wid h o M2 > 4.0 nun;
a p esen known only o he p o inces o Misiones and Chaco A ibeus imb ia us
7 (5'). Lingual idge o lowe mola s no se a ed; lowe inciso s usually bilobed 8
7'. Lingual idge o lowe mola s se a ed; lowe inciso s usually ilobed S u ni a lilium
8 (7). Size la ge, leng h o o ea m> 44 mm; uppe oo h ow almos pa allel; middle
uppe inciso s ending wi h a s aigh ma gin, some imes ligh ly bilobed S u ni a opo aphilum
8'. Size smalle ; leng h o o ea m
<
43 mm; uppe oo h ow sligh ly cu ed; middle
uppe inciso s somewha ounded S u ni a e y h omos
Genus
S u ni a
The e a e 12 species in his genus in he
Neo opics. Li e a u e eco ds o his genus in A gen-
ina e e ed o only one species, S u n i a lilium, un il
analysis o collec ion ma e ial pe mi ed he di e en-
ia ion o h ee o ms,
S.
lilium,
S.
e y h omos, and
S.
opo aphilum (Ba quez e al., 1993; Ma es e al., 1995).
S.
lilium is he mos common;
S.
e y h omos is appa -
en ly e y common; and
S.
opo aphilum is sca ce bu
widely dis ibu ed. The h ee species a e sympa ic in
no hwes e n A gen ina.
S.
e y h o mo s and
S.
opo aphilum p e iously we e known o occu only as
a sou h as Boli ia (Ande son, 1997), whe e hey ap-
pa en ly a e ound abo e he al i udinal limi s o
S.
lilium, wi h which he e is limi ed o e lap. The h ee
species a e e y simila , bu can be eadily di e en i-
a ed mo phologically o gene ically (e.g., Pacheco and
Pa e son, 1991).
S u ni a
e y h omos (Tschudi)
Ph(llos oma) e y h omos Tschudi, The ologie.
Un e suchungen ibe die Fauna Pe uana. Schei lin und
Zolliko e , S . Gallen, Swi ze land, p. 64, 1844.
S u ni a e y h omos, de la To e, Unpublished
Ph.D. disse a ion,Uni e si y o lllinois, Champaign, p.
124, 1961.
This species equen ly has been synonymized
wi h
S.
lilium. Howe e , since de la To e (1961), mos
au ho s ha e conside ed i dis inc .
Type Loca/i y.- Pe u.
Dis ibu ion.- Un il he e ision by Jones and
Ca e (1976), his species was known only in Pe u om
eas o he Andes. Koopman (1993) ga e i s dis ibu-
ion as ex ending om Venezuela o Boli ia. Ande son
e a1. (1982) epo ed he sou he n ange limi s in Bo-
li ia as Ta ija Depa men on he bo de wi h A gen-
ina.
Dis ibu ion in A gen ina (Fig. 31).- Known only
o he p o inces o Sal a, Jujuy, and Tucu nan, bu ap-
pa en ly no limi ed o high ele a ions as some speci-
mens ha e been cap u ed a he limi s o sub opical
o es and chaco ho nsc ub below 500 m. Ande son e
al. (1982) epo ed specimens om locali ies below
1000 m, bu mos we e collec ed abo e 1285
ill.
Desc ip ion (Figs. 32, 33a, 34a, and 35a).- This
is he smalles species o he genus ound in A gen ina,
wi h he leng h o o ea m anging om 38.7 o 43.0
nun ( ew indi iduals ha e measu emen s la ge han
he smalles
S.
liliumi. The skull is always smalle han
in o he S u ni a; maximum g ea es leng h o skull e-
co ded was 21.4 nun. Ex e nal appea ance is simila o
he o he species, bu he bases o he do sal hai s and
he gene al colo a ion o he body a e da ke . Wi h e-
spec o
colo ,
i can be con used only wi h
S.
opo aphilum, bu he pelage is pale and he bands o
he hai s a e only sligh ly ma ked o absen . The in en-
si y o he colo o he bands is a iable, howe e .
In
mos o he specimens, he basal one-hal is da k b own,
ollowed by an ash g ay o whi ish band o abou 3 nun,
and a small ip o da k b own, g ay, o almos black,
abou I o 2 mm.
In
some specimens om Tucuman
P o ince, he e minal colo band is no p esen in many
o he hai s so ha he gene al colo o he middle band
domina es, gi ing hem a pale aspec .
In
ju eniles, he
bands o he hai a e no eadily dis inguishable and a e
gene ally e y da k. All bu h ee specimens ( om Playa
La ga, Tucu nan P o ince) lacked he och aceous shoul-
de .
BARQUEZ ET AL.-THE BATS OF ARGENTINA
a
.(
·:
-z-
'i>
'" '~ , .
.,
..~ . ',' I : .
I I
." /y"
Figu e 37.
S u ni a lilium:
(a) ace; (b) hindqua e s; and
(c) agus.
bu some a e pale , gene ally g ay. One o he mos im-
po an cha ac e s o iden i ying his genus is he ab-
sence o he u opa agium and ail. The hai ex ends o
he p oximal one-hal o he o ea m on bo h he do sal
and en al sides, and on he do sal side o he legs.
Hai s a e spa sely sca e ed o e he plagiopa agium
along he sides o he body, in he bo de whe e his
memb ane a aches o he ankles, and o e he en al
su ace o he p opa agiu n. The snou IS sho . Ea s a e
small and widely sepa a ed, somewha iangula in
shape, and wi h he dis al one-hal o he ou e ma gin
s aigh and he p oximal one-hal con ex; he an i a-
gus is poo ly de eloped. The agus is well de eloped,
iangula , becomes hinne owa d he ip, and has a
small, bu well-de ined basal conca i y. The nose lea
is sho , abou 8 mm, e y wide a he base, na owe
owa d he ip, and lanceola e in o m. A se ies o glands
bo de s he basal po ion o he nose lea owa d he
sides and on o he eyes. The uppe lip and he nose
lea a e sepa a ed by a space. The ma gin o he lip is
53
bo de ed by small papi llae ha a y in numbe . The
lowe lip has a g oup o papillae, wi h a la ge cen al
one su ounded by smalle ones on bo h sides; hei
numbe and loca ion a e a iable. The calca is absen .
The b aincase is in la ed and ele a ed; he leng h
o he os um is less han he leng h o he b aincase;
zygoma ic a ches a e comple e bu weakly de eloped.
A sagi al c es is p esen , bu no well de eloped, pa -
icula ly in emales andju eniles.Incisi e o amina a e
e y la ge. The pala e p ojec s pos e io ly like a ube,
wi h he hamula p ocesses o he p e ygoids ending a
he le el o he glenoid ossa. The caudal spine o he
pala e is undi e en ia ed, gi ing he pos e io bo de
o he ha d pala e he aspec o a con inuous, unin e -
up ed line. Basila pi s a e shallow. Ty npanic bullae
a e small, co e ing only he an e ola e al pa o he
cochlea.
The den al o mula is 12/2, C 1/1, P 2/2, M 3/3,
o al 32. The uppe Inciso s comple ely ill he space
be ween he canines; II is abou wice he size o 12 and
in con ac wi h i a he base;
II
ba ely exceeds he heigh
o he cingulum o he canine. The uppe inciso s p ojec
medially and ha e a ma ked conca i y, like a u ow,
on he an e io su ace. Canines a e obus , bu ela-
i ely sho and wi h a shallow conca i y along hei
in e nal ace. P emola s a e simple, wi h a single cusp
and a conca i y owa d he lingual side; PI is iangu-
la in shape when iewed do sally and is smalle han
P2; P2 is squa e and highe han he o he mola i o m
ee h. M I and M2 ha e a ba ely isible pa acone and
me acone. The o he cusps a e no e iden and p oduce
a longi udinal u ow on he occlusal su ace o each
mola , he u ow educed o es igial in many indi-
iduals. All mola s a e squa e o almos squa e, and
dec ease in size om M 1 o M3. The p o ocone o he
mola s is elonga e and o ms he inne bo de o he
a o emen ioned u ow.
The i 1 and i2 a e small, subequal in size, ilobed,
and comple ely ill he space be ween he canines. The
pos e io pa o he cingulum o he canine is in con-
ac wi h pi; p I is la ge han p2, he e e se o he
uppe p emola s. The pa aconid o ml is low and p oc-
umben ; he me aconid and en oconid a e also low bu
mo e di e en ia ed, so ha when iewed la e ally he
in e nal su ace o he mola s appea s se a ed ( his is
no obse ed in he o he wo A gen ine species). The
small pa aconid and me aconid o m2 a e in close p ox-

54 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
Figu e 38. Skull and ee h o S u n i a lilium. Adul male om Piqui enda Viejo, Sal a P o ince
(OMNH 18736). Ba
=
I e n. Tee h no o scale.
i ni y; he en oconid is well ma ked; he p o oconid and
hypoconid a e almos absen . The m3 is small, wi h less
han one-hal he occlusal su ace o m2, and wi h he
cusps gene ally undi e en ia ed.
Measu emen s :-: Table 4.
Commen s.-
Mos specimens we e cap u ed in
he lowe s a a o o es s, wi h many apped in mis
ne s a g ound le el. We ha e obse ed hem lying in
g oups along de ined ou es o eed up o 10 km away
om hei shel e s. B asse and Cha les-Dominique
(1990) epo ed ha his species was ne ed in dis u bed
BARQUEZ ET AL.-THE BATS OF ARGENTINA
habi a s in F ench Guiana. This species appea s o be
uncommon in u ban a eas in A gen ina; howe e , skulls
o ju enile and subadul s we e ound in ba n owl (Ty o
alba) pelle s, whe e he owls we e nes ing in buildings
in down own Tucu nan ci y (R. M. Ba quez, pe sonal
obse a ion). Whe he o no he owls we e o aging in
he ci y, o in o es s 20 km away, is unknown. Owls
ha e been shown o cap u e a wide a ay o ba s in he
Neo opics (Mo a Junio and Taddei, 1992).
Rep oduc i e ac i i y in A gen ina is g ea es om
Oc obe o Decembe , wi h mos bi hs occu ing a he
end o No embe . Al hough uncommon, some ep o-
duc i e ac i i y occu s h oughou he yea ; a emale
was ound lac a ing in Ap il (la e all) in he A oyo
Los Noques, Pa que Nacional El Rey, Sal a P o ince.
In F ench Guiana, his species was shown o b eed om
June o Augus (B osse and Dubos , 1967).
S u ni a lilium eeds p ima ily on ui , al hough
hey also may eed on pollen and nec a .
Specimens Examined (420).- CATAMARCA:
Cues a del Cla illo, 5 km S La Bande i a, I (RMB); La
Me ced, 3 ( I MACN, 2 TTU). CHACO: Es ancia San
Miguel, along Hwy 90, 15 km NW je . Hwy 90 and
Hwy 11, 6 ( 3 CML, 2 IADIZA-CM, 1 OMNH);
Resis encia, I (MACN). CORRIENTES: Laguna Pai a,
Bo Las Lomas, 4 (3 CML, I OMNH). FORMOSA:
Clo inda, 1 (TTU); Es ancia Las Me cedes, 1 (TTU);
Es e o de los Pa os, 5 (TTU); Rio Po e io, 5 km S
Es ancia San a Ca alina, 1 (CML). JUJUY: Ab a de
Canas, app. 30 km NW Calilegua, sob e u a a Valle
G ande, 2 (CM); Aguas Blancas, 14 km E San a Cla a,
4 (OMNH); Aguas Neg as, Pa que Nacional Calilegua,
15 (14 CML, 1 RMB); A oyo Sauzali o, Pa que
Nacional Calilegua, 2 (CML); Camino de Comisa, 10
km W Dique La Cienaga, I (CML); El Du aznillo, 3000
m, Ce o Calilegua, 1 (CML); El Simbola , 25 km SW
Palma Sola, 5 (CM); F aile Pin ado, 12 km W, sob e
Rio Ledesma, 1 (MD); Laguna La B ea, 12 (OMNH);
Laguna La B ea, 25 km W Palma Sola, 13 (CML);
Vinali o, 2 km al NW, al bo de del segundo canal, I
(MD); Yu o, 3 (AMNH). MISIONES: A is 6bulo del
Valle, 10 km W by oad along Rio Cu iapi u, 2 (1 CML,
1 IADIZA-CM); Colonia Ma i es, 5 (I MACN, 4
TTU); Gobe nado Lanusse, 4 (CML); Iguazu, 56
(MACN); Misiones (no speci ic locali y), I (TTU);
Misiones, 200 m (no speci ic locali y), 2 (BMNH);
Mon eca lo, 2 (FMNH); Pa aje Pa a iso, 16 km SE, sob e
55
u a 21, 5 (CML); Pue o Pi ay, Ru a 12, 1 (MACN);
San Ped o, 47 km SE, 18 (15 CML, 3 RAO) SALTA:
Agua Blanca, 20 km NW, 2 (TTU); Agua Blanca, 24
km NW, 11 (CM); Agua Blanca, 25 km NW, 5 (TTU);
Agua Blanca, 5 km W, 4 (3 MACN, 1 TTU); Agua ay,
2 (MACN); Aguas Blancas, 4 (CML); Al o Macue a, 2
km N del c uce de Macue a y Campo Lago, 1 (AA);
A oyo Los Noques, Pa que Nacional El Rey, 15 (CM);
Finca El A azayal, Se ania de las Pa as, 2 (I CML, I
OMNH); Ingenio San Ma in del Tabacal, 3 (TTU); I au
(Campo La go), Sie as de Ta agal, 1600 m, 3 (CML);
Jun a de
los
Rios Ta ija y Be mejo, 4 (CML); Jun as de
San An onio, 17 (CM); Me an, 850 m, 1(BMNH); O an,
20 km al NW, 9 (MACN); Pa que Nacional Ba i u, Finca
Jakulica, Angos o del Rio Pescado, 650 m, I (CML);
Paso de la C uz, 2 (CML); Piqui enda Viejo, 30 km N
Ta agal, 19 (11 CM, 8 OMNH); Piqui enda Viejo, 6
km W, 18 (4 CML, 14 OMNH); Queb ada de
Acambuco, 5 km W Dique I iyu o, I (RAO); Rio EI
Na anjo, 14 km W de Ru a p o incial 5, I (CML); Rio
Pescado, 21 (TTU); Tabacal, 4 (3 MACN, 1 TTU);
Tonono, I km E sob e Rio I iyu o, 8 (CML); Unchime,
I (ROM); Vado de A azayal, 20 km NW Aguas
Blancas, 6 (CML). SANTA FE: San a Fe, I (MFA).
TUCUMAN: Agua Colo ada, 11 (9 MACN, 2 TTU);
A oyo El Sal 6n, Rese a P o incial San a Ana, I
(CML); Concepci6n, I (BMNH); Dique San Ignacio, I
(OMNH); El CadiIlal, Es aci6n de Piscicul u a, 1
(OMNH); El Na anjal, I (TTU); Ho co Molle, 15 km
W San Miguel de Tucuman, 7 (CM); Ho co Molle, 900
m, 2 (CML); Ho co Molle, Rio Las Le ias, 2 (CML);
Ho co Molle, Rio Las Pied as, 25 (CML); Ingenio San
Pablo, I (MACN); Los Sa mien os, 1 (CML); Pa que
P o incial El Cochuna, km 40 sob e u a 47, 4 (CML);
Pied a Tendida, 5 km W de Dique EI Caj6n, 1 (CML);
Playa La ga, Rio Los Sosa, Ru a 307 km 19.7, 6
(OMNH); Rese a P o incial "Aguas Chiqui as" El
Cadillal, I (CML); Rio Los Sosa, Ru a 307, km 23.9,
camino a Ta i del Valle, 850 m, I (OMNH); Rio Los
Sosa, Ru a 307, km 24.9, camino a Ta i del Valle, 850
m, 2 (OMNH); Rio Pueblo Viejo, Rese a P o incial
La Flo ida, I (CML); San Miguel de Tucuman, 1
(BMNH); San Ped o de Colalao, I (CML); Tucu nan, 1
(FMNH). NO SPECIFIC LOCALITY: Pa agonia, 1
(BMNH).
Addi ional Reco ds.- CHACO: Resis encia,
ma genes del Rio Pa ana (Ro na ia and Abalos, 1950).
ENTRE RIOS: Conco dia, 32 km N (Ba quez and
Lougheed, 1990). FORMOSA: Pa que Nacional Rio
56
SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
Table 4. Measu emen s o S u ina e y h o nos, S umi a liliu n. and S umi a opo aphilu n. Fo a de-
sc ip ion o he measu emen s, see he Ma e ials and Me hods. Mean, s anda d de ia ion, and sample size a e
ollowed by ange.
Va iable
S u ina e y h omos S u ni a {ilium S u ni a opo aphilum
To al leng h 55.3 ±
3A9, 36
621 i 6.31, 110 67.1 ± 13.73,3
50.0 - 630 50.0 - 90.0 585 - 83.0
Hind oo leng h 109 i 199,36 116±IAI,104 9.5 ± 0.50,3
8.0 - 68 88 - 15.7 90-10.0
Ea leng h 168
±
.36,35 66
±
2.34, 09 17.8iO.77,3
13.0 - 18.7 110 - 20.0 170-18.5
Fo ea m leng h 41.1il.02,35 43.5 ± 1.86, 113 44.7 i 1.15,3
38.7-43.0 392 - 49.0 44.0 - 46.1
Weigh 16.0
±
2A5,
33 21.5
±
2.62, 71 216 i 2.08, 3
12.0-23.0 140-280 20.0 - 24.0
Condylobasal leng h 18.7 ±
OA6, 35 20A
± 0.51, 110 20.6 i 0.36, 3
18.0-197 19.5 - 217 20.3 - 21.0
Leas in e o bi al b ead h 6.2 i 0.36, 19 6.3 ± 0.28, 43 6.5 ± 0.34, 3
5A -
6.9 5.6 - 6.9 63 - 6.9
Zygoma ic b ead h 12.6 ± 0.53,26 13.6 ± 0.73, 78 3.8 ± 0.55, 3
III - 13.6 10.0 - 14.8 13.5 - 14.5
G ea es leng h o skull 20.5 ±
OA2, 36
22.5 ± 0.54,112 22.6
±
0.60, 3
196 -
21A
210-23.9 22.0 - 232
Pos o bi al cons ic ion 5.8 ± 0.18,35 60 ± 0.21, II I 6.1
±
0.11,3
55 - 6.2 55 - 6.5 6.0 - 6.2
B ead h o b aincase 9.8 ± 0.23, 35
lOA
± 0.23, II I
lOA
i 0.25,3
9A -
105 9.7 - 11.1 10.2 - 10.7
Leng h o maxi
lla y
oo h ow 5.7 ± 0.21, 34 65 ± 0.23, 108 6.5 ± 0.20, 3
5.1 - 6.2 5.9 - 7.2
6A -
6.8
Pala al leng h 8.2 ± 0.38,35 93
±
OA5, 05
9.1 ±
OA
1,3
7A - 9A
7.9-11.5 8.8 - 9.6
Mas oidal b ead h 11.3 ± 0.23, 22 12.1 ±
OAO, 75
119 ± 028, 3
110 - 11.9 11.0-12.9 11.8 - 2.3
Leng h o mandibula oo h ow 58 ± 0.60, 34 6.7±0.69,107 73 ± 0.25, 3
50 - 69 5.1 -
8A
7.1 - 7.6
Leng h o mandible 13.3 ± 0.34, 32 148 i 0.76,112 14.9 ±
OA5,
3
12.6 - 14.2 8 I - 15.8 145 -
15A
C-C (wid h ac oss canines) 5.5iO.18,23 6.1 ± 0.15,69 5.8 ±
OA5,
3
5.3 - 60 5.7 - 6.8
5A -
6.3
M-M (wid h ac oss mola s) 7.3 ± 0.23, 21 80 ± 0.30, 68 80 ± 0.35, 3
6.9 - 80 71 - 87
7.7-8A
Pilco nayo (S. Heinonen F. and R. M. Ba quez, in li .).
JUJUY: A oyo La U bana, 45 km E y 5.4 k n SE San
Sal ado de Jujuy, 620 m (Villa-R. and Villa-C., 1971);
Finca EI Re na e, 24 km SE San Sal ado de Jujuy, 740
m (Villa-R. and Villa-C., 1971). MISIONES: Apos oles
(Massoia e aI.,
1989a);
A oyo U ugua-i (Massoia e
al., 1987); Depa amen o Cainguas (Massoia, 1980);
Depa amen o Capi al (Massoia, 1980); Escuela P o-
incial 639, Campo Ramon (Massoia, 1988a); Gua ani
(Massoia, 1980); Los Helechos, Escuela n° 12 (Massoia
e al., 1989b). Rio NEGRO: Rio Neg o (no speci ic
locali y) (Cab e a, 1958). SALTA: Seclan as (Yepes,
1944). TUCUMAN: Rio Los Sosa, Ru a 307, km 19.7,
camino a Ta i del Valle, 700 m (Ma es e aI., 1996) (=
Playa La ga, Rio Los Sosa, Ru a 307 k n 19.7); Rio Los
Sosa, Ru a 307, km 19.7, camino a Ta i del Valle, 750
m (Ma es e aI., 1996)
(=
Playa La ga, Rio Los Sosa,
Ru a 307 km 19.7).
,,
,--
:
"",
,/,'/i -,
1/~------~-------------- 1
...,-...., :....
"c , ....,,--,/ :
'-':'"530. : :
'
....
,:
:
"
,
"
,
/ .123/
i
~,/
:
,
~
:
,
,
,
-,..... --' ~---------
BARQUEZ ET AL.-THE BATS OF ARGENTINA
68
0
;"L
-:>
Figu e 39. Map o locali ies o S u ni o opo aphilum.
S u ni a opo aphilum (Tschudi)
Ph(yllos oma) opo aphilum Tschudi, The ologie.
Un e suchungen ube die Fauna Pe uana. Schei lin und
Zolliko e , S . Gallen, Swi ze land, p. 64, 1844.
S u n i a opo aphilum: de la To e, Unpublished
Ph.D. disse a ion, Uni e si y o Illinos, Champaign,
p. 112,1961.
Type Locali y.- Pe u.
Dis ibu ioa--: Acco ding o de la To e (1961),
his high-ele a ion species occu s om sou he n Pe u
no hwa d in o Cen al Ame ica and Mexico as a no h
as he s a e o San Luis Po osi. In a s udy o a la ge
se ies o specimens om Boli ia and om egions o
he no he n Andes (Ecuado , Colombia, and wes e n
Venezuela), including he ypes o
S l
u
do ici ;
S.
bogo ensis, and S opo aphilum, Ande son e al. (1982)
esol ed he con usion o e he axonomy o hese spe-
cies om wes e n Sou h Ame ica. They no ed ha he
cha ac e s o he specimens om Boli ia a e simila o
hose o he ype o
S
opo aphilum.
Honacki e al. (1982) included his species as a
synonym o
S
ludo ici, bu lis ed eco ds om Boli ia
57
60
0
o
I
100 200
I I
400 km
I
as p obably pe aining o S. bogo ensis . Koopman
(1993) ecognized he speci ic s a us o S bogo ensis,
bu men ioned ha he co ec name was likely S.
opo aphilum, ci ing Ande son e al. (1982). E en hough
he iden i y o he di e en species in no he n Boli ia
is no comple ely clea , he specimens om A gen ina
a e iden ical wi h hose om Boli ia housed in he
AMNH and iden i ied as
S
opo aphilum.
Dis ibu ion in A gen ina.- Figu e 39.
Desc ip ion (Figs. 33c, 34c, 35c, and 40).- Gen-
e ally simila o o he species o he genus. La ge han
S e y h omos and la ge han S lilium o e mos o he
la e 's dis ibu ion, including A gen ina. Leng h o o e-
a n anges om 44 o 46.2 mm in he specimens ex-
amined. Cinnamon shoulde pa ches a e absen . Unlike
S lilium, he do sal hai s appea icolo ed, wi h a wide
da k g ay o b own basal band ollowed by a pale one
o simila size. The e ninal band is na ow and he
same colo as he basal po ion. As in S e y h omos,
he lingual ma gin o he lowe mola s lacks e ical
di isions, and he lowe inciso s gene ally a e bilobed,
al hough hey may be sligh ly ilobed. The skull is simi-
la o S lilium. The s aigh uppe oo h ows uniquely
iden i y his species o S u ni a ( hey a e s ongly cu ed
58
SPECIAL PUBLICATlONS, MUSEUM OF TEXAS TECH UNIVERSITY
,
Figu e 40. Skull and ee h o S u u a opo aphilum. Adul emale om A oyo
Sauzali o, Pa que Nacional Calilegua, Jujuy P o ince (CML 2932). Ba =1 em.
Tee h no o scale.

BARQUEZ ET AL.-THE BATS OF ARGENTINA
60°
28°
,
,
-.
,
"
/<i
o
o
o
,
'
-------------------7
:
,,'
/~-l~
..._--------~y;
"~'I ,~;:'-'-----,-
-'I,
Figu e 41. Map o he locali ies o Pla y hinus linea us.
in he o he species). The os um is na owe han in
o he species o he genus and mo e in la ed a he o -
bi s. The an e io ma gin o he uppe inciso s is ei he
s aigh o , in some specimens, sligh ly bilobed, unlike
he o he species in which he an e io ma gin o he
inciso s is ounded.
Measu emen s.- Table 4.
Commenls.- The specimen om Tucu nan P o -
ince was ob ained in Janua y and had sc o al es es; a
emale om Se ania de las Pa as, Sal a P o ince, was
lac a ing in mid-No embe .
Specimens Examined
(7).-
JUJUY: A oyo
Sauzali o, Pa que Nacional Calilegua, 2 (CML); El
Monoli o, I (CML); F aile Pin ado, 12 km W, sob e
Rio Ledesma, 1 (MD); Laguna La B ea, 1 (OMNH).
SALTA: Se an ia de las Pa as, I (OMNH).
TUCUMAN: Casa de Pied a, Rio Los Sosa, u a 307,
k n 24.9,850 n, 1 (OMNH).
59
o
I
100 200 400 km
I I I
Addi ional Reco ds.- TUCUMAN: Pied a
Tendida, 12 km WNW Bu uyacu along Rio Cajon,
2,500 . (Ma es e al., 1996).
Genus Pla y hinus
Pla y hinus linea us (E. Geo oy S .- Hilai e)
Phy//os oma /inea um
E.
Geo oy Sl.- Hilai e,
Annales du Museum d'His oi e Na u elle, Pans, 15:180,
1810.
Pla y hinus /inea us: Saussu e, Re ue e Magasin
de Zoologic, se ie 2,12:429,1860.
Jones and Ca e (1976) conside ed his species
o be mono ypic, bu Koopman (1976) included
Vampy ops
(=
P/a y hinus) nigel/us as a subspecies o
P linea us based on c anial simila i ies. La e , Ande -
son e al. (1982) lis ed bo h subspecies
(P I. linea us
and P I. nigel/us) in Boli ia whe e hey do no occu in
sy npa y Ga dne and Fe ell (1990) cla i ied he no-
60 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
a
I .....
c ~~
.:
.1;;~;.,. ' '
.,
;:'
Figu e 42.
Vampy ops linea/us:
(a) ace; (b) do sal iew;
and (c) agus.
nencla u al s a us o his species. The A gen ine sub-
species is
P l. linea/us.
TypeLocali y.-
Pa aguay, Asuncion.
Dis ibu ion -«
This species is ound om in
no he n Colombia, F ench Guiana and Su iname
h ough Ecuado and cen al Pe u o no h-cen al Bo-
li ia, hence o he B azilian coas and sou hwa d o
eas e n Pa aguay, no heas e n A gen ina, and no he n
U uguay (modi ied om Koopman, 1993).
Dis ibu ion in A gen ina.-
Figu e 41.
Desc ip ion
(Figs.
42
and
43).-
Size is medium,
sligh ly la ge han S.
lilium,
wi h he leng h o he o e-
ann be ween 45 and 50 mm. The p ominen mid-do sal
s ipe is no ound in any o he A gen ine phyllos omid.
Only
Noc ilio
has a simila s ipe.
In
Pla y hinus
he
s ipe is pale colo ed, gene ally whi e, and ex ends om
be ween he ea s o he base o he ail.
In
some speci-
mens i is e y pale and no clea ly e iden , bu i is
always p esen . Two pai s o acial s ipes a e no mally
p esen . The sup ao bi al s ipe ex ends om he uppe
side o he nose lea o he in e nal ma gin o he ea .
The subo bi al s ipe ex ends om he pos e io ma -
gins o he lips o he ea . The specimens om Chaco
P o ince a e ligh b own, wi h he ips o he hai s
sligh ly da ke , he en e pale , and he hai s unicolo ed.
Specimens om Pa aguay, Depa amen o Pa agua i, a e
sligh ly g aye and he do sal line is immacula e whi e,
whe eas in hose om Chaco P o ince i is yellowish
o c eam. The eyes a e su ounded by da k, almos black,
hai s p oducing a cha ac e is ic eye ing. The agus is
yellow; in F ench Guiana, B osse and Cha les-Domi-
nique (1990) epo ed he agus colo as pink. The mem-
b anes a e e y da k b own o black. The hai ex ends
do sally a li le beyond he p oximal hal o he o e-
a m, o e one-hal o he p opa agium, o e he ibia,
and o e he u opa agium. Ven ally he hai is dis ib-
u ed as on he do sum, bu comple ely co e s he
p opa agium and sligh ly mo e o he o ea m and he
plagiopa agium. The nose lea is o medium size, wi h
he basal ma gin ee and su ounded by small glands;
i is lanceoJa e wi h a widened median keel. The ea
has a con ex inne ma gin; he uppe hal o he ex e -
nal ma gin is s aigh and he lowe hal is con ex and
ends in a small an i agus ha is wide han i is all.
The calca and u opa agium a e educed; he
u opa agium is bo de ed by a dense inge o hai s.
The e is no ail.
The skull is obus and, in p o ile, is ele a ed in
he mid-b aincase, whe e he sagi al c es is e iden ;
o e he es o i s leng h he sagi al c es is less p o-
nounced. The os um is sho , i s leng h li le mo e han
one-hal o he leng h o he b aincase. Zygoma ic a ches
a e well de eloped, hin, bu wi h a pa ao bi aJ p ocess
sligh ly widened on he pos e io one-hal . The pos o -
bi al cons ic ion is wide and almos equals he wid h
o he leas in e o bi al cons ic ion. The pala e is na -
ow an e io ly and widened a he le el o he las no-
BARQUEZ ET AL.-THE BATS OF ARGENTINA
I.
61
Figu e 43. Skull and ee h
Pla y hinus linea us.
Adul emale om Resis encia, Chaco P o ince
(CML 1813). Ba
=
I cm. Tee h no o scale.
la s; i s pos e io ma gin is smoo h, a ched, and wi h-
ou a caudal spine. The hamula p ocess o he p e y-
goids is hin. The basisphenoidal pi s a e shallow and
almos indis inguishable.
The den al o mula is I 2/2, C 1/1, P 2/2, M 3/3,
o al 32. The II is well de eloped and mo e han wice
he size o 12, which is educed o a ew ba ely isible
simple spicules. The i s uppe inciso s a e sepa a ed
a he base bu in con ac a he ips. The 12 is sepa a ed
om bo h 1I and he canines, bu in specimens om
a eas o he han A gen ina can be in con ac wi h hem
and ill he in e ening space. The canines a e no mal,
62
SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
sligh ly ecu ed, and sha ply poin ed. The P I is smaJl,
sligh ly mo e han one-hal he size o P2, and wi h i s
an e io mos poin ex ending o e he pos e io p ojec-
ion o he cingulum o he canine. The P2 is sepa a ed
om P I by a small bu isible space; he pa acone is
well de eloped and he mesos yle is low, gi ing he ap-
pea ance o a spicule in he pos e io labial egion o
he p emola . The M I and M2 a e well de eloped; M3
is educed. The cusps o M I and M2 a e la e ally com-
p essed, gi ing he appea ance o a cu ing blade di-
ided in o wo lobes (one an e io and he o he pos e-
io ) whose poin s co espond, espec i ely, o he
pa as yle and me as yle. The p o ocone and he hypo-
cone a e g ea ly educed. The i1 and i2 a e small and
sligh ly bilobed; a smaJl space sepa a es he i s lowe
inciso s; i2 is in con ac wi h he lowe bo de o he
cingulum o he canine. The canines a e no mal. The
p I is smalle han p2. The m 1 and m2 a e e y low
c owned. The an e io edge o m I is poin ed, wi h he
cusps o he igonid g ouped oge he ; pos e io ly, only
he en oconid is isible. The an e io ma gin o m2 is
s aigh due o he alignmen o he p o oconid and me a-
conid and he posi ion o he en oconid on he lingual
side o he pos e io ma gin o he mola . The m3 is
g ea ly ed uced and lac ks di e en ia ed cusps.
Measu emen s.- Table 5.
Commen s.- Al hough he die o P linea us in-
cludes p ima ily ui , Ruschi (1953) ound mo hs o
he Family Sphingidae in s omachs. Sazima (1976) ob-
se ed his species eeding on nec a o he lowe s o
Musa acumina a and La oensia paca i. WiJlig (1983)
no ed a s ong endency o eed on he ui o Vismia.
Males o en main ain small ha ems o 7 o 15 emales,
and each ha em occupies a di e en oos (Willig, 1983).
Li le is known abou ep oduc ion. Gonzalez and
Vallejo (1980) ound p egnan emales in ea ly Feb u-
a y and la e Ma ch ( he la e wi h wo nea - e m e-
uses). B osse and Cha les-Dominique (1990) no ed
ha his species is a e; i is known om Su iname and
F ench Guiana om only a single specimen in each
coun y.
I
was i s epo ed o A gen ina in
1966
(Fo nes and Massoia, 1966).
Specimens Examined (II ).-CHACO:
Resis encia, 5 (I CML, 3 MACN, l TCWC).
CORRIENTES: Laguna Pai a, BO Las Lomas, 2 (1
CML, I OMNH). MISIONES: Colonia Ma i es, I
(MACN); Posadas, 3 (MACN).
Addi ional Reco ds.- FORMOSA: Pa que
Nacional Rio Pilcomayo (S. Heinonen F. and R. M.
Ba quez, in li .). MISIONES: A oyo Zai nan (Massoia,
1980); Depa amen o Capi al (Massoia, 1980).
Genus Vampy essa
Vampy essa pusilla (Wagne )
Phyllos oma pusillum Wagne , Abhandlungen de
Ma hem.-physikaliscchen Classe de Ko ni glich
Baye ischen Akademie de Wissenscha en, Muenchen,
5: 173, 1843.
Vampy essapusilla: Mille , Bulle in o he Uni ed
S a es Na ional Museum, 57: 156, 1907.
TypeLocali y.- B azil, Rio de Janei o, Sapi iba.
Dis ibu ion.- The genus con ains i e species
in Sou h Ame ica (Koopman, 1993), only one o which,
V pusilla, is ound in A gen ina. This species occu s
(modi ied om Koopman, 1993) om sou he n Mexico
o Boli ia and no heas e n A gen ina.
Dis ibu ion in A gen ina (Fig. 44).-
In
A gen-
ina, his species is known only om a single specimen
om he p o ince o Misiones.
Desc ip ion (Figs. 45 and 46).- Body size is me-
dium. Do sal colo a ion is pale b own, and he do sal
hai s a e icolo ed, wi h a medium band ha is pale
han he bases and o he ips. The en al hai s a e a
unicolo ed ligh b own. The mos dis inc i e ea u es
o his species a e he whi e acial s ipes, and he b igh
yellow colo on he agus and on he bo de s o he
ea s. The e is no mid-do sal s ipe. The u opa agium is
na ow, and he calca s a e e y sho .
The den al o mula is I 2/2, C 1/1, P 2/2, M 2/2,
o al 28. The uppe inciso s a e well sepa a ed om one
ano he . II is mo e han wice as la ge as 12. The cu -
ing edges o I I a e no bi id in he specimen examined.
The canine is o no mal shape, wi h he cingulum well
de eloped pos e io ly and almos in con ac wi h he
an e io su ace o Pl. The PI has a single cusp and is
abou one-hal he size o he P2. P2 is sepa a ed om
bo h P I and M2 by dias emas; P2 has wo cusps-a
well-de eloped and high pa acone, and a lowe
mesos yle. The labial cusps o he uppe mola s a e el-
e a ed o ming a cu ing edge; he lingual cusps a e less
de eloped, bu ex end labially o o m a pla o m. Lowe
BARQUEZ ET AL.-THE BATS OF ARGENTINA
a
Figu e 50.
A ibeus li u a us:
(a) ace; (b) agus; and (c)
u opa agium.
cingulum o he canine. The uppe inciso s comple ely
ill he space be ween he canines. The canines a e gen-
e ally no dis inc i e, al hough he e is a small canal on
he in e nal ace be ween wo c es s. The P 1 is abou
one-hal as high as he canine; he pos eno bo de o
PI is sha p and he oo h is abou one-hal he size o
P2; he mesos yle is small and ba ely isible as a low
cusp o ien ed owa d he labial side o he oo h ow and
in con ac wi h he an e io ma gin o P2. The P2 is
much like PI, bu has an in e nal pla o m ( alon) sup-
po ing a small p o ocone on i s an e olingual ma gin.
Viewed om abo e, he M I is iangula and posi ioned
wi h i s base owa d he labial side and i s ip owa d
he lingual side; i is wide han long, wi h he alon
expanded o e he bony pala e; he pa acone and me a-
69
cone a e he main cusps and he o me
IS
highe ; he
p o ocone and hypocone a e well de eloped, he anne
posi ioned in he an e olinguaJ ma gin o he alon, and
he la e also loca ed on his ma gin, bu lowe . Se -
e al specimens ha e a se ies o w inkles and olds on
he pla o m be ween he ex e nal and
in e nal
cusps.
The M2 is smalle han Ml and, al hough i has he
same cusps, hey a e less de eloped and i s shape is
o oid.
Bo h i1 and i2 a e small, subequal in size, and ill
he space be ween he canines comple ely; hey may be
bilobed, bu his is a iable. Canines a e well de el-
oped, bu he cingulum is ha dly isible. The p l is small,
iangula in la e al iew, and has only one well-de el-
oped cusp (p o oconid). The p2 is much highe han he
o he mola i o m ee h, and i s an e olingual
ma gin
has
a small pa aconid and a well-de eloped p o oconid; a
small hypoconid is p esen owa d he ex e nal ma gin
and almos in con ac wi h he an e io ex eme o m I.
The m I is longe han wide, wi h i s an e io ma gin
ounded and i s pos e io almos s aigh ; he p o oconid
and hypoconid a e clea ly isible, he i s la ge han
he second; a well-de eloped an e io me aconid and a
low en oconid a e aligned on he lingual ma gin. The
m2 is smalle , almos squa e; he en oconid is displaced
lingually o ming a ci cula bo de ha con inues pos-
e io ly; he hypoconid is almos absen ; me aconid and
en oconid a e well de eloped; an appa en pa aconid is
loca ed be ween he p o oconid and me aconid. The m3
is small and ounded, wi h poo ly di e en ia ed cusps.
Measu emen s.-
Table 6.
Commen ss-s A ibeus li u a us
inhabi s he eas -
e n o es s, aking shel e p incipally in he oliage o
all ees, bu also in ca es and ock issu es. In F ench
Guiana, B asse and Cha les-Dominique (1990) also
ound his species o be common in o es ed habi a s.
I s die consis s mainly o ui , bu also includes o he
plan pa s, such as lea es and lowe s; some au ho s
ha e epo ed insec s (Ga dne ,
1977a).
A leas 67 di -
e en species o plan s ha e been iden i ied in he s om-
achs o his species h oughou i s dis ibu ion (Ga dne ,
I
977a).
The s omachs o a ious specimens con ained
uniden i ied ui and seeds (R. M. Ba quez, pe sonal
obse a ion). In Iguazu, Misiones P o ince, his spe-
cies eeds no only on
wild
ui s, bu also on cul i a ed
ui s, such as bananas and mangoes (C espo, 1982).
B osse and Cha les-Dominique (1990) epo ed his
species eeding in locks on la ge ui ees.

70
SPECIAL PUBLICATIONS, MUSEVM OF TEXAS TECH UNIVERSITY
•
"~
-
I .;. ,
. 1
[,
'
• 1
---
Figu e 51. Skull and ee h o A ibeus li e a us Adul male om Pa aje Pa a iso, Misiones
P o ince (CML 2172). Ba
=
I
em. Tee h no o scale.
Wilson (1979) summa ized wha is known abou
ep oduc ion. He no ed ha i can ep oduce all yea ,
al hough in he no he n pa o i s dis ibu ion only one
young is p oduced yea ly, whe eas owa ds he sou h
ep oduc i e ac i i y is less p o ac ed. C espo (1982)
obse ed ep oduc i e ac i i y in Pa que Nacional
Iguazu be ween Sep embe and Feb ua y, and p egnan
emales we e ound be ween No embe and Feb ua y;
he also no ed ha males we e cap u ed mo e equen ly
han emales.
Specimens Examined (45).- CORRIENTES:
Es ancia Pues o Valle, 25 k n E I uzaingo, I (AMNH).
FORMOSA: Bou ie , I (ROM). MISIONES: Ca a a as
del Iguazu, I (MPB); Colonia Ma i es, 3 (MACN); Dos
de Mayo, 2 (l CEM, I CML); Es ancia San Jo ge, 3
(MACN); Gobe nado Lanusse, 2 (CML); Iguazu, 18
BARQUEZ ET AL.-THE BATS OF ARGENTINA 71
Table 6. Measu emen s o A ibeus imb ia us, A ibeus li u a us, and A ibeus plani os is. Fo a
desc ip ion o he measu emen s, see he Ma e ials and Me hods. Mean, s anda d de ia ion, and sample size a e
ollowed by ange.
Va iable
A ibeus imb ia us A ibeus li u a us A ibeus plani os is
To al leng h 86.0 ± 5.47, 5 94 1
±
17.03, 6 87.8 ± 8.63, 40
80.0 - 90.0 75.0 - 120.0 750 - 110.0
Hind oo leng h 16.0, I 16.5 ± 1.93,6 16.6 ± 3.36, 40
14.0 - 18.8 12.0-257
Ea leng h 2J.5 ± 2.82,5 223± 1.61,6 21.9 ± 3.26, 40
18.5 - 25.0 20.0 - 24.3 15.0 - 26.3
Fo ea m leng h 66.1 ± 1.14, 6 71.4 ± I. 66, 13 65.7 ± 1.89, 43
64.0 - 67.2 68.0 - 74.0 62.0 - 69.0
Weigh 56.5, 1 76.6 ± 10.69, 3 52.3 ± 6.67,36
70.0 - 89.0 390 - 690
Condy1obasa11eng h 28.4 ± 0.34, 5 281 i 0.58,13 27.2 i 0.66,33
279 - 28.7 270 - 28.9 26.0 - 288
Leas in e o bi al b ead h 8.3 ± 0.48, 5 7.7 ± 0.34,8 85 ± 0.44, 12
7.8 - 89 71 - 81 78 - 9.1
Zygoma ic b ead h 18.9 ± 0.38,6 191 ±0.36, 13 19.1 ± 0.64, 34
185-196 183-197 17.7 - 20.4
G ea es leng h o skull 31.2 ± 0.51,5 31.6±0.61,13 30.2 ± 068, 36
307 - 31.8 30.3 - 33.0 29.0 - 320
Pos o bi al cons ic ion 7.3 ± 0.24,6 66 ± 0.25,13 7.4 ± 0.26, 36
7.1 - 7.7 6.4 - 7.2 6.9 - 7.9
B ead h o b aincase 13.3 ± 0.43, 5 13.7 ± 0.49,13 135 ± 0.38,35
130 - 14.1 13.2 - 15.0 12.4 - 14.2
Leng h o maxilla y oo h ow 11.3 ± 0.28, 6 11.0 ± 0.26,12 11.0 ± 031, 34
10.9 - 11.7 106 - 115 10.5-117
Pala al leng h 15.4 ± 0.20, 6 14.9 ± 057,13 14.4
±
0.44, 36
152 - 158 14.1 - 15.7 13.7 - 15.2
Mas oidal b ead h 16.8 ± 0.46, 5 16.6 ± 0.43,13 163
±
0.50, 34
16.2 - 17.5 160 - 17.5 155-17.4
Leng h o mandibula oo h ow 108
±
0.25,5 105iO.33,12 10.8 ± 1.04,35
105-11.1 97 - 109 9.6 - 12.9
Leng h o mandible 2J.9±0.41,6 2J.9
±
0.45, ]3 21.2 ± 0.53, 36
21.4 - 225 21.0 - 22.5 20.2 - 22.5
C-C (wid h ac oss canines) 8.9 ± 0.28, 4 89±022,12 8.9 ± 0.32, 35
86 - 93 85 - 9.3 82 - 9.6
M-M (wid h ac oss mola s) 140
±
0.33,5 133 ± 0.36, 12 12.9 ± 0.35,33
13.6 - 14.4 126 - 13.9 12.2 - 136
(MACN); Je . Hwy 2 and A oyo Pa aiso, 3 (I CML, 1
IADIZA-CM, 1 OMNH);Je .Hwy21 andA oyoO eja
Neg a, app ox. 2 km W Pa que P o incial Mocona, 6
(4 CML, 2 IADIZA-CM); Pa aje Pa aiso, 16 km SE, 1
(CML); Pa que Nacional Iguazu, 2 (MACN); San Ped o,
47 km SE, 2 (CML).
(Mas so
ia
e al., 1987); Depa amen o Cainguas
(Massoia, 1980); Depa amen o Candela ia (Massoia,
1980); Depa amen o Capi al (Massoia, 1980);
Depa amen o Gua ani (Massoia, 1980).
Addi ional Reco ds.- CORRIENTES: I a Iba e
(Lo d e aI., 1973). MISIONES: A oyo U ugua-i
72 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
60°
443 352
,
~.
707-+-. 536,537,538
495~,~4. 359,360
In'
7
776~.711
.
31.~', 598
" ..,I"~
1
04 ••
:-
402 403
i
280'~·,
·,:279 ' :-"'"
366
-7
".'58 ,/"
106,493 .602 / ""',-..."""
'------------' 601'- 519 / -,
~~/
.:..
-----~~-----------
' ,,d
715
i
578~~.!16
i
411,412
.l.
~30, 53,584,60~
541
424
i
,
,
,
~
.•
_,--;~
~
----------
-
---
-
-----~
'-~..- _...-"'_,.:-.. !
Figu e 52. Map o he locali ies o A ibeus plani os is.
A ibeus plani os is
(Spix)
Phyllos oma plani os e Spix, Simia um e
Vespe ilionum B asiliensium species no ae ou his oi e
na u elle des especies nou elles de singes e de chau e-
sou is obse ees e ecueillies pendan Ie oyage dans
l'in e ieu du B esil execu e pa o d e de S. M. Ie Roi
de Ba ie e dans les annees 1817, 1818, 1819, 1820.
F ancisci Se aphici Hubsch nanni, Monachii, p. 66,
1823.
A ibeus plani os is: Dobson, Ca alogue o he
Chi op e a in he Collec ion o he B i ish Museum, B i -
ish Museum (Na u al His o y), London, p. 515, 1878.
Type Locali y.- B azil, Bahia, Sal ado .
Ol og (1979) epo ed A. jamaicensis he cules in
Ce o Calilegua, Jujuy P o ince. This subspecies was
g ouped wi h A. plani os is and associa ed wi h A.
allax (Koopman, 1978). The axonomic si ua ion o he
A. li u a us-jamaicensis-plani os is species complex
is complica ed in no he n A gen ina (Handley, 1989,
1991). Un il u he s udy, we ecognize specimens om
he No hwes as A. plani os is. The ollowing sub-
species occu s in A gen ina.
o
100 200
I I I
400km
I
A ibeus plani os is allax Pe e s
A ibeus allax Pe e s, Mona sbe ich e de
Koniglich P eussischen Akademie de Wissenscha en
zu Be lin, p. 355,1865.
A ibeus plani os is allax: Ande sen, P oceed-
ings o he Zoological Socie y o London, 1908 :242,
1908.
Type Locali y.- Su inam, Diepe ink.
Dis ibu ion.- Sou he n Venezuela and he
Guianas o no he n A gen ina (Koopman, 1993).
Dis ibu ion in A gen ina.- Figu e 52.
Desc ip ion (Figs.
53
and
54).-
Size is la ge, bu
sligh ly smalle han A. li u a us. Leng h o he o ea m
anges om 62 o 69 mm. The pelage is so and dense.
Do sal colo is g ayish b own; he basal band is wide
and pale, and he ips a e da k; some hai s ha e ashy o
whi e os ed ips. The en e is ligh b own and mos
hai s a e unicolo ed, bu os ed ips mo e common han
on he do sum. The acial s ipes a e only sligh ly e i-
den o absen in many specimens. Hai is sca ce o e
he u opa agium, legs, and oes. Wing ips ha e a whi e
spo . O he cha ac e is ics a e simila o A. li u a us .
BARQUEZ ET AL.- THE BATS OF ARGENTINA
a
Figu e 53. A ibeus
plani os is:
(a) ace; (b) agus; and
(c) u opa agium,
The skull is simila o A. li u a us bu less obus ,
he pos o bi al cons ic ion is wide, and he pos o bi al
p ocesses a e no as dis inc .
The den al o mula is 2/2, 1/1, 2/2, 2-3/3, o al
30-32. Den i ion is simila o ha o A. li u a us; M3 is
minu e, bu is usually p esen .
Measu emen s.- Table 6.
Commen s.- Li le is known abou he biology
o his species in A gen ina. Ou da a show ha i is
common, bu no abundan , and inhabi s he o es s o
no hwes e n A gen ina as a sou h as Tucuman P o -
ince, whe e i is a e. Rep oduc i e da a a e sca ce.
Males ha e been cap u ed wi h sc o al es es in he
middle o Oc obe inAguas Chiqui as,
Tucu nan
P o -
ince. Specimens collec ed in he middle o May in
Piqui enda Viejo, Sal a P o ince, and a he end o June
in Laguna La B ea, Jujuy P o ince, we e no ep oduc-
73
i ely ac i e. Indi iduals we e ep oduc i ely ac i e a
he middle o No embe in Se ania de las Pa as, Sal a
P o ince.
Specimens Examined (135).- JUJUY: Ab a de
Canas, EI Monoli o, 1700 m, 1 (CML); Aguas Neg as,
Pa que Nacional Calilegua, I (CML); Calilegua, I
(FCM); Camino de Co nisa, 10 km W Dique La
Cienaga, I (CML); Laguna La B ea , 3 (I MD, 2
OMNH); Laguna La B ea, 25 km W Palma Sola, 9
(CML); On Highway 9 a bo de wi h Sal a, a camp-
g ound on he way o EI Ca men, 3 (l IADIZA-CM, 2
OMNH); Rio de Zo a y c uce can u a 34, I (CML);
Yu o, 4 (3 AMNH, I CML). SALTA: Ab a G ande, I
(OMNH); Aguas Blancas, 2 (CML); A oyo Los
Noques, Pa que Nacional EI Rey, I (CM); I au (Campo
La go), Sie as de Ta agal, 1600 n, 1 (CML); Jun a de
los Rios Ta ija y Be mejo, I (CML); Jun as de San An-
onio, 13 (CM); La Calde a, I (ROM); Los Mad ejones,
Jun a de los Rios Ta ija e I au, 1400 n, I (CML); O an,
15 km S, 15 km W, along Rio San a Ma ia, 4 (CM);
Paso de la C uz, 2 (CML); Piqui enda Viejo, 30 km N
Ta agal, 12 (CM); Piqui enda Viejo, 6 km W, I
(OMNH); Piqui enda Viejo, 8 kID W, 2 (CML); Rio EI
Na anjo, 14 km W de Ru a p o incial 5, I (CML); Rio
Guanaco, 32 km al NE de Lumb e a, 1 (CML); Se ania
de las Pa as, 3 (I CML, 2 OMNH); Tabacal, 41 (40
MACN, 1 TTU). TUCUMAN: Agua Colo ada, 8
(MACN); Aguas Chiqui as, Sie as de Medina, 800 m,
I (CM); A oyo Aguas Chiqui as, Rese a P o incial
Aguas Chiqui as, 1 (CML); Las Jun as, 22 km W
Cho omo o, I (CML); Las Jun as, 22 km W Cho omo o
on Hwy 312, 3,500
.,
I (OMNH); Las Tipas, Pa que
Biol6gico, I (CML); Playa La ga, Rio Los Sosa, Ru a
307 km 19.7, I (CML); Queb ada del To o, I (CML);
Rese a P o incial "Aguas Chiqui as" EI Cadillal, 4
(CM); Rio Lo o, 3 (RMB); Taco Yana, I (CML).
Addi ional Reco ds.- FORMOSA: Pa que
Nacional Rio Pilcomayo (S. Heinonen
F
and R. M.
Ba quez, in li .). JUJUY: Finca EI Rema e, 24 km SE
San Sal ado de Jujuy, 740 m (Villa-R. and Villa-C.,
1971, as A. li u a us); Finca La Ca olina, Los Pe ales,
San Sal ado de Jujuy, 1310 m (Villa-R. and Villa-C.,
1971, as A. li u a us).
74 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
/
Figu e 54. Skull and ee h o A ibeus plani os is. Adul male om Se ania de las Pa as,
Sal a P o ince (OMNH 18944). Ba
=
1 em. Tee h no o scale.

BARQUEZ ET AL.-THE BATS OF ARGENTINA 75
24°
J
Figu e 55. Map o he locali ies o Pygode ma bilabia um.
Genus Pygode ma
Pygode ma bilabia um (Wagne )
Phyllos oma bilabia um Wagne , Wiegmann's
A chi u Na u geschich e, Jah gang 9, Bd. 1:366,
1843.
P [ygode maJ bilabia um: Pe e s, Mona sbe ich e
de Kbniglich P eussischen Akademie de
Wissenscha en zu Be lin, p. 357, 1866.
TypeLocali y.-
B azil, Sao Paulo, Ipanema.
Dis/ ibu/ion.- This species was conside ed mo-
no ypic, un il Owen and Webs e (1983) ecognized wo
subspecies, P b. bilabia um and P b. magna, based p in-
cipally on size di e ences. P b. magna was desc ibed
o no hwes e n A gen ina and Boli ia, and P. b.
bilabia um was assumed o ex end om no heas e n
A gen ina no hwa d h ough B azil and Pa aguay o
Su iname (see also Webs e and Owen, 1984).
Dis ibu ion in A gen/ina.- Figu e 55.
Desc ip ion (Figs.
56
and
57).-
Size is medium;
leng h o o ea m anges om 38.0 o 44.2 nun. The
uppe lip has a longi udinal old ha gi es i a double-
lipped appea ance. A whi e pa ch on each shoulde is
o
I
200 400 km
I I
cha ac e is ic o his species. The bo de s o he ea s
and he agus a e yellow. Gene al colo is g ayish
b own, ligh e and mo e uni o m on he en e . Hai s
o he do sum a e icolo ed wi h a da k b own basal
band, a pale , almos whi e, middle band, and b own
ips sligh ly ligh e han he basal band. Hai s o he
en e a e unicolo ed. The pelage is long and so , uni-
onnly dis ibu ed o e he body, and ex ends o e mo e
han one-hal o he o ea m along he do sal and en-
al sides, and o e he en i e do sal su ace o he
u opa agiu n and ibia (in lesse amoun s on he en al
side). Memb anes a e da k and he small
dac ylopa agium is anspa en .
The nose lea is well de eloped, la ge, and wide.
The u opa agium is semici cula and suppo ed by small
calca s. The medium-sized ea s ha e ounded ips and
wide bases. Nume ous glands occu on he os um nea
he nose lea and a ound he eyes, (pa icula ly in males),
gi ing a swollen appea ance. Nume ous papillae a e
ound on he lowe lip. A ail is absen .
The skull is dis inc i e, wi h a sho os um (mo e
in la ed in emales and in specimens om Misiones
P o ince compa ed wi h hose om Sal a P o ince).
O bi s a e small; zygoma ic a ches a e comple e bu hin.
The pala e is sho and he oo h ows a e semici cula .
The p e ygoids a e sho and di e ge pos e io ly; he
76
SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
a
c
Figu e 56.
Pygode ma bilabi a um;
(a) ace; (b)
u opa agium; and (c) agus.
hamula p ocess is sho and blun . The basisphenoid
pi s a e elonga e and deep.
The b aincase is ounded. A low sagi al c es is
p esen and con inuous wi h he lambdoidal c es s,
which a e well ma ked in emales and less so in males.
The den al o mula is 2/2, II I, 2/2, 2/2, o al 28.
The II is well de eloped, iangula and poin ed, and
he inciso s a e in con ac only a hei bases; hei ex-
e nal bo de is sha p and se a ed. The 12 is small, no
eaching he cingulum o he canines, inclined inwa d,
and wi h he dis al bo de ounded and illing he space
be ween he I I and he canines. Canines a e well de el-
oped and wide wi h a cu ing la e al bo de and a e y
sha p ip; he cingulum is expanded inwa d owa d he
pos e olingual side whe e i o ms a small cusp. The PI
and P2 a e well de eloped. The P I is he wides oo h
in he oo h ow in la e al iew, is iangula in la e al
iew and he cusps, excep o he ou e one, a e li le
de eloped. The M I is wide, wi h he pa acone well de-
eloped and in he o m o a cu ing blade, and lingually
wi h a low p o ocone and a less de eloped hypocone.
The M2 is educed and wi hou well-di e en ia ed
cusps.
The mandibula oo h ow is semici cula . Inciso s
a e subequal in size, small, bilobed, and deeply g oo ed
along he an e io ace. Canines a e no mal. The in e -
nal ace o he p emola s has a median sep um ha di-
ides i an e io ly and pos e io ly. Mola s wi h well-
de eloped inne cusps, he second sligh ly less han one-
hal he size o he i s . Occasionally a hi d lowe o
uppe mola may be p esen on one o bo h sides (Owen
and Webs e , 1983).
Measu emen s.-
Table 5.
Commen s.-
Ol og (1967) i s epo ed his ba
o A gen ina. This species p e e s humid o es s. Fo nes
and Delpie o (1969) cap u ed specimens in human
dwellings nea " ala" ees
(Cel is).
Specimens in Bo-
li ia we e cap u ed in low woodlands wi h palms and
in he denses pa o he o es in swampy a eas (R. M.
Ba quez, pe sonal obse a ion). This species is p eyed
upon by S ygian owls
Asio s ygius)
in he ce ado sa-
anna o B azil (Mo a Junio and Taddei, 1992). I s
ood seems o be mainly ui , bu ew da a a e a ail-
able. S omachs o cap u ed specimens we e emp y o
solids, bu liquids appea o pe ain o ui s. Yellowish
seeds we e ound in he eces o one specimen om
Piqui enda Viejo, Sal a P o ince. C espo (1982) no ed
ha specimens o his species app oached cul i a ed ui
ees.
Li le is known abou hei ep oduc ion; C espo
(1982) no ed no ac i i y in specimens cap u ed in
Misiones P o ince in Sep embe and No embe . Se -
e al emales we cap u ed in Misiones (San Ped o, 47
km E) on 3 I Oc obe had nea - e m e uses, one ga e
bi h in he ne . A Gobe nado Lanusse, Misiones P o -
ince, II lac a ing emales and a ju enile we e cap u ed
in No embe . In Piqui enda Viejo, Sal a P o ince, a lac-
a ing emale was collec ed in mid-May. Specimens
om Boli ia we e no ep oduc i ely ac i e in July (R.
M. Ba quez, pe sonal obse a ion). Mye s (1981) cap-
u ed p egnan emales in Ma ch, July, and Augus in
Pa aguay. This species may ep oduce aIIyea .
Specimens Examined
(37).- CORRlENTES: San
Ca los, I (MACN). MISIONES: 6 km NE by Highway
2 o jc . Highway 2 and A oyo Pa aiso, I (lADIZA-
BARQUEZ ET AL.-THE BATS OF ARGENTINA
•."0"
,
,
77
~
.
~
~~
"-
~,
,
Figu e 57. Skull and ee h o
Pygode ma bilabia um.
Adul emale om 47 km SE San Ped o,
Misiones P o ince (CML 2162). Ba
=
1 em. Tee h no o scale.
CM); AJma ue e, J (MACN); A is obu!o del Valle, 10
km W by oad along Rio Cu iapi u, 1 (CML); Colonia
Ma i es, 5 (MACN); Es ancia San Jo ge, 1 (MACN);
Gobemado Lanusse, 13 (CML); Misiones (no speci ic
locali y), 1 (BMNH); P o undidad, I (MACN); San
Ped o, 20 km al SE, 1 (RAO); San Ped o, 47 km SE, 7
(5 CML, 2 IADIZA-CM). SALTA: Agua Blanca, 2
(CML); Piqui enda Viejo, 30 km N Ta agal, 1 (CM);
Vado de A azayal, 20 km NW Aguas Blancas, 1 (CML).
Addi ional Reco ds.-
MISIONES: A oyo
U ugua-i (Massoia e al., 1987); Ca a a as del Iguazu
(Massoi a, 1980); Depa amen o Capi al (Massoia,
1980); Depa amen o Gua ani (Massoia, 1980).
78 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
SUBFAMILY DESMODONTINAE
Vampi es ha e been conside ed bo h a dis inc
amily and a ibe (Desmodina) o he Family
Phyllos omidae (G ay, 1866). Recen wo k (Fo man e
al., 1968; Jones and Ca e , 1976), howe e , ecognizes
ampi es as a sub amily. Th ee species a e included,
which a e qui e dis inc om o he phyllos omids. All
a e cha ac e ized by a educed, almos indis inguish-
able, nose lea , and he e y specialized ee h needed
o blood eeding, wi h he uppe inciso s and canines
being sha pened blades.
KEY TO THE SPECIES
1. Thumb sho , wi h wo dis inc pads; wing ips whi e; co onoid p ocess mo e
ele a ed han he mandibula condyle gi ing a iangula appea ance Diaemus youngi
I'. Thumb la ge , wi h h ee la ge pads; wing ips no whi e; co onoid p ocess
mo e o less equal o he mandibula condyle gi ing a squa e appea ance Desmodus o undus
Genus Desmodus
Desmodus
o undus
CE.
Geo oy S .- Hilai e)
Phyllos oma o undum
E.
Geo oy S .- Hilai e,
Annales du Museum d'His oi e Na u elle, Pa is, 15:181,
1810.
Desmodus o undus: Thomas, Annals and Maga-
zine o Na u al His o y, se . 7, 8:194,1901.
Type Locali y.- Pa aguay. Res ic ed o Asuncion
by Cab e a (1958).
Dis ibu ion.- Widely dis ibu ed om no he n
Mexico sou hwa d h ough cen al and pe haps sou h-
e n A gen ina, wi h ew eco ds om he Amazon ba-
sin. Two subspecies (D. . mu inus and D. . o undus)
a e ecognized, bu hei di e ences a e no clea and
hey may no be alid axa. Only D.
.
o undus is ound
in A gen ina.
Dis ibu ion in A gen ina (Fig. 58).- This spe-
cies is common h oughou mos o i s ange in A gen-
ina.
Desc ip ion (Figs. 59 and 60).- Size medium,
wi h he leng h o he o ea m anging om 60 o 67
mm, and he body s ong and muscula . The pelage is
sho and s i . Colo a ies, bu is gene ally b own o
g ay; some specimens a e eddish and o he s a e golden.
The en e is pale han he do sum. Basal bands o he
do sal hai s a e whi ish, while hose o he en e a e
da k. Do sally, he pelage ex ends o e he en i e o e-
ann, p opa agium, u opa agium, plagiopa agium, and
legs. Dis ibu ion o hai is simila do sally and en-
ally, bu he o ea m is no densely u ed and is na-
ked in many specimens. The nose lea is educed and
almos impe cep ible. The snou has a se ies o glands
and a cen al p o ube ance wi h a V-shaped inden ion;
he nasal openings a e ellip ical and elonga ed. The
lowe lip has se e al callosi ies ha o m a "V" wi h a
cen al g oo e. The ea s a e sho , sepa a ed, ha e a
con ex ex e io ma gin, and ounded ips. The inne ace
o he pinna has p onounced idges owa d i s ex e nal
ma gin; he agus has a con ex in e nal bo de , a poin ed
ip, and a cons ic ed base. Tail and calca a e absen .
The humb is long and has h ee pads, di e ing om
Di
aemus youngi , which has only wo. The
plagiopa agium a aches a he base o he oo ; he
u opa agium is educed o a na ow memb ane ha bo -
de s he legs and he pos e io end o he body.
The b aincase is la ge and ounded; he os um
is sho ; zygoma ic a ches a e comple e and widened
medially. Sagi al and lambdoidal c es s a e poo ly de-
eloped o absen . The pala e is sho and conca e, wi h
la ge incisi e o amina, and i s pos e io ma gin ends
a he midpoin o he zygoma ic a ches; he caudal spine
o he pala e is ba ely de eloped. Tympanic bullae a e
well de eloped (bu smalle han in D. youngi) and co e
abou one-hal o he cochlea.
The den al o mula is
I
112,
C I
II,
P
112,
M
II
I,
o al 20. The
II
and C I a e bladelike, acu ely poin ed,
and wi h sha p pos e io ma gins. The M1 is educed
and smalle han PI; bo h a e la e ally comp essed and
bladelike. A wide space sepa a es iI and i2 om he
canines. The i1 and i2 a e bilobed (in
D.
youngi only
he ou e lowe inciso s a e bilobed). The pi, p2, and
m1 a e educed, as in he uppe oo h ow. The co onoid
BARQUEZ ET AL.-THE BATS OF ARGENTINA
.•...•.
85
Figu e 63. Skull and ee h o
Diaemus
youngi .
Adul emale om Agua Salada, Jujuy P o ince
(CML 1343) Ba
= ]
em. Tee h no o scale.
li ia, o om a eas wes o he Andes in Pe u, Ecuado ,
and sou he n Colombia.
Dis ibu ion in A gen ina.-
Figu e 61.
Desc ip ion
(Figs. 62 and 63 ).-
Diaemus youngi
is gene ally simila o
D.
o undus,
bu is sligh ly smalle ,
wi h mo e dis inc i e whi e wing ips, and a sho e
humb ha ing only wo callosi ies. The specimen om
Jujuy P o ince is golden blond bo h do sally and en-
ally; wing memb anes a e da k, almos black, making
he whi e wing ips e en mo e dis inc i e; he
u opa agium is pale and co e ed do sally and en ally
wi h blond hai s, wi h a inge a he ma gin. The nose
lea is educed, as in
D. o undus,
bu he
"y"
is mo e

86
SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
open. Nasal openings a e ounded, ins ead o elonga ed
as in Desmodus. The pos e io leshy p ocess o he nose
lea is almos absen and he glands bo de ing he nasal
p ocess a e well de eloped, as is he u ow be ween
hese and he ex e io ma gin o he p ocess. Ea s a e
simila o Desmodus; he agus is less poin ed. This
species is cha ac e ized by la ge glands inside he mou h
ha a e clea ly isible in li e animals when hey open
he mou h.
The skull is simila o Desmodus, bu wi h la ge
ympanic bullae, a mo e ele a ed os um, and a di e -
en mandibula con igu a ion in which he ma gin be-
ween he co onoid p ocess and he mandibula condyle
is ele a ed, gi ing i a iangula aspec .
The den al o mula is I 1/2, C 111,P 1/2, M
J
-2/1,
o al 20-22. The i I and i2 gene ally a e no bilobed,
al hough i2 is bilobed in some indi iduals.
In
he speci-
men om Jujuy P o ince, inciso s a e no bilobed and
he o al numbe o ee h is 20.
In
a specimen om
Misiones P o ince, he ou e inciso s a e bilobed and
he e a e 22 ee h, bu his appea s o be aju enile; pos-
sibly he small second mola s a e los wi h age.
Measu emen s.- Table 7.
Commen s.- Goodwin and G eenhall (1961)
ound his species li ing in g oups o mo e han 30 in-
di iduals in a hollow ee and one indi idual was iso-
la ed in a well-ligh ed ca e. Fe nandez (198 I) indica ed
ha he species is sca ce in Venezuela; a ew indi idu-
als we e obse ed eeding on domes ic chickens, mak-
ing incisions on hei ee , which may kill he bi d
h ough blood loss. The Jujuy specimen (Ba quez,
1984b) was apped in a mis ne abou 4 m abo e he
g ound. Abou 60 e n below he ba , a small owl (O us
choliba) was also apped in he ne and, upon exami-
na ion, was ound o ha e spo s o blood p obably p o-
duced by bi es om he ampi e. This ba is also ea en
by ba n owls (Ty o alba) in A gen ina (Massoia e aI.,
1989c).
In
spi e o in ensi e e o s o cap u e and collec
specimens, we ha e only been able o cap u e a single
animal; his species is a e in no he n A gen ina. The
en i onmen whe e i was cap u ed was a complex mix
o chacoan ho n sc ub in e mixed wi h ansi ional o -
es , and loca ed only a ew kilome e s om he Yungas
o es . The specimen, cap u ed a he beginning o Au-
gus , was a emale wi h an open agina.
Specimens Examined (3).- JUJUY: Agua Salada,
app. 15 k n SE Agua Calien e, lado o ien al del Rio San
F ancisco, 1(CML). MISIONES: Eldo ado, 1 (MACN);
Picada Vieja, 1 (CML).
Addi ional Reco ds.- MISIONES: Bonp land
(Massoia e al., 1989c); Dos de Mayo (Massoia, 1980);
Eldo ado, 3 km SE (Delpie o e aI., 1973).
FAMILY VESPERTILIONIDAE
This amily has a wo ldwide dis ibu ion, being
absen only om he A c ic and An a c ic.
I
is com-
posed o 6 sub amilies, 34 gene a, and abou 315 spe-
cies. Many gene a a e sha ed among a ious geog aphic
egions, bu some, such as Tomopeas in Pe u, a e qui e
es ic ed. Some species a ain he sou he nmos dis i-
bu ions known o ba s, eaching Tie a del Fuego, A -
gen ina, and Isla Na a ino, Chile. Only he Sub amily
Vespe ilioninae, wi h 5 gene a and 20 species, is ound
in A gen ina. The species in his sub amily can be iden-
i ied by a long ail ha is included o ally in he
u opa agiu n and eaches he pos e io bo de o he
memb ane, a wide e na gina ion a he an e io bo de
o he pala e ha sepa a es he uppe inciso s, and by
he lack o a nose lea (a cha ac e sha ed wi h he
Noc ilionidae and Molossidae). The agus is well de-
eloped.
In
A gen ina, he ea s, which a e o a ious
shapes and sizes, p o ide cha ac e s diagnos ic o he
di e en gene a. The ea s a e long, wide, and ounded
in His io us; medium sized, na ow, and gene ally
poin ed in Myo is and Ep esicus; and sho (almos la -
ened agains he head) and ounded in Lasiu us and
Dasyp e us.
All A gen ine species a e insec i o ous. Al hough
a ious oos s a e u ilized, he e is a endency o oc-
cupy u ban and subu ban a eas, dwellings, b idges, is-
su es in buildings and ocks, ee holes, and ca es. Some
species a e mig a o y, o he s hibe na e. The ecology and
beha io o espe ilionids in A gen ina is essen ially
unknown.
BARQUEZ ET AL.-THE BATS OF ARGENTINA
I.
I'.
2 (1).
2'.
3 (2').
3'.
4 (3).
4'.
5 (1').
5'.
6 (5).
6'.
7 (6').
7'.
8 (7).
8'.
9 (7').
9'.
10(5').
87
KEY TO THE SPECIES
Do sal su ace o u opa agium densely u ed; ea s sho and ounded 2
Do sal su ace o u opa agium nea ly naked o spa sely hai ed; ea s ela i e~
la ge and o di e en shapes 5
P oximal one-hal o do sal su ace o u opa agium co e ed wi h hai s; gene al
colo a ion yellowish g ay o yellowish b own, wi h oli e ones Dasyp e us ega
Do sal su ace o u opa agium almos comple ely co e ed wi h hai s 3
Gene al colo a ion wi h os ed appea ance, wi h mix o g ayish o eddish hai s 4
Gene al colo a ion s ongly eddish (b ick ed), bu no os ed Lasiu us a ius
Leng h o o ea m > 50 mm Lasiu us cine eus
Leng h o o ea m 37-42 mm Lasiu us blosse illii
Ea s la ge, gene ally> 25 mm, ex ending well beyond ip o nose when
laid o wa d 6
Ea s o medium size,
<
20 mm, gene ally no ex ending beyond ip o nose
when laid o wa d 10
Ea s iangula , uni ed by a band ac oss he o ehead; colo a ion da k b own;
ips o do sal hai s con as ing li le wi h bases; only known om
Misiones P o ince His io us ela us
Ea s o oid; connec ing band be ween ea s wide, na ow, o absen ; gene al
colo a ion pale han abo e, bu colo o ips o do sal hai s clea ly
con as ing wi h bases 7
Ra io o leng hs o head and body e sus ea anging om 2.3 o 3.1;
leng h o he bullae> 4 mm 8
Ra io o leng hs o head-body e sus ea anging om 1.4 o 2.1; leng h o he
bullae
<
4 mm His io us mac o us 9
Leng h o ea
<
22 mm, no connec ed by a band; gene al colo a ion da k
do sally and en ally His io us magellanicus
Leng h o ea > 22 mm, some imes connec ed by a low band ac oss he o ehead;
colo a ion a iable, bu en e g ayish o whi ish, pale
han do sum His io us mon anus
Tips o hai s o en e g ayish o whi ish; ea s da k, almos black; connec ing
band be ween ea s na ow o ba ely isible; os um wide; b ead h ac oss
he hi d uppe mola s> 7 mm His io us mac o us mac o us
Tips o hai s o en e yellowish; ea s ela i ely pale; connec ing band
be ween ea s clea ly isible; os um na ow; b ead h ac oss he hi d
uppe mola s
<
7 mm His io us mac o us laepho is
Do sal su ace o base o u opa agium hai ed; ip o agus poin ed; 12 abou
same size as Il; PI and P2 small; a "space" made up o wo g ea ly educed
p emola s is p esen be ween P3 and he canine 11
88 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
10'. Do sal su ace o base o u opa agium naked o nea ly so; ip o agus ounded;
12 la ge han he inne ; no "space" p esen be ween he p emola and he canine 20
II (10). Sagi al c es p esen 12
II'. Sagi al c es absen 15
12 (II). P2 displaced o he lingual side o he oo h ow 13
12'. P2 no displaced om oo h ow 14
13 (12). Fu sho and el e like; do sal colo a ion usually b igh o ange, eddish,
o yellowish; leng h o o ea m 36.5-41.0 mm; pos o bi al cons ic ion
4.0-4.1 mm Myo is simus
13'. Fu sho , bu longe han abo e, and woolly; do sal colo a ion cinnamon b own,
some hai s wi h yellowish ips; leng h o o ea m 34-36 mm; pos o bi al
cons ic ion 3.4-3.5 mm Myo is ipa ius
14 (12'). Gene al colo a ion b igh ed wi h some och aceous ones; leng h o o ea m
37-40 mm; g ea es leng h o skull mo e han 14.5 mm; es ic ed o no heas e n Misiones
P o ince Myo is ube
14'. Colo a ion da ke , no ed o och aceous; leng h o o ea m> 40 mm;
g ea es leng h o he skull
<
14.3 mm; known only om he p o inces
o Sal a and Tucu nan Myo is keaysi
15 (11 '). U opa agium wi h ligh ly pigmen ed bo de and a inge o hai s on ma gin,
a ying in deg ee o de elopmen 16
15'. U opa agium wi hou ligh ly pigmen ed bo de o inge o hai s on ma gin 17
16 (15). Ven e , especially he pe ianal egion, pale o almos whi e; do sum wi h mix
o hai s gi ing a golden o sil e y os ed appea ance; pos o bi al cons ic ion 2:.
4.0 mm; ee la ge and ounded; os um sho ela i e o b aincase; lowe
oo h ow 5.0-5.6 mm Myo is albeseens
16'. Colo a ion o en e pale bu no whi ish; do sum may be os ed bu no
as equen ly as abo e; pos o bi al cons ic ion
<
4.0 mm (wi h a e
excep ions); ee small o la ge bu no ounded; os um elonga ed;
lowe oo h ow 5.5-6.1 mm Myo is le is 19
17 (15'). Do sal hai s icolo ed wi h whi e bases 1.0 mm long; leng h o o ea m
37.7-41.2 mm; known only om he p o inces o Chubu and
Rio Neg o Myo is aelleni
17'. Do sal hai s uni- o bicolo ed 18
18 (17').Leng h o o ea m 37.0-41.0 m n: do sal hai s usually bicolo ed wi h ips
pale han bases; known only om he p o inces o Chubu , Neuquen,
and Rio Neg o Myo is chiloensis
18'. Leng h o o ea m 32.0-36.0 mm; do sal hai s usually unicolo ed o mode a ely
bicolo ed; widely dis ibu ed in no he n A gen ina
Myo is
nig icans
BARQUEZ ET
AL.-
THE BATS OF ARGENTINA
89
19 (16'). Tips o do sal hai s yellowish; u opa agium wi h a ligh bo de ; leng h o o ea m
35.0-38.0 mm; pos o bi al cons ic ion 3.3-3.8 mm; om no hwes e n
A gen ina sou h o sou hwes o Buenos Ai es P o ince Myo is le is dinellii
19'. Tips o do sal hai s pale, bu no as yellowish as abo e; u opa agium wi h ligh
bo de and a inge o hai s; leng h o o ea m 35.0-39.3 mm; pos o bi al
cons ic ion 3.6 o 4.0 mm; only known om he p o inces o Buenos Ai es
and En e Rios Myo is le is le is
20 (10'). Size small; leng h o o eann
<
35 mm; leng h o mandible
<
10.1 mm; some
indi iduals wi h os ed appea ance, pa icula ly hose om no heas e n
A gen ina Ep esicus diminu us
20'. Size la ge; leng h o o eann > 35 mm; leng h o mandible > 10.1 mm 21
21 (20'). Leng h o o eann > 41 mm (nea 45 mm); g ea es leng h o skull > 16.7 mm;
zygoma ic b ead h> 12 mm; leng h o mandible> 12.9 mm Ep esicus b asiliensis
21'. Leng h o o ea m
<
41 mm; g ea es leng h o skull
<
16.5 mm;
leng h o uppe oo h ow
<
6.8 mm; mandible
<
13.0 mm Ep esicus u inalis
Genus Myo is
This is he la ges and mos complex genus o ba s
in he Neo opics. Re isions a e hose o Mille and
Allen (1928) and La Val (1973). The la e p esen ed a
cla i ica ion o he species o
Myo is,
bu lacks a com-
ple e and de ailed analysis o he A gen ine species. We
basically ollow La Val (1973), wi h modi ica ions gi en
in each species accoun . The di icul y in iden i ying
A gen ine Myo is is due p ima ily o poo knowledge
abou popula ion and geog aphic a ia ion, and he dis-
ibu ion o each species. Some o he axonomic cha -
ac e s a e no diagnos ic, hus ecological da a o en mus
be used o cla i y he s a us o a species. Some dis inc-
i e cha ac e s a e: h ee lowe and uppe p emola s,
medium-sized ea s wi h a sha ply poin ed agus, and
he p esence o hai s on he do sal base o he
u opa agium (di e ing in his om Ep esieus). The
skulls o many species a e simila , so sligh di e ences
a e o en used as diagnos ic cha ac e s, al hough hese
a e equen ly inconsis en . Fo his eason only hose
c anial cha ac e s ha help in iden i ica ion a e men-
ioned in a species accoun and compa isons a e made
only wi h simila species.
The alues ob ained om specimens used in de-
eloping he key o species e lec he a e age alues
o specimens we examined.
Some species a e easily dis inguishable, e.g.,
Myo is le is, M. albeseens, and M. ube , al hough in
some a eas he i s wo species can be con used wi h
one ano he . Myo is nig icans exhibi s conside able
a ia ion and could be con used wi h o he species.
Myo is chiloensis could be con used wi h M. aelleni;
hei dis ibu ions a e sympa ic and he iden i y o ei-
he is p oblema ic (see discussion o M. chiloensisy.
Myo is ipa ius could be con used wi h M. nig icans i
he mola s a e no examined.
Myo is
simus is mo e eas-
ily iden i ied by i s sho el e like u and b illian o -
ange and yellow colo . Myo is keaysi is simila o M.
nig icans, al hough he o ea m o M. keaysi is longe .
Ou iden i ica ions a e based on an examina ion o A -
gen ine specimens, a compa ison wi h specimens om
o he a eas, and he desc ip ions gi en by La Val (1973).
The den al o mula is I 2/3, C 1/1, P 3/3, M 3/3,
o al 38.
Myo is aelleni Baud
Myo is aelleni Baud, Re ue Suisse de Zoo logie,
86:268, 1979.
Type Loeali y.-A gen ina, Chubu , EI Hoyo de
Epuyen.
Dis ibu ion (Fig. 64 ).-Known only om he
ype locali y and EI Bolson, Rio Neg o P o ince; bo h
locali ies a e in close p oximi y.
Dese ip ion.-We did no examine he holo ype
o o he specimens used in he o iginal desc ip ion; cha -
90 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
Figu e 64. Map o locali ies o Myo is aelleni.
ac e is ics a e om he li e a u e (Baud, 1979). See also
he discussion unde he dis ibu ion sec ion o
M.
chiloensis.
Size is medium; leng h o o ea m is 37.7-41.2
mm. The skull is as wide, bu sho e , han M. l. le is,
and longe han
M. I. dinellii
and
M.
chiloensis, wi h
ma ked on al cu a u e. The occipi al egion is well
de eloped and ele a ed abo e he sagi al c es (which
is poo ly de eloped in ju eniles, bu mo e p onounced
in olde animals). Gene ally, M. aelleni appea s sligh ly
mo e obus han ei he M. l. dinellii o M. chiloensis.
The dis al one- hi d o he ou e edge o he ea s is con-
ca e; when olded o wa d, he ea s each o sligh ly
exceed he ip o he nose. The agus is well de el-
oped, i s ou e edge sligh ly wa y. The pelage is da ke
abo e and pale below. The leng h o he hai s on he
do sum and he sides is a leas 6 mm, bu no mo e
han 8 mm; he hai is no silky. The do sal hai s a e
icolo ed wi h a whi e basal band abou 0.6 mm-I.O
mm in wid h, ollowed by a da k ash-b own band, and
a e minal band o sepia-b own o cinnamon-b own
(wi h a sligh eddish one nea he ump and
u opa agium). The u opa agium is no immed wi h
whi e. The e is li le sexual dimo phism.
Commen s.- In o ma ion on he biology o his
species is gi en in Baud (1979).
Specimens Examined.- None.
Addi ional Reco ds.- CHUBUT: Hoyo de
Epuyen (Baud, 1979). RioNEGRO: EI Bols6n (Baud,
1979).
Myo is albescens
(E.
Geo oy S .- Hilai e)
Vespe ilio albescens
E.
Geo oy S .- Hilai e,
Annales du Museum d'His oi e Na u elle, Pa is, 8:204,
1806.
Myo is albescens: Thomas, Annali di Museo
Ci ico di S o ia Na u ale di Geno a, se . 2, 20:546,
1900.
Type Locali y.- Acco ding o Cab e a (1958), he
ype locali y is "Es ancia San Solano" in ex eme sou h-
e n Pa aguay, adjacen o he Es e o de Ibe a. This spe-
cies has had a con used axonomic his o y and nei he
a holo ype no a ype locali y was speci ied (La Val,
1973). Adding o he con usion, Acos a y La a (1950)
placed he ype locali y in Co ien es P o ince, A gen-
ina. La Val (1973) designa ed an adul emale, skin and
skull, AMNH No. 205195, om Yagua 6n, Pa agua i,
Pa aguay, as he neo ype.
Dis ibu ion.- This species is dis ibu ed om
sou he n Mexico o no he n A gen ina; i is no ound
in Chile o wes e n Pe u (Koopman, 1993).
Dis ibu ion in A gen ina.- Figu e 65.
Desc ip ion (Figs. 66,67, and 77g).- Leng h o
he o ea m anges om 31.0 o 37.3 mm. Adul s a e
easily iden i ied, al hough specimens a e simila o
M.
l. le is whe e hey a e sympa ic in Buenos Ai es and
En e Rios p o inces. Measu emen s o specimens om
no hwes e n A gen ina and om he p o ince o
Misiones a e sligh ly la ge han hose o he neo ype,
bu he cha ac e s a e clea ly hose o he species. The
pelage is so and dense. Many do sal hai s ha e whi e
ips, gi ing his ba a sil e y o golden in ( his os ed
appea ance is ound in ew o he species, excep ing
some M. l. le is). The en e is ligh colo ed, almos
pu e whi e on he abdomen and pe ianum and on he
sides o he body.
In
gene al, specimens om Misiones
P o ince a e da ke han hose om no hwes e n A -
gen ina; he inge o hai s on he bo de o he
u opa agium is well ma ked, e en mo e so han in M. l.
dinellii. The ee a e obus and ounded, simila o hose
o molossids. The b aincase is globose; he os um is
sho ; he pos o bi al cons ic ion is wide (gene ally>

BARQUEZ ET AL.-THE BATS OF ARGENTINA 91
: 625
-.' ,."" 270
637,640
I .•
601 ":. '-"" •
,376 ,.: 548 ...•
730:731 739
:'----<---------:
"J,56,157
" ' ,-"L:
.699: '
74~2S -:L-30, 212 :
163-+' :' :
;..~ ~·.362 :
'''',_ /':;5
'J-",'
I '
, , ---------------'
_ '-'.-- .>: ,
476,: 384
198:_
------.'1'"'
Figu e
65,
Map o he locali ies o
Myo is a/beseens
4 nun); he on al cu a u e is accen ua ed; and he e
is no sagi al c es .
Measu emen s.-
Table 8.
Commen s.-
Li le is known abou he biology
o his species in spi e o i s wide dis ibu ion. Speci-
mens ha e been cap u ed in human dwellings. We cap-
u ed indi iduals: as hey le he oo o a u al dwell-
348
89.
o
100 200
I I I
400
km
I
ing in Tucu nan P o ince; o e a gully wi h calm wa e
in mois o es ; in a eas o open chacoan ho n sc ub;
and unde he ea es o a small building in he d y ho n
sc ub o San iago del Es e o P o ince, which hey sha ed
wi h
Myo is nig icans , M. ipa ius,
and Molossus
molossus, Acos a y La a (1950) epo ed ha he spe-
cies li es in human dwellings, ock c e ices, nea un-
ning wa e , and unde ee ba k. Mye s (1977) indica ed
92 SPECIAL PlTBLICATIONS, MUSElTM OF TEXAS TECH UNIVERSITY
..'~i
Figu e 66. Myo is albescens: (a) ace, la e al iew; (b)
ace, on al iew; and (c) agus.
ha in Pa aguay i oos s in buildings da ing om he
las cen u y.
Rep oduc ion is poo ly known. Mye s (1977) sug-
ges ed ha delayed e iliza ion may occu ; ma ing was
obse ed in May and p egnan emales appea ed a he
end o July. Sexual dimo phism has been epo ed
(Mye s and We zel, 1983).
Specimens Examined
(85).-
BUENOS AIRES:
Bosch, 15 km NW BaJca ce, I (MSU); La Pla a, 2
(MLP); Lobos, I (ROM). CHACO: Pozo del Ga o, 2 (I
CML, 1 IADIZA-CM); Rio de O o, Gene al Vedia, 2
(l CML, I FCM); Rio Teuco, 10 km W Ta agal, 2
(CML). CORRIENTES: Goya, 2 (BMNH). ENTRE
RIOS: Es acion Pa anaci o, I (FCM); Islas de Ibicuy,
Pa anaci o, 13 (TTU). FORMOSA: Clo inda, 1 (TTU);
Colonia km 503, 2 (I CML, I IADIZA-CM); Colonia
km 503, 16 km S, 2 (I CML, 1IADIZA-CM); Es anislao
del Campo, I (TTU). MISIONES: Misiones (no spe-
ci ic locali y), I (BMNH); San Ped o, 26 km NE, sob e
u a 16, 1 (CML); San Ped o, 47 km SE, 2 (1 CML, I
IADIZA-CM). SALTA: Aguas Blancas, 27 km W, I
(MACN); Hickman, Luna Mue a, I (CML); La
Me ced, 2 (TCWC); O an, 15 km S, 15 km W, along
Rio San a Ma ia, 1 (CM); Pa que Nacional Ba i u, Finca
Lipeo, a 15 km de Las Jun as, 600 m, 1 (CML); Rio EI
Na anjo, 14 km W de Ru a p o incialS, 1 (CML).
SANTIAGO DEL ESTERO: San o Domingo, 24 (7
CML, 5 IADIZA-CM, 12 OMNH). TUCUMAN: Aguas
Chiqui as, Sie as de Medina, 800 m, I (CM); A oyo
EI Sal on, Rese a P o incial San a Ana, 1 (CML);
Concepcion, I (MACN); Dique San Ignacio, 4 (3 CML,
1 OMNH); EI Cadillal, Usina, 3 (RMB); Las Talas, 4
km al N de Bella Vis a, 1 (OMNH); Mon eagudo, 4 (2
FCM, 1 MACN, 1TTU); Ticucho, 3 km E, I (OMNH);
Ticucho, en ando po cola del Dique EI Cadillal, I
(OMNH); T anqui as, 1 (MACN).
Addi ional Reco ds.- FORMOSA: Pa que
Nacional Rio Pilcomayo (S. Heinonen F. and R. M.
Ba quez, in li .). SALTA: EI B eal (Ma es e al., 1981);
Sal a (La Val, 1973); Sal a, casa habi ada (Villa-R. and
Villa-C., 197 I). SANTIAGO DEL ESTERO: La Banda
(Ro na ia and Abalos, 1950); La Pue a (Romaiia and
Abalos, 1950). TUCUMAN: Tucu nan, ciudad (Roma ia
and Abalos, 1950).
Myo is chiloensis (Wa e house)
Vespe ilio chiloensis Wa e house, Mammalia, In
C. Da win (ed.). The zoology o he oyage o RM.S.
Beagle, unde command o Cap ain Fi z oy, R. N., du -
ing he yea s 1832 o 1836.5 p s., 97 pp. Smi h, Elde ,
and Co., London, p. 5, 1840.
Myo is chiloensis: Cab e a, Re is a Chilena de
His o ia Na u al, 7:295, 1903.
Type Locali y.:-: The holo ype was om a small
island eas o he Isla de Chiloe, Chile. I has no been
loca ed in he BMNH and appa en ly is los (Mille and
Allen, 1928; La Val 1973; Ca e and Dolan, 1978). Con-
sequen ly, La Val (1973) designa ed a neo ype ( emale
FMNH 24029), om Cucao, Isla de Chiloe, Chile.
BARQUEZ ET AL.-THE BATS OF ARGENTINA
,
'.
1
I
93
Figu e 67. Skull and ee h o Myo is albescens. Adul emale om Dique San Ignacio, Tucuman
P o ince (OMNH 18877). Ba
=
I em. Tee h no o scale.
Disl ibulion.- Res ic ed o Chile by La Val
(1973), bu included in A gen ina by C espo (1974).
Cab e a (1958) conside ed M chiloensis a poly ypic
species, lis ing as subspecies M. c. al e and M. c.
chiloensis, among o he s no analyzed in his s udy; he
also conside ed M. dinellii as a synonym o M. c.
a acamensis. La e , La Val (1973) ecognized M
chiloensis and M. a acamensis as alid mono ypic spe-
cies, es ic ing he dis ibu ion o he o me o cen al
and sou he n Chile and he la e o he coas al dese o
no he n Chile and Pe u. La Val (1973) also ecognized
M. le is as a poly ypic axon, including as a subspecies
M. I. dinellii; M. c. al e was ecognized as a synonym
o M
I.
le is.
94
SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
761
346,347
75,392
391
Figu e 68. Map o he locali ies Myo is chiloensis.
Thus, La Val (1973) conside ed M. chiloensis o
be a mono ypic species es ic ed o Chile, bu wi h a
p obable ex ension o Isla Na a ino sou h o Tie a del
Fuego. This ex ension was based on a specimen in he
AMNH men ioned by Koopman (1967). This specimen,
as well as hose men ioned by Ol og (1950) in he CML,
and a specimen om Rio Douglas om Tie a del Fuego,
Chile, a e all M. chiloensis (R. M. Ba quez, pe sonal
obse a ion).
F om an examina ion o specimens om he p o -
inces o Chubu , Rio Neg o, and Neuquen, C espo
(1974) included M. c. chiloensis in A gen ina and ex-
ended he dis ibu ion o M. c. a acamensis (= M. le is
dinelliii o Isla Vic o ia, Neuquen P o ince whe e, ac-
co ding o him, he e is a clea sepa a ion be ween bo h
geog aphic o ns. We conside M. chiloensis and M.
le is o be dis inc species and do no ag ee ha he
o ne should be es ic ed o Chile, as s a ed by La Val
(1973). We also doub he ange ex ension o M. l.
dinellii gi en by C espo (1974), since we ha e no been
able o ind his specimens in he MACN.
We ha e examined many specimens o
M.
chiloensis om di e en locali ies. All a e qui e di e -
en om
M. l. dinellii,
bo h in appea ance and in c a-
nial and ex e nal measu emen s. Specimens om San
Ma in de los Andes, 71 k n SE, a e ligh e do sally han
o
200 400 km
I
I
I
specimens om o he locali ies, bu hey a e no com-
pa able o M. I. dinellii; he colo a ion may be a colo
phase, o seasonal o en i onmen al a ia ion. These
specimens esemble M. aelleni and ha e simila mea-
su emen s, bu di e in o he cha ac e s. We ha e no
been able o examine he ype and ype se ies o
M.
aelleni, bu om an examina ion o many specimens
om he egion, including some collec ed a he same
locali y, on he same da e, and by he same collec o as
hose o he ype o M. aelleni, we eel ha he alidi y
o M. aelleni is ques ionable.
All specimens examined pe ain o M. chiloensis,
including hose om El Hoyo ( he ype locali y o M.
aelleni). Al hough he c anial and ex e nal measu e-
men s o e lap hose o M. aelleni, he specimens Jack
he icolo ed do sal hai s ha cha ac e ize M. aelleni.
Dis ibu ion in A gen ina.- Figu e 68.
Desc ip ion (Fig.
69).-
Specimens om A gen-
ina ha e a o ea m leng h anging be ween 37.0 and
39.8 mm. Colo a ies om blond in he no h o da k
b own in he sou h. Do sal hai s measu e be ween 4
and 7 mm; he basal h ee- ou hs a e ligh b own o
black and he ips a e blond, ligh b own, o g ay-b own.
Ven ally, hai bases a e da k b own o black and he
ips a e ligh b own, g ay, o whi ish. The con as be-
ween he bases and he ips is mo e accen ua ed in speci-
BARQUEZ ET AL.-THE BATS OF ARGENTINA
,
101
"
,j
Figu e 73. Skull and ee h o
Myo is le is dinellii .
Adul male om Dique San Ignacio, Tucu nan
P o ince (CML 3162). Ba
=
1 em. Tee h no o scale.
Type Locali y.-
A gen ina, Tucu nan, 456 m.
Dis ibu ion in A gen ina.-
Figu e 74.
Measu emen s.-
Table 9.
Specimens Examined
(298).-
CATAMARCA:
Andalgala, 2 (CM); Andalgala, 4 km S, 3 (2 CM, I
MSB); Balnea io EI Caolin, 6 km NW Chumbicha, 4
(OMNH); Balnea io Municipal Capayan, Chumbicha,
I (OMNH); Chumbicha, 1km Nand W o balnea io by
oad, 2,600 ., 34 (9 CML, 9 IADIZA-CM, 16 OMNH);
Cues a del Cla illo, 5 km S La Bande i a, 2 (CML);
Cues a La Sebila, Hwy 60, 2 (CM); EI Rodeo, 900 m, I
(CML); EI Rodeo-La Jun a, 2 (CML); Es ancia Al o las
Jun as, 2 (CML); Las Jun as, Es ancia de los Figue oa,
50 km NW Ca ama ca Ciudad, 3 (OMNH); Molleci o,

102
SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
",;
~
..
",
37, 638, 664'''11'3, 295, 31:2 """"
,,9,~,??,~.
.376,377,593,629
308,309. -----':--------,
100,284."" !.325,728 :
'I':" -. :
176,568 ';
,,'
40, 4 ,758, 759 •• , ~
---""_'__~!?
,'26'0,413,414
'., 6'411-78":••• 2416~
",768 ~~--:"68 69 143
: •• 139 '>':",.491 ,
'---', " " ·442------ '
342
e" •
258
~
., " .172 "
425.
!;
36 .410, 652
!
:,757 :•• .166 .:
526. ,-__
'c, __."":,::,,:.
720 "
753 .·-' /""':575. 481~ .704
, • " .' 378
132,217,466 - ._"',586:: 387_":
97" 428" •• 636. •
• "
6:
I
4~_~/: ,
480, 583 ... : 66, 254, 596
:
~
-
-
-
- -
-
-
--;
!
'298.~
:
..
,
'
397.:
",,--,------,'.
:
103'<
Figu e 74. Map o he locali ies o Myo is le is dine/Iii.
30
k n
SE Andalgala, 3 (CML); Pues o F on e izo 6
k n
W La Bande i a, 6 (CML); Rio San Pablo, 3
k n
NW
Concepci6n, I (CM); Villa El Po e o, I (CML); Villa
EI Po e o (N An dalgala) , EI Po e o, 1 (CM).
CORDOBA: Baigo ia, 1 (CUNRC); Chancani, 1
(CUNRC); C6 doba, 9 (USNM); C uz del Eje, 1
(CUNRC); La Paz, 6 (MACN); Las Es ancias, 2 (CML);
Los Cisnes, 4 (MACN); Los Hoyos, 1 (MACN); Rio
,----------._.--..;
65
487 678,712
150~~ ,- ••433_----(
'l; '. --- ..
',_," ,."'- • _ ' 20, 372
: •• ; 771
38...,~
.---i-
28, 30
367~, " 320,321
621~ • ,:'
421-;-:.:.... ~744, 745, 746
- 518....;.l!'".
X
385
55-:"'~ ~422
190/'~ __... 163
/>: ~
198
o
100 200 400 km
I I I I
Cua o, 3 (SUVM); San Es eban, 1100 m, 3 (2 CML, I
OMNH); Tala Canada, 1250 m, 2 (MACN); Villa
Dolo es, 6 (TTU). LA PAMPA: Caleu Caleu, 8 (I
FMNH, 7 MACN); Gene al Pico, 1 (MACN); Laguna
Colo ada G ande, Caleu Caleu, 2 ( MACN). LA
RlOlA:
Chi1eci o, 6 (MACN); Es aci6n Pa quia, Guayapa, 1
(CML); San Bias de los Sauces, 4
k n
SE San BIas, 9
(CML); Villa Un
i
on, 9 (MACN). MENDOZA:
BARQUEZ ET AL.-THE BATS OF ARGENTINA 103
Table 9. Measu emen s ojMyo is le is dinellii, Myo is le is le is, and Myo is nig icans. Fo a desc ip-
ion o he measu emen s, see he Ma e ials and Me hods. Mean, s anda d de ia ion, and sample size a e
ollowed by ange.
Va iable
Myo is
le is dine/Iii
Myo is
le is le is
Myo is
nig icans
To al leng h
86.0±4.81,39 89.0 ± 7.78, 15 81.3±5.01,20
no -
970 72.0 - 99.0 74.0 - 92.0
Tail leng h
38.1 ± 3.95, 34 38.0 ± 4.08,15 35.2 ± 2.88,20
22.0 - 44.0 27.0 - 41.0 30.0 - 40.0
Hind oo leng h
75 ± I. II, 38 7.8 ± 1.30, 14 66 ± 0.72, 20
4.0 - 11.0 6.0 - 11.0 5.7 - 80
Ea leng h
138 ± 2.20,39 14.1 ± 1.27, 15 13.0 ± 1.87, 20
7.0 - 19.0 12.0-16.0 10.0 - 15.7
Fo ea m leng h
364 ± 092,39 369 ± 1.20, 16 33.7± 1.21, 19
34.3 - 38.5 35.0 - 39.3 32.0 - 36.2
Weigh
5.2 ± 0.93,31 7.0 ± 1.30,7 5.2 ± 1.08, 18
4.0 - 8.0 50 - 9.0 3.8 - 8.0
Condylobasalleng h
134 ± 0.35, 40 140 ± 0.34,15 12.7 ± 0.29,23
125 - 144 13.2 - 144 12.2 - 13.5
Leas in e o bi al b ead h
4.6 ± 0.15, 9 4.9 ± 0.16,15 4.6 ± 0.12, 19
4.4 - 4.8 4.7 - 5.2 4.4 - 4.8
Zygoma ic b ead h
86 ± 0.23, 26 9.2 ± 0.13,6 84 ± 0.22,10
83 - 9.0 9.1 - 9.5 8.2 - 8.8
G ea es leng h o skull
14.3 ± 0.29, 40 14.7 ± 0.27, 15 134±0.31,23
138-150 14.3 - 152 129-143
Pos o bi al cons ic ion
3.5±0.10,40 3.8±0.11,15 34 ± 0.12, 23
33 - 38 36 - 4.0 32 - 3.7
B ead h o b aincase
6.9 ± 0.25, 40 7.3 ± 0.30,15 6.5 ± 0.23, 23
64 -74 6.9 - 7.7 6.1 -7.1
Leng h o maxilla y oo h ow
54 ± 0.17, 39 5.5±0.15,15 5.0 ± 0.13, 23
5.2 - 5.9 5.3 - 5.9 4.8 - 54
Pala al leng h
7.1 ± 0.31, 32 7.2±0.21,14 63 ± 0.22, 19
66-79 7.0 - 76 60 - 6.9
Mas oidal b ead h
7.2±0.14,37 7.6±0.18,15 6.9±0.12,23
69 - 7.5 7.2 - 7.9 6.7 - 7.2
Leng h o mandibula oo h ow
58 ± 0.14,27 59 ±0.14, 15 5.3 ± 0.18,21
5.5 - 6.1 57 - 62 5.0 - 56
Leng h o mandible
104 ± 0.25, 37 10.8 ± 0.30,15 9.8 ± 0.20, 21
9.8 - 10.8 102 - 11.2 94 - 10 I
C-C (wid h ac oss canines)
3.5 ± 0.10,36 3.8 ± 0.13, 14 34±0.12,19
33 - 3 7 3.6 - 4.0 3.2 - 3.7
M-M (wid h ac oss mola s)
5.6 ± 0.18, 37 5.9±0.16,14 5.3 ± 0.13,22
5.2 - 60 5.8 - 63 5.1 - 56
Cacheu a, 2 (I BMNH, I FCM); Ce o de la Glo ia, 2
(IADIZA-CM); La alle, I (IADIZA-CM); Mala gue,
4 (MACN); Mendoza, 3 (I MCZ, 2 USNM); Nacuiian,
17 (16 IADIZA-CM, I OMNH); Rese a Ecol6gica
Nacu ian , I (IADIZA-CM); Rese a Tel eca, I
(OMNH); Uspalla a, 4 (MACN). NEUQUEN:
Neuquen, 400 m, 1 (BMNH). SALTA: Buena Vis a, I
(TTU); Cachi, I (CML); Campo San o, I (ROM); Gen-
e al Gi.iemes, Ru a 34 km 1363,2 (I ROM, 1 TCWC);
Guachipas, I (MACN); Guachipas, Iglesia, 2 (TCWC);
Gi.iemes, I (MACN); Hickman, 3 (CML); Ho cones, I
(ROM); La Me ced, 2 (TCWC); Rosa io de Le ma,
Escuela Gu uchaga, I (MACN); Sal a Capi al, 2 (I
TCWC, I TTU); San Lo enzo, I (TCWC); San a
Vic o ia Oes e, 2200 m, 2 (MACN); Uppe Cachi, I
(BMNH); Valle de Le ma, 1200 m, 1 (BMNH). SAN
JUAN: Ischigualas o, I (CML); Las Tumanas, along
Hwy 510 a c ossing o Rio Tumanas, I (IADIZA-CM);
104 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
i~"'",
,/,/
-,
,/'------"------------'1
:
,
'
,
'
'
I
i
,
,
,
/-------------------I
:' 1',"--
:
,
'
-----, j
i
y ,
,
'
, - -- -----,
, ,
.
"',
,
/ II ~
,
'
,
'
,( 691
7--'/
-,
~
,
.
"
'
,/ ' 256 •
" >, •
,/ ':-6~3·'"•••
,
"
/------
/ / 257"
__L '
119
438.
630,
455
721.
261.
437.
o
100 200
I I I
400 km
I
Figu e 75. Map o he locali ies o Myo is le is le is.
Pede nal, 1 (OMNH). SAN LUIS: La Toma, 13
(MACN); Naschel, 1 (ROM); Queb ada de Lopez, San
F ancisco del Mon e de O o, 1(OMNH); Saladillo, Rio
Quin o, 1(MACN); San Luis, 4 (MACN). TUCUMAN:
Agua Rosada, San Ped o de Colalao, 2 (CML); Aguas
Chiqui as, 25 km NE San Miguel de Tucuman, 800 m,
5 (CM); Amaicha del Valle, 2000 m, 1 (OMNH); A -
oyo El Sal on, Rese a P o incial San a Ana, 4 (CML);
Colalao del Valle, 2500 m, 1 (BMNH); Concepcion, 4
(3 BMNH, 1 FMNH); Dique Escaba, 3 (CML); Dique
San Ignacio, 1 (CML); Ho co Molle, Pa que Biologico
Sie a de San Ja ie , 10 (CML); Ho co Molle,
Residencia Uni e si a ia, 1 (CML); La Cienaga, 2500
m, 1 (BMNH); La Higue a, 1 (CML); La Rinconada, 1
BARQUEZ ET AL.-THE BATS OF ARGENTINA
(CML); Las Pa as, Aconquija, 1(MACN); Las Talas, 4
km al N de Bella Vis a, 3 (SIG); Leocadio Paz, 1
(MACN); No co, Vipos, 2500 m, 2 (BMNH); Pa que
P o incial EI Cochuna, km 40 sob e ni a 47, 1 (CML);
Rio Pueblo Viejo, Rese a P o incial La Flo ida, 2
(CML); San Ped o de Colalao, sou h o , a k n ma ke
42, on
Hwy
364, 4,700 , 13 (3 CML, 3 IADIZA-CM,
7 OMNH); Tacanas, 1 (CML); Ta i de Valle, 2200 m, 1
(CML); Tucuman, 400 m, 3 (BMNH); Tucu nan, 450
m,2 (MACN); Tucuman, 456 m, 2 (BMNH); Tucu nan,
no speci ic locali y, 1 (BMNH); Vipos, Es ancia San
Ped o,S (CML).
Addi ional Reco ds.- BUENOS AIRES: Bahia
Blanca (Baud, 1979). CATAMARCA: Las Jun as, 50
km NW Ca ama ca Ciudad (Ma es e aI., 1996) (= Las
Jun as, Es ancia de los Figue oa, 50 k n NW Ca ama ca
Ciudad). CORDOBA: Espinillo (Ti an i and To es,
1998); Paso del Du azno (Ti an i and To es, 1998);
Rio Cua o (Ti an i and To es, 1998); Segunda Usina
(Ti an i and To es, 1998); Villa Walca de (MVZ, no
examined). LA PAMPA: Agua de To es (San is and
Jus o, 1978). MENDOZA: Depa amen o San Ca los
(Roig, 1965); El Challao (Roig, 1965); Mendoza (MVZ,
no examined); Pues o Lima (Roig, 1965). SALTA:
Ca aya e (La Val, 1973; Ro na ia and Abalos, 1950);
Depa amen o San Ca los (Yepes, 1944); Finca La Rosa,
Ca aya e, 2000 m (Villa-R. and Villa-C., 1971); La
Me ced, 1200 m (La Val, 1973); Rio Blanco, 35 km
SW Sal a (Villa-R. and Villa-C., 1971); Sal a (La Val,
1973); Te nas de Rosa io de La F on e a (Ro naiia and
Abalos, 1950). SANTIAGO DEL ESTERO: Ojo de
Agua (Ro na ia and Abalos, 1950).
Myo is le is le is
(I.
Geo oy S .- Hilai e)
Vespe ilio le is I. Geo oy S .- Hilai e, Annales
des Sciences Na u elles, Pa is, se . 1,3 :444-445, 1824.
Myo is le is le is: La Val, Los Angeles Coun y
Museum Na u al His o y Science Bulle in, 15:38, 1973.
Type Locali y.-"Sou hem B azil."
Dis ibu ion in A gen ina.- Figu e 75.
Measu emen s.- Table 9.
Specimens Examined (81).- BUENOS AIRES:
Bahia Blanca, 1 (MACN); CanaliS, 1 (ROM); Capi al
Fede al, 9 (MACN); Chasco nus, I (MACN); Del a del
105
Pa ana de las Pal nas, 1 (TCWC); Es ancia El Casali o,
Ru a 226 ca. k n 98, 3 (MSU); Es ancia La Espe anza,
5 (CML); Ezpele a, I (FCM); La Pla a, 3 (MLP);
La alle, I (USNM); Lib es del Su , I (MACN); Lobe ia,
15 k n SW, 2 (MACN); Lobos, I (MACN); Maipu, I
(TTU); Ma del Pla a, I (ROM); Pun a Rasa, Cabo San
An onio, 1 (OMNH); Rancho Los Yngleses, Aj6, 10
(BMNH); Ru a 2 y puen e del Canal 2, 2 (MACN); San
Ped o, uel a de Rocha, I (MACN); Tandil, 2 (MACN).
ENTRE RIOS : B azo La go, 2 (TTU); Depa amen o
Gualeyguachu, 7 (TCWC); Es acion Medanos, 2 (TTU);
Es aci6n Pa anaci o, 2 (FCM); Islas de Ibicuy,
Pa anaci o, 4 (TTU). MISIONES: 6 k n NE by High-
way 2 o jc . Highway 2 and A oyo Pa aiso, I (CML);
Je . Hwy 2 and A oyo Pa aiso, 13 (8 CML, 5 IADIZA-
CM); Je . Hwy 21 and A oyo O eja Neg a, app ox. 2
km W Pa que P o incial Mocona, 1 (CML). SANTA
FE: San a Fe, 1 (MFA).
Addi ional Reco ds.- BUENOS AIRES: Gene al
La alle, 20 mi S (La Val, 1973). ENTRE RIOS: Pa que
Nacional EI Palma (Massoia and Di Lo io, 1981).
Myo is nig icans (Schinz)
Vespe ilio nig icans Schinz, Na u geschich e und
Abbildungen de Sauge hie e. Das Thie eich einge heil
nach dem Bau de Thie e als G undlage
i
h e
Na u geschich e und de e gleichenden Ana o nie on
dem He n Ri e on Cu ie . Sauge hie e und Vogel.
J. G. Co a'schen Buchhandlung, S u ga und
Tiibingen, 1:179, 1821.
M o is nig icans. Mille , No h Ame ican Fauna,
13:74, l897.
Type Locali y.- The ype specimen was collec ed
by Wied-Neuwied in Fazenda do Aga, nea he Rio
I i iba, Espi i o San o, B azil. This specimen is appa -
en ly los , as i has no been loca ed in he museums o
Eu ope (Ca e and Dolan, 1978), o in he AMNH
whe e a la ge pa o he Wied-Neuwied collec ion is
deposi ed (La Val, 1973). La Val (1973) designa ed as a
neo ype an adul emale (Los Angeles Coun y Museum,
No. 36877) om 42 k n S Rio de Janei o, B azil.
Dis ibu ion.- This species is widely dis ibu ed
in Sou h Ame ica. The only subspecies known om A -
gen ina is Myo is n. nig icans (Schinz), which ag ees
wi h Wilson and La Val (1974).
I
occu s in almos all
106
SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
! ••
, 273,684
i
688~~--495, 695:
(', ~ --''--776 :
, 1.,230 .:
i
-:>
57.
)402,403 -'" 619
'-,'
"
"-
251~ .34 ,,/
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77
_. - -, __
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699 ,.228 ""
'-' : '.see
:488 .110: "",
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i
120..' 300 597
"abo e' : '
'" 552•• 676:
•
351
-). ::.
383 -' 362 :-----------------,
--,-, ,-, • : : 169,394,405
, ••••- ••_•• ' • II : :
:,- 126 .313 : .:
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,
"
,
"
:' ,I ,//
.
./"
Figu e 76. Map o he locali ies o Myo is nig icans.
opical and sub opical o es s up o 3150 m, bu is
less common abo e 1200 m. I also is ound in sa an-
nas and b ushy habi a s. Two o he subspecies a e ound
in Sou h Ame ica (La Val, 1973): M. n. punensis occu s
on he wes coas om no he n Pe u o cen al Colom-
bia, and M. n. caucensis occu s in he Andes o Colom-
bia, Ecuado , and Pe u.
Dis ibu ion in A gen ina (Fig. 76).- Like M.
albescens, M. nig icans is widely dis ibu ed, bu cap-
u es a e ela i ely uncommon. Many specimens in a i-
ous collec ions we e ound o be misiden i ied o uni-
den i ied.
o
I
100 200
I I
400km
I
Desc ip ion (Figs. 77a, 77b, 77 , and 78).- La
Val (1973) and Wilson and La Val (1974) desc ibed his
species, bu he ollowing desc ip ion is based on A -
gen ine specimens. This species is highly a iable ex-
e nally and c anially. Do sal hai s a e gene ally
unicolo ed. In specimens om Tucu nan P o ince and
Depa amen o O an, Sal a P o ince, he ips a e sligh ly
pale , bu no as much so as in M. le is; hose om Pozo
Hondo, Es ancia el Guapo, San iago del Es e o, a e e en
pale han o he specimens examined. In hese, he do -
sal colo a ion is simila o ha o he ips o he hai s o
M. le is, bu he ma ked con as wi h he bases is lack-
ing. In specimens om Chaco and Co ien es, he

BARQUEZ ET AL.- THE BATS OF ARGENTINA
a
c
9
107
e
o>.
'.:
.:~.:
.
':".:"
.~~
h
Figu e 77. Compa ison o cha ac e is ics o Myo is: (a) on al iew o ace o Myo is nig icans; (b) la e al
iew o ace o Myo is nig icans; (c) agus o Myo is ube , (d) agus o Myo is ipa ius; (e) agus o Myo is
simus; ( ) agus o Myo is nig ieans; (g) u opa agium o Myo is a/beseens; (h) u opa agium o Myo is le is
dinellii; and (i) u opa agium o Myo is simus .
108 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
Figu e 78. Skull and ee h o Myo is nig icans. Adul male om Rio Po e io, Fo mosa P o ince
(OMNH 18889). Ba
=
J
em. Tee h no o scale.
bicolo a ion o he do sal hai s is dis inc , making hem
appea simila o
M.
ipa ius om Misiones and o some
a ypical specimens o M. albescens. Ven ally he hai s
a e bicolo ed, wi h he bases da k b own o black and
he ips pale ( a ying be ween yellow, c eam, o whi -
ish). The memb anes a e da k, almos black, and p ac-
ically naked.
The skull is small, he os um sho , and he sag-
i al c es no mally absen , al hough i may be p esen
in a ypical specimens. The p esence o a sagi al c es
and a P2 displaced o he lingual side o he oo h ow
a e cha ac e is ics o M. ipa ius. These cha ac e s no -
mally a e no ound in M. nig icans, al hough bo h spe-
cies can be con used ex e nally.
BARQUEZ ET
AL.-
THE BATS OF ARGENTINA
Appa en ly much geog aphic and indi idual a ia-
ion exis s. La Val (1973) indica ed ha in aspeci ic
a ia ion is mo e p onounced in leng h measu es, in a-
ios in ol ing measu emen s o skull wid h and shape,
and in colo o he pelage.
Measu emen s.- Table 9.
Commen s.- This is a well-s udied Neo opical
species, ye in o ma ion on i s axonomy and biology
in cen al and sou he n Sou h Ame ica is incomple e
and con used. Specimens om A gen ina sugges a a i-
able species wi h nume ous aces, bu insu icien ma-
e ial is a ailable o s a is ical analyses. Wilson (1971)
s udied his species on Ba o Colo ado Island, Panama,
and o he s udies ha e also examined i s biology (Ibanez
Ula gui, 1981; Ma es and Wilson, 1971; Mye s, 1977;
Willig, 1983; Wilson and Findley, 1970, 1971), bu none
e e o A gen ina.
This species is mos common in a eas modi ied
by people, bu i also occu s in o es s. We ha e col-
lec ed hem om oo s o u al dwellings, below b idges
(see commen s abou G. so icina), and benea h ee ba k
in Piqui enda Viejo, Sal a. J. R. Con e as (pe sonal
communica ion) cap u ed specimens om Capi an
Sola i, La Fo es al, Chaco, unde nea h oo s o u al
sheds cons uc ed o palm unks.
Wilson and Findley (1970) ound ha he ges a-
ion pe iod las s abou 60 days ( his is appa en ly is e-
la ed o ood a ailabili y) and weaning occu s abou he
second o hi d week. Males become sexually ac i e
a e 15 o 17 weeks. In he o es s o Sal a, we ound
ep oduc i ely inac i e males in Augus and ac i e ones
III
No embe .
Specimens Examined (107).- CHACO: Capi an
Sola i, La Fo es al, 2 (OMNH); El Mang ullo, 20 Ian
NNW by oad and 11 Ian NE by oad, 5 (2 CML, 2
IADIZA-CM, I OMNH); Gene al Vedia, I (TTU);
No he n Chaco, 1(BMNH); Rio de O o, Gene al Vedia,
3 (FCM); Taco Pozo, 77 Ian NE, sob e picada 20, 1
(CML). CORRlENTES: Co ien es, I (CML); I a i, 1
(BMNH); Laguna B a a, 2 (OMNH); Laguna Pai a,
BO Las Lomas, 16 (CML). FORMOSA: Bou ie , 2
(TTU); Clo inda, I (MACN); Es ancia Las Me cedes,
I (TTU); Es anislao del Campo, 2 (l ROM, I TTU);
Rio Po e io , 5 km S Es ancia San a Ca alina, 1
109
(OMNH). JUJUY: Laguna La B ea, I (CML); Laguna
La B ea, 25 km W Palma Sola, 4 (2 CML, 2 OMNH);
San a Ba ba a, I (AMNH). MISIONES: Colonia
Ma i es, Pue o Candela ia, 1 (MACN); Libe ad, 3
(MACN). SALTA: Alga obal, 20 km E Gene al
Gue nes, I (OMNH); Escuela Gu uchaga, Rosa io de
Le ma, 1 (TTU); Finca Ab a G ande, 1 (OMNH); O an,
15 Ian S, 15 km W, along Rio San a Ma ia, 1 (CM);
Piqui enda Viejo, 6 Ian W, 2 (I CML, I OMNH); San
Ramon de la Nue a O an, 2 (CML); San a Ma ia, sob e
Rio San a Ma ia, 1 (CML); Tonono, I Ian E sob e Rio
I iyu o, I (CML). SANTIAGO DEL ESTERO: Nue a
Espe anza, 15 (8 MACN, I TCWC, 6 TTU); Pozo
Hondo, Es ancia El Guapo, 2 (CML); San Ped o, 1
(TTU); San o Domingo, 21 (5 CML, 6 IADIZA-CM,
10 OMNH). TUCUMAN: Ache al, I (CML); Agua
Colo ada, I (TTU); Ce o del Campo, 900 m, I (CML);
El Cadillal, 25 Ian N San Miguel de Tucu nan, I (CM);
El Na anjo, I (TTU); Pied a Tendida, 12 Ian WNW
Bu uyacu along Rio Cajon, 2,500 ., I (OMNH); Senda
del Plu io ne o, 800 m, Ho co Molle, I (CML); Ta
Viejo, I (CML).
Addi ional Reco ds.-CATAMARCA: Ca a-
ma ca, Ciudad (Ro na ia and Abalos, 1950); La Pue a
(Ro na ia and Abalos, 1950). FORMOSA: Ba olome
de Las Casas (Massoia, 1970); Pa que Nacional Rio
Pilcomayo (S. Heinonen F. and R. M. Ba quez, in li .).
JUJUY: A oyo La U bana, 45 Ian E Y 5.4 Ian SE San
Sal ado de Jujuy, 620 m (La Val, 1973); El Monoli o
(Ol og, 1979); Yu o (Ol og, 1959). MISIONES:
Depa amen o Cainguas (Massoia, 1980); Depa amen o
Capi al (Massoia, 1980); Dos de Mayo, Pa aje Ian 51
(Massoia and Chebez, 1989); Los Helechos, Escuela n°
12 (Massoia e al., 1990). SANTIAGO DEL ESTERO:
Campo Gallo (Ro naiia and Abalos, 1950); EI Zanj6n
(Ro na ia and Abalos, 1950); He e a (Ro na ia and
Abalos, 1950); La Banda (Ro na ia and Abalos, 1950);
La F agua (Roman a and Abalos, 1950); Pozo Hondo,
Es ancia El Guapo (La Val, 1973); Sumampa (Ro na ia
and Abalos, 1950). TUCUMAN: Bu uyacu (Ro na ia
and Abalos, 1950); Mon e Bello (Ro na ia and Abalos,
1950); San Ped o de Colalao (Ro na ia and Abalos,
1950); Ta del Valle (Ro na ia and Abalos, 1950);
Tucuman, ciudad (Ro na ia and Abalos, 1950).
110 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
:"-
53, 174, 1'8~'-'- ,./" ,
----------, ·714 '.
,-'
,-----------,
.
" " • 699:
°":"'.·:531 :
'0""
,~:' : 157, 592 "
621< " :
,
,
,
55'--_-::': ( '
"
:
,-----,
,
y
Figu e 79, Map o he locali ies o Myo is ipa ius,
Myo is ipa ius Handley
Myo is simus ipa ius Handley, P oceedings o he
Uni ed S a es Na ional Museum, 112:466-468, 1960,
Myo is ipa ius: La Val, Los Angeles Coun y Mu-
seum o Na u al His o y Science Bulle in, 15:32-33,
1973,
Type Locali y,- Panama, Da ien, Villa Taca cuna,
Rio Puc o, 975 m. We ha e no modi ied he speci ic
name o ag ee in gende wi h he gene ic name
(Woodman, 1993),
Dis ibu ion.s-:
Acco ding o La Val (1973) and
Koopman (1993), he dis ibu ion includes mos o
Sou h Ame ica, excep wes e n Pe u, sou hwes e n
Boli ia, Chile, and mos o A gen ina,
Dis ibu ion in A gen ina.- Figu e 79,
Desc ip ion (Figs. 77d and 80),- Specimens om
A gen ina a e simila o
M,
nig icans om Chaco and
Co ien es. In con as o M, nig icans, M. ipa ius has
a sagi al c es and he P2 is displaced o he lingual
side o he oo h ow, A specimen om U uguay, 40 km
NW Tacua embo (CML) is simila o hose om
Misiones, al hough he pelage is edde . A gen ine speci-
mens a e cinnamon b own do sally, and he base o he
hai is da ke . The ips o many o he hai s a e b igh
o
I
400 km
--.J
yellow, gi ing a os ed appea ance, al hough his is less
p onounced han inM a/bescens. La Val (1973) epo ed
ha colo a ies om da k g ay o b igh cinnamon.
Ven al hai s a e bicolo ed, wi h he hai s da k b own
o black basally and he ips pale, c eam, o yellowish,
Wing memb anes join he body a he base o he pha-
langes. Ea s a e small, na ow, and he ips ounded;
he in e nal ma gin is s aigh ; he agus is s aigh o
he dis al one-hal and hen widens, becoming con ex
in he basal one-hal , whe e i e mina es in a small lobe.
The calca is well de eloped, ex ending abou one-hal
he dis ance be ween he ip o he ail and he oo ;
ibia a e long,
The skull is simila o o he Myo is, di e ing by
he p esence o a low sagi al c es and by he small P2
ha is displaced o he lingual side o he oo h ow. Be-
cause o his a angemen , he P2 is no isible in la -
e al iew, a ai ha also is ound in
M,
simus.
M,
ipa ius om A gen ina can only be con used (ex e -
nally) wi h some M nig icans , and wi h some M, ube
ha a e no b igh ed, al hough M ube is la ge .
Measu emen s,- Table 10,
Commen s.- The biology o his species is poo ly
known and no da a a e a ailable o A gen ina. B osse
and Cha les-Dominique (1990) ound his species o be
a e in F ench Guiana, bu epo ed i as widesp ead in
BARQUEZ ET AL.-THE BATS OF ARGENTINA
Genus Ep esicus
This genus, which is almos wo ldwide in dis i-
bu ion, con ains abou 33 species. Fi e species a e
known om Sou h Ame ica and h ee occu in A gen-
ina. I is simila o Myo is, bu la ge (excep E.
diminu us). Two cha ac e s a e use ul o ield iden i i-
ca ion: he absence o hai om he base o he
u opa agium, and he blun agus. The agus is poin ed
in Myo is. The snou o Ep esicus is mo e swollen han
ha o Myo is; I I and 12 a e unequal in size, and only
PI is p esen . In Myo is, he inciso s a e almos equal in
size and h ee uppe p emola s a e p esen . The dis ance
be ween he canine and he i s mola is g ea e in
Myo is han in Ep esicus, in which his space is educed.
The den al o mula is I
2/3,
C Ill, P
1/2,
M
3/3,
o al 32. The den al o mula, while di e ing om Myo is
(which has 38 ee h), is like His io us. I canno be con-
used wi h His io us, howe e , because o ex e nal and
c anial cha ac e s. The skulls o he h ee A gen ine spe-
cies o Ep esicus a e e y simila , di e ing mainly in
size and in he de elopmen o he sagi al c es .
Da is (1966) and Williams (1978) e iewed he
Sou h Ame ican species o Ep esicus, and Mye s e al.
(1983) sugges ha all h ee species a e sympa ic in
he Rio Pa ana d ainage basin. They can be dis inguished
by size, al hough o e lap may exis : E. b asiliensis is
la ge; E. u inalis is medium-sized; and E. diminu us is
small. Misiden i ica ions a e likely due o a ia ion in
cha ac e s such as colo and size, and because o sexual
dimo phism. Small males o E. u inalis can be con-
used wi h la ge emales o E. diminu us, and la ge e-
males o E. il inalis can be con used wi h small males
o E. b asiliensis.
Williams (1978) ound ha he bes cha ac e s o
species disc imina ion a e he leng h o he mandible
(measu ed om he ex e nal ace o he inciso s o he
pos e io poin o he angula p ocess) and he leng h o
he mandibula oo h ow (measu ed om he al eolus
o he canine o ha o he las mola ). When hese mea-
su emen s a e plo ed, he e is li le o e lap be ween
species. Howe e , many o he specimens examined a e
di icul o iden i y because hei cha ac e is ics a e in-
e media e be ween one species and ano he , especially
be ween E. u inalis and E. b asiliensis.
Villa-R. and Villa-Co (1969) iden i ied specimens
om Jujuy P o ince as E. innoxius using he key o
117
Da is (1965). La e , Villa-R. and Villa-Co (1971) e-
iden i ied hem as
Ei . u inalis,
ollowing Da is (1966).
G eenhall e al.
(1983b)
conside ed hese specimens and
hose inhabi ing he a id and semia id a eas om Jujuy
o Mendoza as E. innoxius, including a specimen om
Nacu ian s udied by Massoia (1976). Howe e , we be-
lie e hese specimens a e E. u inalis.
Ep esicus b asiliensis (Desma es )
Vespe ilio b asiliensis Desma es , Nou eau
dic ionnai e d'his oi e na u elle, appliquee aux a ,
p incipalemen
i
lag icul u e, e
i
I'economie u ale e
domes ique; pa une socie e de na u alis es. Nou elle
edi ion, p esqu en ie emen e ondue e
conside ablemen augmen ee. Ch. De e ille, Pa is,
35:478,1819.
Ep esicus b asiliensis: Thomas, Annals and Maga-
zine o Na u al Hiso y, se . 9, 5:367, 1920
Type Locali y.- B azil, Goias.
Dis ibu ion.- Fou subspecies a e ound in
Sou h Ame ica: E. b. melanop e us in mos o he Ama-
zon basin, he Guianas, and Venezuela; E. b. homasi in
he wes e n Amazon basin o B azil, Pe u, and Ecua-
do ; E. b. b asiliensis in eas e n B azil; and E. b.
a gen inus in sou he n B azil, Pa aguay, U uguay, eas -
e n A gen ina, and ex eme eas e n Boli ia. E. b.
a gen inus is he only one ound in A gen ina. The gen-
e al dis ibu ion gi en abo e is ha o Da is (1966),
al hough he ci es no Boli ian eco ds and la e s udies
do no men ion i in Boli ia (Ande son, 1997; Ande -
son e al., 1982; Ande son and Webs e , 1983). I s seems
o be a e in Pa aguay and was no ci ed o he coun y
by Mye s and We zel (1983), al hough Mye s e al.
(1983) eco ded i s p esence. The dis ibu ion o his
subspecies gi en by Da is (1966) appea s exagge a ed
conside ing he ew da a ha a e a ailable and e o s
ha occu ed in lis ing locali ies (below).
Ep esicus b asiliensis a gen inus Thomas
Ep esicus a gen inus Thomas, Annals and Maga-
zine o Na u al His o y, se . 9, 4:365, 1920.
Ep esicus b asiliensis a gen inus: Cab e a,
Re is a del Museo A gen ino de Ciencias Na u ales
"Be na dino Ri ada ia," e Ins i u o Nacional de
In es igaci6n de las Ciencias Na u ales, Ciencias
Zool6gicas, 4: 105, 1958.

118 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
,,,,,,
.,
,
"
.:
..•.•.........
-,
'''
55746-'",
"
,
:_----------~-------;
, ,
, ,
,
,
,
,
, ,
,
,
!
(~·305, 306
.
,
,
,
, ,
-;- --- : i
i -..-_..-..,
,
.
,
••••••. "
:
"
.•.
.
,
.
,
,
.-
:
/
Figu e 85. Map o he locali ies o Ep esicus b asiliensis a gen inus.
Type Locali y.- A gen ina, Co ien es, Goya.
Dis ibu ion in A gen ina (Fig. 85).- A e com-
pa ing specimens wi h he ype a he BMNH, only he
ype se ies and specimens om Puen e sob e el Rio
Be mejo, Chaco (Massoia, 1976) can be assigned o his
subspecies. The collec ion o skins a BMNH con ains
eigh specimens iden i ied as E. a gen inus (p obably
by Thomas): se en belong o he ype se ies. O hese,
ou a e nales (a pa a ype) and h ee a e emales ( wo
pa a ypes). Two o he males a e ela i ely young wi h
a el -like pelage, ca ilaginous phalanges, and o ea m
leng hs o 36.0 and 36.5 mm. The o he specimen is a
male om C uz del Eje, Co doba, iden i ied on he ag
as Ep esicus c . a gen inus which is E.
u inalis .
O he
specimens such as a male om Yu o, Jujuy (BMNH)
men ioned by Thomas (1920b) as E. hila ei, a emale
om La Maya, Co doba (MVZ), p obably hose men-
ioned by Massoia (1976) o Nacu ian (CEM),
Mendoza, and hose (MSB) om Po e o Ri e Dike,
Ca ama ca (Williams, 1978) a e he e ea ed as E.
u inalis.
Specimens examined om hese las locali-
ies a e indi iduals wi h measu emen s a he uppe
o
I400 km
I
ange o he measu emen s o
E. u inalis.
Specimens
in he CML collec ion om Huyamampa, San iago del
Es e o and San An onio de la Falda, Ca ama ca (Ol og,
1959) also pe ain o his species. The e is an appa en
e o in Da is (1966) in he placemen o he locali ies
o E. b. a gen inus; only wo locali ies in Co ien es
P o ince (Goya and Concepcion) a e men ioned, bu
he la e is placed in Tucuman P o ince. The locali y
name "Concepcion" exis s o owns in bo h Co ien es
and Tucu nan. We we e unable o ind specimens o E.
b. a gen inus om ei he loca ion a he BMNH whe e
specimens o
E. u inalis
om Concepcion, Tucu nan,
we e examined (and we e iden i ied by Da is in 1965).
Specimens om Gualeguaychu, En e Rios, in he
TCWC, TTU, and ROM collec ions a e E.
u inalis;
some a e he same specimens ci ed by Fo nes and
Massoia (1967) as E. b asiliensis.
Desc ip ion (Figs. 86g).- Size is la ge o A -
gen ine Ep esicus, wi h he leng h o he o ea m> 41
mm in adul s, g ea es leng h o skull> 16.7, and zygo-
ma ic b ead h> 12.0 mm. The do sal colo a ion is ligh
b own, wi h he bases o he hai s da ke . Some spec i-
BARQUEZ ET AL.-THE BATS OF ARGENTINA
-
9
119
de
h
Figu e 86. Ep esicus u inalis (a, b, c, d, , and h), Ep esicus b asiliensis (g), and Ep esicus diminu us (e and i): (a) ace,
on al iew; (b) ace, la e al iew; (c) u opa agium; (d) agus; (e) agus;
( )
hind oo ; (g) o ea m; (h) o ea m; and
(i)
o ea m.
120 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
mens o
E u inalis
om a id and semia id egions,
pa icula ly hose om Huyamampa, San iago del
Es e o, a e simila in colo o hose o he ype se ies o
E b. a gen inus om Goya.
In gene al e ms, he ex e nal and c anial cha ac-
e s a e simila o hose desc ibed o E u inalis; hey
can only be disc imina ed by size (see measu emen s).
Measu emen s.- Table 11.
Commen s.- Biology unknown. Da a in he li -
e a u e a e sca ce and iden i ica ion o he specimens
o which hey e e is unclea .
Specimens Examined
(9).-
CORRIENTES:
Goya, 5 (BMNH); Goya, 600 m, 4 (BMNH, including
he ype o Ep esicus a gen inus).
Addi ional Reco ds.- CHACO: Puen e sob e el
Rio Be mejo (Massoia, 1976). CORRlENTES: Es ancia
El Pila , 2 km de pa ada Ta a e (Ro naiia and To anzos,
1947).
Ep esicus diminu us Osgood
Ep esicus diminu us Osgood, Field Museum o
Na u al His o y, Zoological Se ies, 10:197, 1915.
Dobson (1885) desc ibed Vespe ugo (Vespe us)
do ianus om Misiones P o ince, bu Thomas (1920b)
said ha his was a synonym o
E u inalis.
Osgood
(1915) desc ibed E diminu us based on specimens om
B azil, and Thomas (1920a) desc ibed E idelis based
on specimens om Espe anza, San a Fe, A gen ina.
Cab e a (1930) synonymized E do ianus wi h E
u inalis
and main ained
E
idelis as a alid species;
la e , Cab e a (1958) conside ed E diminu us and E
idelis
o be dis inc species. Da is (1966) included
E
do ianus in he E innoxius species g oup. Using he
cha ac e s ha iden i y his g oup, such as small o e-
a m size, and he measu emen s o he specimens o
Dobson and hose o Thomas, Da is (1966) concluded
ha
E
do ianus and
E idelis
we e conspeci ic. Da is
main ained E do ianus as he alid name o he small
species om Sou h Ame ica and ecognized wo sub-
species; E d. do ianus (Dobson, 1885) and E. d.
diminu us. Da is conside ed E jidelis Thomas o be a
synonym o E do ianus.
Williams (1978) sugges ed ha he bes way o
ea he name E do ianus would be o designa e i as a
nomen dubium, based on he holo ype being oo la ge
o he small species o he genus. Williams (1978) u -
he sugges ed ha he measu emen s o he holo ype
ha he ob ained om he Museo Ci ico di S o ia Na u-
ale, Geno a, co esponded o E b asilensis. Howe e ,
he measu emen s gi en by Williams (1978) do no ag ee
wi h hose o Dobson, al hough Dobson did no p o-
ide skull measu emen s; he body measu emen s a e
oo la ge o E diminu us, oo small o E b asiliensis,
and a e only simila o hose o E u inalis. The c anial
measu emen s o he holo ype o E do ianus (Williams,
1978) co espond o E b asiliensis. The co ype o E
do ianus om San Ignacio, Misiones P o ince, in he
BMNH (BMNH 86.11.3.13), which was ecei ed in an
exchange wi h he Museo Ci ico di S o ia Na u ale,
Geno a, has no had he skull emo ed, bu he speci-
men is iden ical o
E u inalis
(R. M. Ba quez, pe -
sonal obse a ion). The specimen used by Dobson may
be a composi e specimen and in his case he nomen-
cla u e p oposed by Williams (1978) should be e ained:
E. do ianus would be conside ed nomen dubium and
he emaining a ailable names would be E diminu us
and E jidelis. Since E diminu us has p io i y, he wo
subspecies would be: E d. diminu us and E d.
idelis.
The la e is ound in A gen ina.
Type Locali y.- B azil, Bahia, Rio P e o, Sao
Ma cello.
Dis ibu ion.- The dis ibu ion is poo ly known,
o he e a e ew eco ds and he limi s be ween he wo
subspecies a e unclea . The gene al dis ibu ion o he
species includes no he n A gen ina, U uguay, and
sou he n B azil, ex ending no hwa d in B azil as a na -
ow s ip o app oxima ely 5° S la i ude. The no he n
a ea o dis ibu ion co esponds o E d. diminu us.
Ibanez Ula gui (1981) epo ed his species in Venezu-
ela, bu i s subspeci ic s a us is unclea in iew o i s
appa en isola ion om he o he popula ions.
Ep esicus diminu us idelis Thomas
Ep esicus
idelis
Thomas, Annals and Magazine
o Na u al His o y, se . 9, 5:366, 1920.
Ep esicus diminu us jidelis: Williams, Annals o
Ca negie Museum, 47:380-382, 1978.
Type Locali y:- A gen ina, San a Fe, Espe anza.
Dis ibu ion in A gen ina.- Figu e 87.
BARQUEZ ET AL.-THE BATS OF ARGENTINA
----
--
':"::,'.~;~, 30, 2:10
'.i.
362 : :
,-,,',"( :::
351
,
,
--',,,:' - ~ y
':253 ,:'---:
:'
..
,:
(.,
,','
/';-"
,-
~
-
--
1-- ... ,
:452:
------. :
Figu e 87. Map o he locali ies o Ep esicus diminu us.
Desc ip ion (Figs. 86e, 86i, and 88).- Specimens
om he Pa ana and U uguay i e d ainages a e ligh e
in colo and la ge in size han hose om no hwes e n
A gen ina; one indi idual om I a i, Co ien es P o -
ince (BMNH), is an excep ion. The ligh b own ips o
he hai s gi e a ligh , os ed appea ance o he do sum;
hai s o he en e a e ligh e , wi h he ips g ayish o
b own. Specimens om he p o inces o Sal a and
Tucuman a e smalle and ha e unicolo ed hai s. Those
om Buenos Ai es, Del a del Pa ana, and U uguay, can
be con used easily wi h specimens o M. a/bescens om
no hwes e n A gen ina because o hei os ed colo ,
he same as he ype o
Ec
idelis. In specimens ha ing a
os ed appea ance, he basal one-hal o he do sal hai s
is e y da k, almos black, and he dis al one-hal is
yellow, wi h he ips pale. Those om he No hwes
a e simila in colo o
E [u inalis,
bu can be dis in-
guished by size. The bes cha ac e is ics o iden i y his
species a e leng h o o ea m, leng h o he mandible,
and leng h o he oo h ow. In he specimens examined,
leng h o he o ea m a ied be ween 32.1 and 34.8 mm,
he leng h o he mandibula oo h ow be ween 4.8 and
5.4 nun, and he leng h o he mandible om 9.3 o
121
o
400 km
I I
10.1 mm. The ex e nal cha ac e s a e simila o hose
o
E
u inalis,
bu he calca is p opo ionally longe
(mo e han double he leng h o he oo ), he ips o he
ea s a e mo e ounded, and he agus has a less-de el-
oped basal lobe.
The skull is simila , bu less obus , and wi h less-
de eloped sagi al and lambdoidal c es s.
Measu emen s.- Table II.
Commen s.- The biology o his species h ough-
ou i s ange is almos unknown. We cap u ed a lac a -
ing emale and a ju enile male in he o es s o Aguas
Chiqui as, Tucuman, a he end o Janua y. Ibanez
Ula gui (1981) no ed ha all specimens ob ained om
Venezuela we e ound inside dwellings in open a eas.
S omachs con ained insec s, mainly coleop e ans. Speci-
mens om A gen ina inhabi a wide a ie y o habi a s
including g asslands, dis u bed u ban a eas, and dense
o es s.
Specimens Examined (16).- BUENOS AIRES:
Pa ana de Las Palmas y Canal 6, I (CML).
122 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
Figu e 88. Skull and ee h o Ep esicus diminu us. Adul male om Del a de Pa ana, Buenos
Ai es P o ince (CM L 1820). Ba
=
I cm. Tee h no o scale.
CORRlENTES: I a i, I (BMNH). JUJUY: Laguna La
B ea, 25 km W Palma Sola, I (CML). LA PAMPA: Luan
To o, 25 km S, 5 (El). SALTA: Agua Blanca, 24 km
NW, 1 (CM); Agua ay, 700 m,
J
(MACN); Queb ada
de Acambuco, 5 km W Dique l iyu o, I (RAO). SANTA
FE: Espe anza, 1 ( ype o E. id el is ; BMNH).
TUCUMAN: Aguas Ch
i
qu
i
as, 1 (CM); Aguas
Chiqui as, Sie as de Medina, 800 m, 2 (CM); EI
Cadillal, Es aci6n de Piscicul u a, I (CM).
Addi ional Reco ds.-
JUJUY: Palma Sola (Villa-
R. and Villa-C., 1971). MISIONES: Rio Vic o ia
(Massoia, 1980). SANTIAGO DEL ESTERO: La
Banda (Ro naiia and Abalos, 1950).

BARQUEZ ET AL.-THE BATS OF ARGENTINA
123
Table ll . Measu emen s o Ep esicus b asiliensis, Ep esicus diminu us, and Ep esicus u inalis. Fo a
desc ip ion o he measu emen s, see he Ma e ials and Me hods, Mean, s anda d de ia ion, and sample size a e
ollowed by ange,
Va iable Ep esicus b asiliensis Ep esicus diminu us Ep esicus
u inalis
To al leng h 108,8 i 6,27, 6 86, I i 3,68, 6 97,2 i 9.58, 41
1010 - 117,0 8 I,0 - 91.0 80,0 - 140,0
Tail leng h 438 ± 4.30,6 339 i 1.83. 7 40,0 i 3,81, 42
37,0 - 49.0 320 - 37,0 32,0 - 49,0
Hind oo leng h 9.lil.47,6 6.3 i 057,7 8, I i 1.0I, 39
80 - 11.0 56 - 7,0 6,0 - 10 I
Ea leng h 16,8 i 0,98, 6 13.8 i 0.48, 7 14,2 ±1.82,42
160 - 18,0 13,3 - J4,6 100-185
Fo ea m leng h 430il,91,6 332 ± 0.98, 8 39,8 i 1.54, 59
405 - 450 321 - 34,8 365 - 42,5
Weigh 59 i 0,65,5 10.6i2,22,31
4.9 - 65 70 - 140
Condylobasal leng h 15,8i031,5 12,7±0,19,7 14.7±0.47,51
J
5,6 - 16.4 12.4 - 130 136-157
Leas in e o bi al b ead h 6.4 i 0.47, 5 50 i 0.33, 6 5,7 i 0.31, 53
5,9 - 7,2 45 - 5.3 5.1 - 6,5
Zygoma ic b ead h 12.0 ± 0,07, 2 9, I ± 0,15, 5 10,6 i 0.43, 44
120 - 12,I 89-93 9,8 - lI8
G ea es leng h o skull 17,0 i 0,33, 5 132 i 022, 7
j
5.4 i 0.47, 52
16.7-17.4 129-136 143 - 16,3
Pos o bi a I cons ic ion 4.0 i 0,12,5 3,7iO.15,7 38i018,54
38 - 41 3.5 - 4,0 3,6 - 4.4
B ead h o b aincase 7,7i021,5 6,6iO.17,7 73 i 0,30, 52
7,5 - 8, I 65 - 7,0 67-80
Leng h o maxilla y oo h ow 6.4 i 0,20, 5 4.8 i 0, 13,7 5,H 0.19, 55
6,2 - 6.7 46 - 5,0 53 - 6,3
Pala al leng h 7,6 i 0,21, 5 61 ± 0.10,3 7,0 i 0.34, 41
7,5 - 80 6,0 - 62 6,5 - 7,9
Mas oidal b ead h 9, I i 0,30,5 7,2±0.19,7 8.4 i 0,35, 50
87 - 95 6.9 - 7,5 77 - 9.1
Leng h o mandibula oo h ow 6,8 i 0, 11,5 5IiO,21,6 6, I i 0.22, 54
6,7 - 7,0 48 - 5.4 5,6 - 68
Leng h o mandible 131 i023,5 98 i 0,30, 7 11.9 i 0,38,54
129-13,5 9.3 - JO I 110 - 130
C-C (wid h ac oss canines) 55 i 0, J5,5 4,0 i 0,12,7 48iO,21,51
53 - 5,7 3.9 - 4,2 4.4 - 5,6
M-M (wid h ac oss mola s) 7,3 i 0,23, 5 5,6iO,17,7 65 ± 0.29, 50
7.1 - 76 5,5 - 5,9 5.7 - 7.4
Ep esicus u inalis (dO bigny) les annees 1826, 1827, 1828, 1829, 1830, 1831, 1832,
e 1833. Tome Qua ie ne, 2e Pa ie. (A. d'O bigny, ed.).
Pi ois-Le au1 , e cie, Pa is, 413, 1847.
Vespe ilio
[u inalis
d'O bigny, Mammi e es. In
Voyage dans l' Ame ique Me idionale (Ie B esil, Ja
Republique o ien ale de U uguay, 1aRepublique A gen-
ine, la Pa agonie, la Republique du Chili, la Republique
de Boli ia, la Republique du Pe ou), execu e pendan
Ep esicus
u inalis,
Thomas, Annals and Maga-
zine o Na u al His o y, se . 9, 5:365, 1920.
Type Loca/i y.- A gen ina, Co ien es.
Dis ibu ion .-cp esicus
u inalis
is widely dis-
ibu ed in he Neo opics. I is absen om he egion
wes o he Andes in Pe u and Boli ia, and i is no
known om Chile o U uguay. Howe e , some o he
specimens om U uguay (Acos a y La a, 1950) may
be E.
u inalis
and no E. b asiliensis, based on mea-
su emen s gi en in ha publica ion. Da is (1966) in-
cluded h ee subspecies in Sou h Ame ica: E. gaume i
in he Guianas, no he n Venezuela, and Colombia no h-
wa d o Cen al Ame ica; E.
chapmani in he Ama-
zon basin in B azil, sou he n Colombia, and no he n
Boli ia; and E.
u inalis in A gen ina, sou he n Bo-
li ia, Pa aguay, U uguay, and sou he n and eas e n B a-
zil. Mies e a . (1996) ecognized six subspecies o
Sou h Ame ica: E. chopmani, E. chi alensis, E.
indleyi, E. u inalis, E. guame i, and E. mon osus,
wi h E. indleyi and E. i inalis occu ing in A gen-
ina. Williams (1978) had desc ibed E.
indleyi o
specimens om no hwes e n A gen ina eas o he
moun ains in he p o inces o 'Tucuman, Sal a, and Jujuy
based on cha ac e s o size and colo . Howe e , many
specimens examined om his egion exhibi cha ac-
e s o E.
[u inalis a he han E.
indleyi,
and we
conside he axa o be synonymous. Analyses o la ge
se ies om mo e locali ies indica e ha his is a highly
a iable species, hus di e ences in size and colo a e
insu icien o suppo he c ea ion o a di e en sub-
species.
The do sal colo a ion is highly a iable. Speci-
mens om San iago del Es e o can be ligh e han hose
om Tucuman, which would con adic he diagnos ic
cha ac e s o
E.
indleyi.
On he o he hand, colo
may be in luenced by he en i onmen , o specimens
om a id and semia id zones a e gene ally pale, while
hose om o es s a e da k. E. in dley
i,
acco ding o
Williams (1978), is la ge han he o he subspecies.
In
specimens om Tucu nan P o ince, he leng h o he
o ea m anges om 39.3 o 41.1 mm, bu in a se ies
om Huyamampa, San iago del Es e o P o ince, he
maximum measu emen is 42.6; he leng h o he o e-
a m is g ea e han 40 mm in 11 o he 13 indi iduals
examined. Mye s and We zel (1983) compa ed 17 speci-
mens om Jujuy P o ince (assigned o E. indleyi by
Williams, 1978) wi h a se ies om Pa aguay. They
ound ha hose om eas e n Pa aguay we e la ge and
da ke , and hose om he cen al chaco we e ligh e ,
bu wi h a iable do sal colo a ion whe e he hai s a e
banded and he ips ligh e . Consequen ly, hey could
no jus i y ea ing specimens om Jujuy and he cen-
al chaco as di e en subspecies. We ound he same
pa e n when examining specimens om h oughou
hei dis ibu ion in A gen ina. The cha ac e s o bo h
subspecies o e lap conside ably, such ha one can as-
sign specimens a any poin along he ange o one o
ano he subspecies. This si ua ion implies ha ei he
only a single subspecies is p esen , o ha wo e y simi-
la subspecies a e in comple e sympa y.
Ep esicus
u inalis u inalis
(d'O bigny)
Vespe ilio u inalis d'O bigny, Mammi e es.
In
Voyage dans I' Ame ique Me idionale (Ie B esil, la
Republique o ien ale de U uguay, la Republique A gen-
ine, la Pa agonie, la Republique du Chili, la Republique
de Boli ia, la Republique du Pe ou), execu e pendan
les annees 1826, 1827, 1828, 1829, 1830, 1831, 1832,
e 1833. Tome Qua ieme. 2e Pa ie. (A. d'O bigny, ed.).
Pi ois-Le aul , e cie, Pa is, 4:13,1847.
Ep esicus
u inalis
[u inalis.
Da is, The Sou h-
wes e n Na u alis , 11:265, 1966.
Type Locali y.- A gen ina, Co ien es.
Dis ibu ion in A gen ina (Fig. 89).- Cab e a
(1958) conside ed Vespe ugo do ianus Dobson a syn-
onym o his species. Williams (1978) ea ed i as
nomen dubium (see discussion in E. di ninu usi, bu
he co ype om he BMNH is he e included in E.
u inalis.
Desc ip ion (Figs. 86a, 86b, 86c, 86d, 86 , 86h,
and 90).-Size medium o A gen ine Ep esicus, wi h
he leng h o he o ea m a ying om 37.0 o 42.6 mm,
and a ely exceeding 41 mm. Colo a ies h oughou
he ange, bu is uni o m wi hin popula ions. Specimens
om Fo mosa ha e a da k b own do sum and he bases
o he hai s a e sligh ly ligh e han he ips. Those om
Tucu nan and Sal a p o inces a e simila o hose om
Fo mosa P o ince, especially in en al colo a ion, bu
he hai s o he do sum a e s ongly bicolo ed. The bases
a e almos black and he ips a e ligh b own. Indi idu-
als om San iago del Es e o P o ince a e e en ligh e
and he ips o he do sal hai s a e yellowish b own.
The colo o he en e is simila in almos all speci-
mens examined; he bases o he hai s a e da k b own
o black and he ips a e yellow b own. The memb anes
a e da k, wi h black in s in some specimens. The pel-
age does no ex end o e he memb anes on he do sal
124
BARQUEZ ET AL.-THE BATS OF ARGENTINA
13,43
22
494 : 537,538
695~"15 e736:
598, , :
~"B88e~..-776
i
<: ~-,~
I
,
402,403: ,e 567
431 ,J~07 L' ~ -.
- '~,,-1 /625 ',-,
_ 30,53 499,/ """
-----------, 286 .: -,
,_: ~ e
,-----!----------,
-'157,592
"419,' : -,
745,747', "",- __--.' : e 228 '-',
163~""e
i
e 653 : -",
55
<,~
j .:
e 676
i -.
~~/ e330 :
546e ' 301e
i
'-'- '422
,:-----------------i
'--', ~46e : e
! :
647':e :773313 •• 39: :
"':68, 143,145'-' _, .... 710 769/
'
: e768 " '~_---__e e: '
.._--......... ' / --...~------ ' :
-. e 258
/' ' ,
"",,>-,-------',:
: el72 :
"
,
"
,
:, : e84 .:
.>
.__~_..._~_l.. (
'--' i
",~378 , 737:
, : -,e :
': ,: 374e,:'
,
'
,)..
480e // //-,.: """
~
:
,--'-------'
1
----------l
, ,
'
/ 1
I
--------c-~-22-~.
452
'---
,
,
...----T-~;
:
...•.•.......
,//'
/,,,,,,
/./"
..
,
125
"
Figu e 89. Map o he locali ies o Ep esicus u inalis.
side, excep o e he base o he u opa agium, bu ne e
as much as in Myo is. Ea s a e medium-sized and well
sepa a ed, ha e ounded ips, and ha e an in e nal keel
ex ending om he uppe conca i y o he conch hal -
way up he an e io ma gin; he uppe one-hal o he
ex e nal ma gin o he ea s is s aigh and lowe one-
o
I
200
I
400
km
I
hal is con ex. The agus is well de eloped, blun , and
sligh ly widened a he base. The lips a e inged wi h
ine hai s and, excep o a iangula -shaped papilla on
he lowe lip, papillae a e absen . A dis inc i e wa
co e ed wi h long, ine black hai s can be obse ed in
esh o alcoholic specimens. In d y specimens, his
126 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
s uc u e is seen only as a u o long, black hai s. The
u opa agium is wide and con ains he ail, which ex-
ends beyond he dis al ma gin abou 3.5 o 4.0 mm.
The calca is well de eloped, sligh ly mo e han double
he leng h o he oo , and wi h a leshy la e al expan-
sion a i s mid-poin .
The skull is obus , i s g ea es leng h be ween
15.2 and 16.5 mm; he os um is sligh ly mo e han
one-hal he wid h o he b aincase; he sagi al c es is
isible (gene ally mo e de eloped in adul emales) and
mo e ele a ed on he pos e io one- hi d o he skull.
Lambdoidal c es s a e e iden and zygoma ic a ches a e
hin bu s ong and sligh ly widened medially. An e i-
o ly, he pala e has a deep in agina ion ha sepa a es
he inciso s o each side; pos e io ly, he pala e ex ends
beyond he las mola . The p e ygoids a e well de el-
oped, and he ympanic bullae a e la ge and co e mo e
han one-hal o he cochlea.
The 11 is well de eloped and 12 is small, i s ip
ba ely eaching he cingulum o he II. A small space
abou equal o he leng h o 12 is p esen be ween he
inciso s and he canines. Canines a e no mal, bu wi h
a pos e io keel aligned wi h he oo h ow. P I is close
o he canine and well de eloped. The M I and M2 a e
simila in appea ance and size, wi h ou commissu es
o ming he "W" pa e n ypical o insec i o ous mo-
la s; he p o ocone is well de eloped, bu he hypocone
is educed o absen ; M3 is educed wi h only h ee
commissu es; he p o ocone is p esen bu he hypo-
cone is absen .
Lowe inciso s a e ilobed, o e lapping and com-
ple ely illing he space be ween he canines. The ca-
nines ha e a pos e io p ojec ion ha is in con ac wi h
p I. The p I is abou one-hal o he size o p2, bo h wi h
a single well-de eloped cusp; p2 has a small me aconid
in con ac wi h he pa aconid o m!. The mola s a e
s uc u ally iden ical, wi h all he cusps well de eloped,
pe mi ing a clea dis inc ion o he alonid and igo-
nid; mola s di e only in size, he second being he la g-
es and he hi d he smalles . The inciso s o ju eniles
a e occasionally bi id, a condi ion a ely obse ed in
adul s.
Measu emen s.- Table II.
Commen s.- The biology o his species is poo ly
known. Pa asi es o his species we e s udied by Roma ia
and Abalos (1947) om specimens collec ed om a i-
ous dwellings. The specimen om Colonia k.m 503, 16
k.m S, Fo mosa P o ince, was cap u ed unde he ba k
o a ee (Ba quez and Ojeda, 1992). This species has
been aken in many habi a s, including we o es , gal-
le y o es , chacoan ho n sc ub, and ansi ional o es
(Ma es e al., 1995). B osse and Cha les-Dominique
(1990) ound
E.
gaume i in dis u bed and subu ban
a eas in F ench Guiana. Rep oduc i e pa e ns s udied
by Mye s (1977) in Pa aguay indica e wo pe iods o
ep oduc i e ac i i y--one in mid-July and Augus and
a second sho e pe iod in Janua y. Pos pa um copula-
ion and o ula ion also we e de ec ed; lac a ing emales
we e ound o be p egnan . In Rio Po e io, Fo mosa,
we ound a male wi h sc o al es es in mid-May and a
Angos o del Pescado, Se ania de las Pa as, Sal a P o -
ince, in mid-No embe . Lac a ing emales we e ound
in mid-No embe a Angos o del Pescado, Se ania de
las Pa as and a Queb ada de Acambuco, 5 k.mW Dique
I iyu o, Sal a P o ince. This species is known o be
p eyed upon by S ygian owls (Asio s ygius) in he
ce ado sa anna o B azil (Mo a Junio and Taddei,
1992).
Specimens Examined (21 0).- BUENOS AIRES:
Canal 6, Del a bonae ense, I (FCM); La Pla a, 4 (I
BMNH, 3 MLP); Los Cisnes, Del a del Pa ana, 5
(BMNH); Pla a ios, I (MACN). CATAMARCA:
Balnea io EI Caolin, 6 k.mNW Chumbicha, 8 (OMNH);
Chumbicha, I k.mNand W o balnea io by oad, 2,600
.,4 (I CML, I IADIZA-CM, 2 OMNH); Chumbicha,
a balnea io, 2,400 ., I (OMNH); Po e o Ri e Dike,
EI Po e o, I (MSB); San An onio de la Falda, I (CML).
CHACO: EI Mang ullo, 20 k.m NNW by oad and II
km NE by oad, 40 (21 CML, 16 IADIZA-CM, 3
OMNH); Rio Teuco, 10 km W Ta agal, 2 (CML).
CORDOBA: Biale Masse, 2 (MACN); C uz del Eje, I
(BMNH); La Maya, 45 k.mSE Bell Ville, I (MVZ); La
Paz, 3 (MACN); Villa Walca de, I (MVZ).
CORRIENTES: San Bo gi a, 3 (1 BMNH, 1MACN, I
TTU); San Bo gi a, Es e o de Ibe a, 5 (CEM). ENTRE
RIOS: Del a, B azo La go, I (CEM); Gualeguaychu, 5
(2 ROM, 2 TCWC, I TTU). FORMOSA: Colonia k.m
503, 16 k.m S, I (CML); EJ Chu cal, app ox. 10 k.m
SSE Buena Vis a, 3 (I CML, 2 IADIZA-CM); Es ancia
Las Me cedes, I (MACN); Pues o Di isade o, 35 k.m
S, 5 k.m E Ing. Guille mo N. Jua ez, I (CML); Rio
Be mejo, 10 mi S Colonia km 503, I (CML); Rio
Po e io, 5 km S Es ancia San a Ca alina, 2 (CML).
JUJUY: Laguna La B ea, I (CML); Laguna La B ea,
25 km W Palma Sola, I (CML); Ledesma, I (AMNH);
BARQUEZ ET AL.-THE BATS OF ARGENTINA
Banda (Ro na ia and Abalos, 1950). TUCUMAN: Las
Jun as, 22 km W Cho omo o (Ma es e aI., 1995) (=Las
Jun as, 22 km W Cho omo o on Hwy 312, 3,500
..).
His io us mac o us mac o us (Poeppig)
Nyc iceius mac o us Poeppig, Reise in Chile, Pe u
und au demAmazonens ome wah end de Jah e 1827-
1832. F ied ich Fleische , Leipzig, IA5 I, 1835.
His io us mac o us mac o us: Cab e a, Re is a del
Museo A gen ino de Ciencias Na u ales "Be na dino
Ri ada ia," e Ins i u o Nacional de In es igacion de las
Ciencias Na u ales, Ciencias Zoologicas, 4:108,1958.
Type Locali y.- Chile, Bio Bio, An uco.
Dis ibu ion in A gen ina (Fig.
94).-
Cab e a's
(1930) eco d o H. mac o us om he Depa men o
Belen, Ca ama ca P o ince, p obably co esponds o his
subspecies, al hough we ha e no ound specimens om
ha locali y among he specimens examined.
Desc ip ion (Fig.
95).-
The band be ween he
ea s seems o be a iable, bu in he specimens exam-
ined i is less e iden han in H. m. laepho is. The a-
gus is well de eloped, as in he o he subspecies. The
gene al colo a ion is simila o ha o H. mon anus om
he same a ea. The hai s a e s ongly bicolo ed: do sal
hai s ha e black o da k b own bases and yellowish ips;
en ally, bases o he hai s a e da k b own and he ips
a e whi ish, and in some specimens his is p onounced
( hese specimens esemble H mon anus ex e nally, al-
hough he size o he ea s dis inguishes he wo spe-
cies in li ing o esh specimens).
The skull is mo e obus and he os um wide
han inH. m. laepho is o H mon anus. F om he o me ,
hey a e di e en ia ed by he ex e nal b ead h ac oss
he hi d uppe mola s being g ea e han 7 mm. F om
he la e , his subspecies can be dis inguished by i s
g ea e size and la ge ympanic bullae.
Measu emen s.- Table 12.
Commen s.- This subspecies gene ally has been
con used wi h H. m. laepho is and H mon anus, hus
he da a ha can con iden ly be aken om he li e a-
u e a e sca ce. Ma es e al. (1995) eiden i ied many
specimens om no hwes e n A gen ina as Hm.
mac o us and cla i ied he dis ibu ion o his species
in wes e n A gen ina. Specimens om Rio Quin o, San
133
Luis P o ince (MACN and TTU) we e cap u ed in a
na u al ca e 8 m deep by 3 m wide, whe e hey we e
oos ing wi h Tada ida b asiliensis and Myo is sp. All
we e p egnan emales and one (MACN) had wo ull-
e m e uses acco ding o no es made by he collec o
on he skin ag. Pea son and Pea son (1989) ound a
colony o hese ba s in an a ic a Es ancia EI Condo ,
Rio Neg o P o ince, du ing he summe mon h o De-
cembe ; p egnan and lac a ing emales, and nu sing
young we e included in he small colony (con aining
abou 20 ba s). A isi o he same colony 5 yea s la e
ound h ee adul emales, wo wi h a ached young.
Specimens Examined
(56).-
CATAMARCA:
Andalgala, I (MACN); Balnea io EI Caolin, 6 km NW
Chumbicha, 2 (OMNH); Choya, 13 km NNW o
Andalgala, 4,000
.,
I (OMNH); Chumbicha, I km N
and W o balnea io by oad, 2,600 ., 8 (3 CML, 2
IADIZA-CM, 3 OMNH); Cues a del Cla illo, 5 km S
La Bande i a, 5 (CML); Dique EI Po e o, 1 (CM); El
Rodeo, 900 m, I (CML); La Bande i a, 1 (CML); La
Bande i a, Ru a 62 camino a Las Es ancias, I (CML);
Las Es ancias, I (CML); Los Naci nien os, u a 40, km
869, I (CML); Po e o Ri e Dike, El Po e o, 4 (MSB);
Sie a de Ancas i, Ru a 2, 1700 m, I (CML); Tinogas a,
1 (MACN). CORDOBA: Cu a B oche o, 1 (MACN);
Rio Cua o, 1 (MACN). JUJUY, T es C uces, 8 km al
SE, 3600 m, I (CML); T es C uces, 8 km S, sob e Ru a
9,2 (CML). NEUQUEN: Villa La Angos u a, 19 km N
along Hwy. 234, 3 (I CML, 2 IADIZA-CM).
Rio
NE-
GRO: Es ancia EI Condo , 22 km ESE Ba iloche, 7
(MVZ). SALTA: Ca aya e, 20 km NW, 1700 m, I
(CML). SAN JUAN: Complejo As on6mico EI
Leonci o, 7 km W, I (OMNH). SAN LUIS: Paso del
Rey, I km N, along A oyo de la Canada Honda, I
(OMNH); Rio Quin o, I (TTU); Saladillo, Rio Quin o,
2 (1 MACN, 1 TTU). TUCUMAN: EI Nogala , Ru a
307, 1700 m, 4 (CML); Ru a 307, km 43, 1700 m, I
(CML); Ru a 307, km 46, 1700 m, I (CML).
Addi ional Reco ds.- CATAMARCA: Depa -
amen o Belen (Cab e a, 1930).
Rio
NEGRO: Ba iloche,
20 km E (Pea son, 1995).
His io us magellanicus (Philippi)
Vespe ilio magellanicus Philippi, A chi u
Na u geschich e, 32(1): 113, 1866.
His io us magellanicus: Thomas, Annals and
Magazine o Na u al His o y, se . 8,17:273,1916.

134 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
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Figu e 94. Map o he locali ies o His io us mac o us mac o us.
200
I
400 km
I
BARQUEZ ET AL.-THE BATS OF ARGENTINA 135
Figu e 95. Skull and ee h o
His io us mac o us
mac o us.
Ba
=
1 em. Tee h
no o scale.
136 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
o
400 km
I I I
Figu e 96. Map o he locali ies o His io us
magellanicus.
Type Locali y.- S ai s o Magellan.
Dis ibu ion (Fig. 96).- His io us magellanicus
is known only om sou he n Chile, Tie a del Fuego,
and Neuquen P o ince. Cab e a (1958) epo ed i om
sou he n San a C uz P o ince, bu specimens examined
om his a ea a e
H.
mon anus. Osgood (1943) lis ed
his species as a subspecies o H. mon anus (H.
mon anus magellanicus).
Few specimens o his subspecies a e deposi ed
in collec ions. We ha e only been able o examine i e
specimens, h ee ha we collec ed in Neuquen P o -
ince in 1991. The specimen om Es ancia Viamon e,
Tie a del Fuego, A gen ina (BM 30.10.9.1) is a young
indi idual.
Desc ip ion (Figs. 92b and 97).- Do sal colo a-
ion is da k b own, wi h hai s da ke a he base and
ligh e a he ip. Ven ally, he bases o he hai s a e
almos black, and he ips yellowish o c eamish, con-
as ing s ongly wi h he da k base. All memb anes and
ea s a e da k b own. The ea s a e well sepa a ed, wi h-
ou any ace o a connec ing band, a cha ac e ha
clea ly di e en ia es his species om all o he species
o His io us in A gen ina. The agus is well de eloped,
slende , and almos one-hal he leng h o he ea . The
skull and ee h a e simila o o he species o he genus.
Measu emen s.- Table 13.
Commen s.- The specimens om Neuquen P o -
ince we e cap u ed in he No ho agus ain o es , along
wi hH. mac o us mac o us, Lasiu us a ius, and Myo is
chiloensis. All indi iduals we e adul s, and one emale
was p egnan (one e us = 19 mm CRL). The o he wo
we e males, one wi h la ge sc o al es es, in he sum-
me mon h o Decembe .
Specimens Examined (5).- NEUQUEN: Villa La
Angos u a, 19 k n N along Hwy. 234, 3 (I CML, I
IADIZA-CM, I OMNH). TIERRA DEL FUEGO:
Es ancia Viamon e, I (BMNH). NO SPECIFIC LO-
CALITY: Pue o P a , Las Hope Inle , Pa agonia, 5
m, I (BMNH).
His io us mon anus (Philippi and Landbeck)
Vespe ilio mon anus Philippi and Landbeck,
A chi ii Na u geschich e, 27(1):289,1861.
His io us mon anus: Mille , Bulle in o he Uni ed
S a es Na ional Museum, 57:214, 1907.
Type Locali y.- Chile, Co dille a de San iago.
Dis ibu ion.- Acco ding o Koopman (1993),
his species is dis ibu ed in a na ow s ip ha ex ends
sou hwa d om wes e n Venezuela and Colombia,
h ough cen al Ecuado and Pe u, o sou hwes e n Bo-
li ia, widening o include A gen ina, U uguay, and
Chile. As discussed in he H. mac o us accoun , speci-
mens examined ha came om no hwes e n A gen-
ina a e no H. mon anus; consequen ly, he idea ha
he dis ibu ion is con inuous om no h o sou h is no
co ec . A e-examina ion o specimens om o he coun-
ies would help cla i y he si ua ion.
Dis ibu ion in A gen ina.- Figu e 98.
Desc ip ion (Figs. 92c, 99a, and 100).- Gene al
appea ance is simila o H. mac o us mac o us, bu he
size o he body and he leng h o he ea s is less. In
adul s, he a io o he leng h o he head and body and
he leng h o he ea no mally anges be ween 2.3 and
BARQUEZ ET AL.-THE BATS OF ARGENTINA 137
Figu e 97. Skull and ee h o His io us magellanicus. Adul emale om Villa
La Angos u a, Neuquen P o ince (CML 3231).
138 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
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I
400 km
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BARQUEZ ET AL.-THE BATS OF ARGENTINA
Figu e 99. Face o : (a) His io us mon anus mon anus and
(b) His io us ela us.
Figu e 98. Map ( acing page) o he locali ies o His io us
mon anus mon anus
139
3.1. The skull is simila o H. mac o/us, bu is smalle ,
and has smalle ympanic bullae ha a e less han 4.0
mm in leng h. Colo is simila o H. mac o/us mac o/us,
bu he ips o he hai s o he en e a e whi e in
H.
mon anus. The band be ween he ea s is no mally ab-
sen , bu may be p esen in some indi iduals. As is he
case wi h he majo i y o cha ac e s o his and he o he
species o he genus, he ex en o indi idual and geo-
g aphic a ia ion is unknown.
Measu emen s.- Table 13.
Commen s.- Pea son and Pea son (1989) s ud-
ied his species in wes e n Pa agonia. They ound se -
e al oos s in he a ics o houses. Rep oduc i ely ac-
i e emales (wi h ull- e m e uses) we e collec ed in
ea ly No embe and ep oduc ion con inued in o a leas
Decembe . The Pea sons es ima ed ha emales do no
ep oduce be o e one yea o age. These ba s a e occa-
sionally ea en by bam owls, Ty o alba (Massoia e al.,
1993).
Specimens Examined
(56).-
BUENOS AIRES:
Balca ce, 2 (MSU); Buenos Ai es, I (BMNH); Sie a
de la Ven ana, I (MACN). CHUBUT: EI Mai en, 2 km
N, I (MVZ); Leleque, 7 (MVZ); Pico Salamanca, I
(BMNH). CORDOBA: EI Ca izal, C6 doba Range,
1000 m, 2 (BMNH); Espinillo, 2 (SUVM); La Cumb e,
I (CML); Punilla, Valle He moso, I (TTU); San
Es eban, 1100 m, 3 (I CML, 2 OMNH). LA RIOJA:
Aimogas a, I (MACN); San Isid o, I (MACN); Villa
Uni6n, 5 (MACN). MENDOZA: Mendoza, 2 (l
BMNH, I MCZ); Zanj6n Be mejo, I (MACN).
NEUQUEN: Codihue, 4 (MACN); Isla Vic o ia, I
(CML); Neuquen, 2 (I BMNH, I MACN).
Rio
NE-
GRO: El Bols6n, I (MACN); El C6ndo , Lago Pe i o
Mo eno, 800 m, I (MVZ); Lago Pe i o Mo eno, I
(MVZ). SANTA CRUZ: Laguna G ande, II (MVZ);
Pun a Loyola, S. Pa agonia, I (BMNH); San Julian, I
(MACN). SANTIAGO DEL ESTERO: San Sal ado ,
I (FCM).
Addi ional Reco ds.- BUENOS AIRES: Laguna
Chasic6 (Con e as, 1973); Villa La Ven ana (Massoia
e aI., 1993). CHUBUT: Gol o San Jose (Daciuk, 1977).
NEUQUEN: Las Colo adas, 1000 m (Fo nes and
Massoia, 1967). SANTIAGO DEL ESTERO: San Ped o
de Guasayan (Ro na ia and Abalos, 1950).
140 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
Figu e 100. Skull and ee h o His io us man/anus man/anus. Adul male om San Es eban,
Co doba P o ince (OMNH 18984). Ba
=
I em. Tee h no o scale.
BARQUEZ ET AL.-THE BATS OF ARGENTINA
141
Table 13. Measu emen s o His io us magellanicus, His io us mon anus, and His io us ela us. Fo a
desc ip ion o he measu emen s, see he Ma e ials and Me hods. Mean, s anda d de ia ion, and sample size a e
ollowed by ange.
Va iable His io us magellanicus His io us mon anus
To al leng h 111.0 ±14.14,2 112.1i541,25
101.0 - 121.0 100.0 - 1220
Tail leng h 46.5 i 9.19, 2 47.3 i 3.67, 25
40.0 - 53.0 40.0 - 55.0
Hind oo leng h 95iO.71,2 10.2 i 1.01,25
9.0 - 10.0 8.0 - 12.0
Ea leng h 22.5 i 2.12,2 25.7 ±241, 25
21.0 - 24.0 20.0 - 29.5
Fo ea m leng h 45.7 i 1.84,2 45.1 ±1.63,25
444 - 47.0 42.5 - 49.0
Weigh 160, I 13.6 i 2.84, 21
9.0 - 18.8
Condylobasal leng h 17.2, 1 16.7 i 0.33, 26
16.2 - 17.3
Leas in e o bi al b ead h 5.25 i 1.20,2 62iO.21,26
44 - 6.1 5.9 - 6.6
Zygoma ic b ead h 11.7, 1 Il.2i041,25
10.5 - 123
G ea es leng h o skull 17.8, 1 17.5 i 0.35, 26
17.0 - 18.2
Pos o bi al cons ic ion 4.5, I 44 i 0.16, 26
4.2 - 4.8
B ead h o b aincase 8.4, I 8.2 i 0.20, 26
7.9 - 8.7
Leng h o maxi lla y oo h ow 645iO.21,2 6.3 i 0.20, 26
6.3 - 6.6 5.9 - 6.8
Pala al leng h 75, I 8.1 0.31,25
7.5 - 88
Mas oidal b ead h 9.5, I 9.liO.15,26
8.8 - 9.3
Leng h o mandibula oo h ow 7.1, I 6.8 i O.
J
8, 25
6.5 - 7.2
Leng h o mandible 13.3
±
0.14, 2 12.1 i 0.30, 26
13.2 - 134 11.5- 12.8
C-C (wid h ac oss canines) 5.2 ±0.21, 2 4.9 i 0.16, 26
5.0 - 5.3 4.6 - 5.3
M-M (wid h ac oss mola s) 6.8 i 0.28, 2 7.1 i 024, 26
6.6 -7.0 66 - 7.6
His io us e a/us
46.0,1
His io us ela us
(I.
Geo oy S .- Hilai e) Ge ais, ed., Mammi e es. In Animaux nou eaux ou
a es ecueillis pendan l' expedi ion dans les pa ies
cen ales de
I'
Ame ique du Sud, de Rio de Janei o a
Lima, e de Lima au Pa a; execu ee pa o d e du
gou e nemen F ancais pendan les annees 1843
a
1847,
sous la di ec ion du com e F ancis de Cas elnau (F.
Cas elnau, ed.). P. Be and, Pa is, 1(2):77, 1855 (1856).
Pleco us ela us I. Geo oy S .- Hilai e, Annales
des Sciences Na u elles, Pa is, 3:446, 1824.
His io us ela us: Ge ais, Deuxie ne Memoi e.
Documen s zoologiques pou se i
a
la monog aphie
des Chei op e es Sud-A ne icains. Pp. 25-88, in P.
142 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
---------1
--
/~'
.
i',....----....-~,
,
'
./,> "
o
I
Figu e 101. Map o he locali ies o His io us ela us.
Type Locali y.- B azil, Pa ana, Cu i iba.
Dis ibu ion (Fig. 10I).- Cu en ly, his species
is es ic ed o Boli ia (Ande son, 1993), cen al and
sou he n B azil, sou heas e n Pa aguay, and no heas -
e n A gen ina (Koopman, 1993), whe e i is only known
om he p o inces o Co ien es and Misiones.
Desc ip ion (Figs. 92d and 99b ).- Cab e a (1958)
ema ked ha he igu es o His io us ela us published
by Ge ais (1855) do no clea ly p esen he cha ac e s
o his species and he sugges ed ha hey may ha e been
based on a specimen o H. mac o us. The e is an e o
in he lack o an in agina ion on he uppe ou e ma gin
o he ea , which is shown as almos con inuous in he
igu e (Ge ais, 1855: pla e 13). In H. ela us, he ea
is poin ed, gi ing a iangula appea ance. This shape
is one o he ew cha ac e s ha can be used o di e en-
ia e H. ela us om H. alienus (in which he ea s a e
ounded). Thomas (1916) ag eed wi h Pe e s (1875) ha
H. ela us is eadily dis inguishable om o he His io us
by he iangula ea , he inne ma gin o which has a
o wa d-p ojec ing ounded lobe; he lobes o each ea
a e connec ed by a ans e se band 3 um in heigh . In
addi ion o ea shape, H. ela us di e s om o he
His io us by he da k colo a ion o bo h he hai and he
memb anes. No mally he ans e se band be ween he
ea s is p esen (abou 3 mm in heigh ) bu , as in he
o he species, his cha ac e may a y.
200
I
400 km
I
The skull is sligh ly na owe han in he o he
species, and he in e o bi al egion and pala e a e na -
owe (Thomas, 1916), bu hese di e ences a e sub le.
J 1easu emen s.- Table 13.
Commen s.- The biology o his species is al-
mos unknown. I is known o be p eyed upon by S ygian
owls (Asio s ygius) in he ce ado sa anna o B azil
(Mo a Junio and Taddei, 1992).
Specimens Examined (3).- MlSlONES:
F on e a, Tobuna, I (MACN); Gua ani, 1 (CEM); San
Ignacio, I (MACN).
Addi ional Reco ds.- CORRlENTES: Vi aso o
(Vacca o, 1992). MISIONES: Campo Ramon (Vacca o,
1992); Rio Vic o ia (Massoia, 1980).
Genus Dasyp e us
Dasyp e us has long been ecognized as a genus
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M 3/3, o al30, whe eas in Lasiu us i is I 1/3, C 1/1, P
2/2, M 3/3, o al 32. In D. ega, he p oximal one-hal o
he u opa agium is u ed. The ea s a e small and
ounded. I has only one uppe p emola on each side.
I lacks he os ing on he pelage ha cha ac e izes al-
mos all species o Lasiu us, and only he p oximal one-
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Sazima, I., and W. Uieda. 1977.
0
mo cego P omops
nasu us no sudes e b asilei o (Chi op e a,
Molossidae). Ciencia e Cul u a, 29:312-314.
Sazima, M., M. E. Fabian, and I. Sazima. 1982.
Polinizacao de Luehea speciosa (Tiliaceae) po
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42:505-513.
Schinz, H. R. 1821. Das hie eich einge heil nach dem
Bau de Thie e als G undlage ih e
Na u geschich e und de e gleichenden
Ana omie on dem He n Ri e on Cu ie .
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0
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Ciencia e Cul u a, 27:723-734.
BARQUEZ ETAL.-THE BATS OF ARGENTINA 261
S 57° 35' W 119. Capi al Fede al (Dis i o Fede al, Buenos Ai es)
92. B azo La go (Islas del Ibicuy, En e Rios) 33° 51' 34° 38' S 58° 34' W
S 58° 57' W 120. Capi an Sola i, La Fo es al (Sa gen o Cab al,
93. Buenos Ai es (Dis i o Fede al, Buenos Ai es) 34° Chaco) 26° 45' S 59° 49' W
38' S 58° 34' W 121. Ca agua ay (Mon eca lo, Misiones) 26° 37' S 54°
94. Bu uyacu (Bu uyacu, Tucu nan) 26° 30' S 64° 37' W
45'W 122. Ca o Quemado (Lo en ue, La Pampa) 36° 28' S
95. Bu uyacu, Iglesia (Bu uyacu, Tucuman) 26° 30' 65°21'W
S 64° 45' W 123. Casa de Pied a, Rio Los Sosa, u a 307, km 24.9,
96. Bu zaco (Almi an e B own, Buenos Ai es) 34° 850 m (Mon e os, Tucu nan) 27° 03' S 65° 37' W
50' S 58° 24' W 124. Caspichango (Mon e os, Tucu nan) 27° 04' S 65°
97. Cacheu a (Lujan, Mendoza) 33° 01' S 69° 08' W 30'W
98. Cachi (Cachi , Sal a) 25° 07' S 66° 12' W 125. Cas ano Nue o, 9 km NW Villa Nue a
99. Cachi, 2500 m (Cachi , Sal a) 25° 07' S 66° 12' W (Calingas a, San Juan) 31° 02' S 69° 33' W
100. Ca aya e (Ca aya e, Sal a) 26° 06' S 65° 57' W 126. Ca ama ca, ciudad (Capi al, Ca ama ca) 28° 28'
101. Ca aya e, 20 km NW, 1700 m (San Ca los, Sal a) S 65° 47' W
25° 54' S 66° 09' W 127. Ca a a as del Iguazu (Iguazu, Misiones) 25° 41'
102. Caimanci o (Ledesma, Jujuy) 23° 44' S 64° 36' W S 54° 27' W
103. Caleu Caleu (Caleu Caleu, La Pampa) 38° 59' S 128. Ca a a as del Iguazu, 125 m (Iguazu, Misiones)
64° 04' W 25° 41' S 54° 27' W
104. Calilegua (Ledesma, Jujuy) 23° 46' S 64° 47' W 129. Cebila , 20 km W Apolina io Sa a ia (An a,
105. Cama Co ada, ce ca de Los Hoyos (Rio Seco, Sal a) 24° 33' S 64° 30' W
Co doba) 29° 52' S 63° 39' W 130. Ce canias de San Ped o (San Ped o, Jujuy) 24°
106. Camino de Comisa, 10 km W Dique La Cienaga 15' S 64° 50' W
(El Ca men, Jujuy) 24° 29' S 65° 22' W 131. Ce illos (Ce illos, Sal a) 24° 55' S 65° 29' W
107. Campana (Campana, Buenos Ai es) 34° 10' S 58° 132. Ce o de la Glo ia (Capi al, Mendoza) 32° 54' S
57' W 68° 50' W
108. Campana, INTA (Campana, Buenos Ai es) 34° 133. Ce o del Campo, 900 m (Bu uyacu, Tucuman)
10' S 58° 57' W 26° 35' S 64° 57' W
109. Campo Aleg e (Juan F. Iba a, San iago del 134. Ce o San Ja ie (Ye ba Buena, Tucuman) 26°
Es e o) 28° 19' S 61° 54' W 47' S 65° 24' W
110. Campo Gallo (Albe di, San iago del Es e o) 26° 135.
Cha ia
Pozo (Rio Hondo, San iago del Es e o)
36' S 62° 50' W 27° 34' S 64° 41' W
Ill.
Campo Quijano (Rosa io de Le ma, Sal a) 24° 136. Chancani (Pocho, Co doba) 31° 25' S 65° 28' W
54' S 65° 38' W 137. Chasco nus (Chasco nus, Buenos Ai es) 35° 34'
112. Campo Ramon (Obe a, Misiones) 27° 26' S 54° S 58° 2' W
58'W 138. Cha a ia (San Roque, Co ien es) 28° 57' S 58°
113. Campo San o (Gene al Giiemes, Sal a) 24° 32' S 34' W
65° 06' W 139. Chileci o (Chileci o, La Rioja) 29° 10' S 67° 30' W
114. Campo San o, Escuela Nacional 149 (Gene al 140. Chimpay (A ellaneda, Rio Neg o) 39° 09' S 66°
Giiemes, Sal a) 24° 32' S 65° 06' W 11'W
115. Campo Vie a (Obe a, Misiones) 27° 20' S 55° 141. Chos Malal (Chos Malal, Neuquen) 37° 23' S 70°
03' W 16'W
116. CanaliS (Cas elli, Buenos Ai es) 35° 58' S 57° 142. Choya, 13 km NNW o Andalgala, 4,000 .
47' W (Andalgala, Ca ama ca) 27° 32' S 66° 24' W
117. Canal 6, Del a bonae ense (Campana, Buenos 143. Chumbicha, 1 km Nand W o balnea io by oad,
Ai es) 34° 7' S 59° 02' W 2,600
.
(Capayan, Ca ama ca) 28° 52' S 66° 14'W
118. Candela ia (Candela ia, Misiones) 27° 28' S 55° 144. Chumbicha, 7 km N (Capayan, Ca ama ca) 28°
45'W 52' S 66° 14' W

262 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
145. Chumbicha, a balnea io, 2,400 . (Capayan, 50'W
Ca ama ca) 28° 52' S 66° 14' W 170. Co ien es Capi al, Ba io Lomas (Capi al,
146. Ciudad Capi al (Capi al, Tucuman) 26° 50' S 65° Co ien es) 27° 28' S 58° 50' W
13'W 171. Co ien es Capi al, Escuela 599 (Capi al,
147. Clo inda (Pilcomayo, Fo mosa) 25° 17' S 57° Co ien es) 27° 28' S 58° 50' W
43'W 172. C uz del Eje (C uz del Eje, Co doba) 30° 44' S
148. Clo inda, 13 km al S, sob e u a II (Pilcomayo, 64° 49' W
Fo mosa) 25° 23' S 57° 47' W 173. Cucho, San Sal ado de Jujuy (D . Manuel
149. Codihue (Picuches, Neuquen) 38° 28' S 70° 30' W Belg ano, Jujuy) 24° 04' S 65° 15' W
150. Cola lao del Valle, 2500 m (Ta i del Valle, 174. Cues a del 25, 3 k n N en ada al Dique EI Cadillal
Tucuman) 26° 22' S 65° 57' W (Ta i Viejo, Tucuman) 26° 37' S 65° 12' W
151. Colas ine Su (La Capi al, San a Fe) 31° 42' S 175. Cues a del Cla illo, 3 km SW La Bande i a
60° 40'W (Andalgala, Ca ama ca) 27° 29' S 66° 06' W
152. Colonia Beni ez (P ime o de Mayo, Chaco) 27° 176. Cues a del Cla illo, 5 km S La Bande i a
20' S 58° 57' W (Andalgala, Ca ama ca) 27° 29' S 66° 06' W
153. Colonia Beni ez, 20 km N Resis encia, 30 m 177. Cues a del To o al (Paclin, Ca ama ca) 28° 0
:
S
(P ime o de Mayo, Chaco) 27° 20' S 58° 57' W 65° 35' W
154. Colonia Do a (A ellaneda, San iago del Es e o) 178. Cues a La Sebila, Hwy 60 (Capayan, Ca ama ca)
28° 37' S 62° 57' W 28° 43' S 66° 22' W
155. Colonia EI Pa ao, 41 k n N Comandan e Fon ana 179. Cue a del Chacho, Pa quia (Independencia, La
on Rd 95 (Pa ino, Fo mosa) 24° 55' S 59° 36' W Rioja) 30° 03' S 66° 53' W
156. Colonia k n 503 (Pa ino, Fo mosa) 25° 23' S 60° 180. Cue a del Indio, 4 km W Ca aya e, 1800 m
19'W (Ca aya e, Sal a) 26° OS'S 65° 58' W
157. Colonia k n 503, 16 km S (Pa ino, Fo mosa) 25° 181. Cue a del Mu cielagallo, 15 km SSW San a
26' S 60° 15' W Vic o ia, 2000 m (San a Vic o ia, Sal a) 22° IS'S
158. Colonia Ma i es (Candela ia, Misiones) 27° 26' 65° 02' W
S 55° 23' W 182. Cue a del Tig e, 74 km N Pampa Blanca, 700 m
159. Colonia Ma i es, Chac a Fe ei a, 125 m (Ledesma, Jujuy) 23° 57' S 64° 47' W
(Candela ia, Misiones) 27° 26' S 55° 23' W 183. Cue a del Tig e, El Milag o (EI Ca men, Jujuy)
160. Colonia Ma i es, Pue o Candela ia (Candela ia, 24° 22' S 65° 02' W
Misiones) 27° 26' S 55° 23' W 184. Cue a Ma ia An onia (San Ignacio, Misiones) 27°
161. Comandan e Fon ana (Pa ino, Fo mosa) 25° 19' 17' S 55° 32' W
S 59° 30' W 185. Cu a B oche o (San Albe o, Co doba) 31°40' S
162. Complejo As ono nico EI Leonci o, 7 k n W 65° 00' W
(Ca1ingas a, San Juan) 31° 48' S 69° 20' W 186. Desaguade o, Escuela 208, Ru a P o incialS, a
163. Concepcion (Chicligas a, Tucuman) 27° 20' S 65° 20 k n de Co ien es (Capi al, Co ien es) 27° 34'
35'W S 58° 39' W
164. Conco dia, 32 k n N (Fede acion, En e Rios) 31° 187. Dique EI Cadillal, 25 k n N San Miguel de Tucuman
09' S 57° 58' W (Ta i Viejo, Tucu nan) 26° 37' S 65° 12' W
165. Co azon de Ma ia (Colon, Co doba) 31° 30' S 188. Dique EI Po e o (Andalgala, Ca ama ca) 27° 32'
64° OO'W S 66° 20' W
166. Co doba (Capi al, Co doba) 31° 25' S 64° 12' W 189. Dique EI Po e o, 13 km N Andalgala (Andalgala,
167. Co onel 1. F. Gomez (Gene al Roca, Rio Neg o) Ca ama ca) 27° 32' S 66° 20' W
39° 03' S 67° 37' W 190. Dique Escaba (Albe di, Tucuman) 27° 40' S 65°
168. Co onel Vidal (Ma Chiqui a, Buenos Ai es) 37° 46'W
48' S 57° 44' W 191. Dique I iyu o (San Ma in, Sal a) 22° 10' S 63°
168a. Co al de Pied as (Coli on Cu a, Neuquen) 40° 50'W
26' S 70° 36' W 192. Dique I iyu o, 5 k n W (San Ma in, Sal a) 22°
169. Co ien es (Capi al, Co ien es) 27° 28' S 58° 10' S 63° 55' W
BARQUEZ ET AL.-THE BATS OF ARGENTINA 263
193. Dique La Cienaga (EI Ca men, Jujuy) 24° 25' S 220. EI Colmena (Capi al, Tucuman) 26° 50' S 65°
65° 17' W 13'W
194. Dique La Cienaga, 13
Ian
SSW S. S. de Jujuy 221. EI Colo ado (Pi ane, Fo mosa) 26° 18' S 59° 22' W
(EI Ca men, Jujuy) 24° 25' S 65° 17' W 222. EI C6ndo , Lago Pe i o Mo eno, 800
ill
195. Dique La Cienaga, casa del club nau ico, 30
Ian
(Ba iloche, Rio Neg o) 41° 08' S 71° 26' W
SSW San Sal ado de Jujuy, 1000 m (EI Ca men, 223. EI Co e (Ye ba Buena, Tucu nan) 26° 49' S 65°
Jujuy) 24° 25' S 65° IT W 26'W
196. Dique Las Made as (EI Ca men, Jujuy) 24°
21'
S 224. El Du aznillo, 3000 m, Ce o Calilegua (Valle
65° 15' W G ande, Jujuy) 23° 28' S 64° 55' W
197. Dique Las Made as, 42
Ian
San Sal ado de Jujuy, 225. EI Hoyo (Cusha nen, Chubu ) 42° 03' S 71° 31' W
920 m (EI Ca men, Jujuy) 24° 27' S 65° 15' W 226. El Mai en, 2 km N (Cusha nen, Chubu ) 42° 02'
198. Dique San Ignacio (La Cocha, Tucuman) 27° 44' S 71° 09' W
S 65° 40' W 227. El Manan ial (Lules, Tucu nan) 26° 51' S 65°
199. Dola on (Gaiman, Chubu ) 43° 20' S 65° 43' W
IT
W
200. Dos de Mayo (Cainguas, Misiones) 27° 02' S 54° 228. EI Mang ullo, 20
Ian
NNW by oad and II
Ian
39'W NE by oad (Almi an e B own, Chaco) 26° 09' S
201. Dos de Mayo, Pa aje
Ian
51 (Cainguas, Misiones) 61° 15' W
27° 02' S
54°
39' W 229. EI Moj6n (Pelleg ini, San iago del Es e o) 26°
202. D agones (San Ma in, Sal a) 23° 16' S 63° 21' W 09' S 64° 18' W
203. El Ba eal (Capi al, La Rioja) 29° 38' S 66° 02' W 230. EI Monoli o (Valle G ande, Jujuy) 23° 40' S 64°
204. EI Bo1s6n (Ba iloche, Rio Neg o) 41° 58' S 71° 54'W
31'W 231. EI Na anjal (Mon e os, Tucu nan) 27° 02' S 65°
205. EI Bols6n, 3
Ian
N (Ba iloche, Rio Neg o) 41 ° 41'W
58'S7I031'W 232. EI Na anjo (Bu uyacu, Tucuman) 26° 40' S 65°
206. EI B eal (Ri ada ia, Sal a) 22° 14' S 62° 56' W 03'W
207. EI B eal, 6 km SW San a Vic o ia Es e 233. EI Nogala , Ru a 307, 1700
ill
(Mon e os,
(Ri ada ia, Sal a) 22° 14' S 62° 56' W Tucuman) 27° 0 I' S 65° 40' W
208. EI Cadillal (Ta i Viejo, Tucu nan) 26° 37'S 65° 234. EI Palma del Rio San F ancisco (San a Ba ba a,
12'W Jujuy) 24° 08' S 64° 33' W
209. EI Cadillal, 25
Ian
N San Miguel de Tucuman 235. EI Pa aiso (Ye ba Buena, Tucuman) 26° 49' S 65°
(Ta i Viejo, Tucuman) 26°
31'
S 65° 12' W 26'W
210. EI Cadillal, Es aci6n de Piscicul u a (Ta i Viejo, 236. EI Po eni (Me an, Sal a) 25° 23' S 64° 55' W
Tucuman) 26° 37'S 65° 12' W 237. EI Po e o (Rosa io de La F on e a, Sal a) 25°
211. EI Cadillal, Rio Lo o (Bu uyacu, Tucuman) 26° 53' S 64° 40' W
37' S 65° 10' W 238. E1 Po e o, 13 km N Andalgala (Andalgala,
212. EI Cadillal, usina (Ta i Viejo, Tucuman) 26°
31'
Ca ama ca) 27° 33' S 66° 20' W
S65° 12'W 239. EI Re i o (Jimenez, San iago del Es e o) 27° 04'
213. EI Ca men (EI Ca men, Jujuy) 24° 24' S 65° 15' W S 64° 28' W
214. EI Ca izal, C6 doba Range, 1000 m (San Ja ie , 240. EI Rinc6n (Jimenez, San iago del Es e o) 26° 43'
C6 doba) 31°
51'
S 65° 12' W S 64° 40' W
215. EI Ca izal, Villa Dolo es, 1000 m (San Ja ie , 241. EI Rodeo, 900 m (Amba o, Ca ama ca) 28° 13' S
C6 doba) 31° 57' S 65° 12' W 65° 52' W
216. EI Chagua al (O an, Sal a) 24° 00' S 64° 00' W 242. EI Rodeo-La Jun a (Amba o, Ca ama ca) 28° 13'
217. EI Challao (Las He as, Mendoza) 32° 50' S 68° S 65° 52' W
52'W 243. El Simbola , 25 km SW Palma Sola (San a
218. EI Chu cal, app ox. 10 km SSE Buena Vis a Ba ba a, Jujuy) 24° II'S 64° 28' W
(Be mejo, Fo mosa) 23° 30' S 61° 33' W 244. EI Tuscal (Tulumba, C6 doba) 29° 40' S 64° 31' W
219. EI Cogoik, 5
Ian
an es de (Pa ino, Fo mosa) 24° 245. EI Zapalla , Finca Las T es Ma ias (Gene al
48' S 59° 10' W Gue nes, Sal a) 24° 31' S 65° 00' W
264 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
246. Eldo ado (Eldo ado, Misiones) 26° 26' S 54° 41' W 270. Es anislao del Campo (Pa ino, Fo mosa) 25° 03'
247. Eldo ado, 3 km SE (Eldo ado, Misiones) 26° 26' S 60° 06' W
S54°41'W 271. Es anislao del Campo, 40 km N (Pa ino,
248. Emba cacion, Finca T es Pozos, 247 km NE San Fo mosa) 24° 43' S 60° 06' W
Sal ado de Jujuy (San Ma in, Sal a) 23° 13' S 272. Ezpele a (Quilmes, Buenos Ai es) 34° 45' S 58°
64° 07' W 14'W
249. Ene gia (San Caye ano, Buenos Ai es) 38° 34' S 273. Finca Ab a G ande (San Ma in, Sal a) 22° 40' S
59° 21' W 63° 41' W
250. Escuela 149,30 km NE Sal a (Gene al Gue nes, 274. Finca Belg ano, Ce illos, 30 k n SW Sal a
Sal a) 24° 32' S 65° 06' W (Ce illos, Sal a) 24° 55' S 65° 29' W
251. Escuela Gu uchaga, Rosa io de Le ma (Rosa io 275. Finca Ca amon a ia, 33 k n SSE San Sal ado
de Le ma, Sal a) 24° 59' S 65° 35' W de Jujuy, 925 m (El Ca men, Jujuy) 24° 23' S 65°
252. Escuela P o incial 639, Campo Ramon (Obe a, 07' W
Misiones) 27° 26' S 54° 58' W 276. Finca El A azayal, Se ania de las Pa as (O an,
253. Espe anza (Las Colonias, San a Fe) 31°27' S 60° Sal a) 22° 33' S 64° 32' W
56'W 277. Finca El Ol ido, 15 km E Palma Sola (San a
254. Espinillo (Rio Cua o, Co doba) 33° 01' S 64° Ba ba a, Jujuy) 24° 00' S 64° 10' W
21'W 278. Finca El Pongo, 45 km E San Sal ado de Jujuy
255. Es acion Banos (Rosa io de La F on e a, Sal a) (El Ca men, Jujuy) 24° 21' S 65° 04' W
25° 50' S 64° 50' W 279. Finca EI Rema e, 24 km SE San Sal ado de
256. Es acion Medanos (Gualeguaychu, En e Rios) Jujuy, 740 m (D . Manuel Belg ano, Jujuy) 24°
33° 25' S 59° 41' W 17' S 65° 06' W
257. Es acion Pa anaci o (Islas del Ibicuy, En e Rios) 280. Finca La Ca olina, Los Pe ales, San Sal ado de
33° 42' S 59° 01' W Jujuy, 1310 m (D . Manuel Belg ano, Jujuy) 24°
258. Es acion Pa quia, Guayapa (lndependencia, La 11' S 65° 19' W
Rioja) 30° 09' S 66° 57' W 281. Finca La C uz, 28 km SSE Sal a (Capi al, Sal a)
259. Es ancia Alicu a (Laca , Neuquen) 40° 30' S 70° 24° 54' S 65° 10' W
45'W 282. Finca La C uz (Capi al, Sal a) 24° 54' S 65° 10' W
260. Es ancia Al o Las Jun as (Amba o, Ca ama ca) 283. Finca La Flo ida, Rosa io de Le ma, 25 km SSW
28° 08' S 65° 54' W Sal a (Chicoana, Sal a) 25° 0 I' S 65° 34' W
261. Es ancia EI Casali o, Ru a 226 ca. km 98 284. Finca La Rosa, Ca aya e, 2000 m (Ca aya e,
(Balca ce, Buenos Ai es) 37° 46' S 58° 00' W Sal a) 26° 06' S 65° 57' W
262. Es ancia EI Ca ao (Bu uyacu, Tucuman) 26° 30' 285. Finca La To na, 25 km SW San Sal ado de Jujuy
S 64° 45' W (San An onio, Jujuy) 24° 20' S 65° 21' W
263. Es ancia La Espe anza (Gene al La alle, Buenos 286. Finca San Lo enzo, Rosa io de La F on e a
Ai es) 36° 30' S 57° 13' W (Rosa io de La F on e a, Sal a) 25° 48' S 64° 58' W
264. Es ancia Las Jun as (Amba o, Ca ama ca) 28° 08' 287. Finca San a C uz, 3 km N Las Me cedes (Rosa io
S 65° 54' W de La F on e a, Sal a) 25° 57' S 64° 40' W
265. Es ancia Las Me cedes (Pilco nayo, Fo mosa) 25° 288. Finca Tabaquillo, 15 km E C uz del Eje (C uz
II'S 57° 54' W del Eje, Co doba) 30° 45' S 64° 52' W
266. Es ancia Pues o Valle, 25 km E I uza ingo 289. Flo encia (Gene al ObJigado, San a Fe) 28° 02' S
(l uzaingo, Co ien es) 27° 36' S 56° 25' W 59° 14' W
267. Es ancia San Miguel, along Hwy 90, 15 km NW 290. Flo es a (Dis i o Fede al, Buenos Ai es) 34° 38'
jc . Hwy 90 and Hwy II (P ime o de Mayo, S 58° 28' W
Chaco) 26° 57' S 58° 59' W 291. F acan (Gua ani, Misiones) 26° 47' S 54° 15' W
268. Es ancia San a Ca alina, 5 km S, km 64 (Pa ino, 292. F aile Pin ado, 12 km W, sob e Rio Ledesma
Fo mosa) 24° 56' S 59° 12' W (Ledesma, Jujuy) 23° 57' S 64° 55' W
269. Es ancia Via mon e (Rio G ande, Tie a del 293. F on e a, Tobuna (San Ped o, Misiones) 26° 26'
Fuego) 53° 59' S 67° 30' W S 53° 54' W
BARQUEZ ET AL.-THE BATS OF ARGENTINA 265
294. Gaspa Campos, 10 km N (Gene al Al ea , (Ye ba Buena, Tucu nan) 26°
41'
S 65° 23' W
Mendoza) 34° 39' S 67° 26' W 321. Ho co Molle, Residencia Uni e si a ia (Ye ba
295. Gene al Giie nes, Ru a 34 km 1363 (Gene al Buena, Tucuman) 26°
41'
S 65° 23' W
Giiemes, Sal a) 24° 31' S 65° 02' W 322. Ho co Molle, Rio Las Canas (Ye ba Buena,
296. Gene al La alle, 20 mi S (Gene al La alle, Tucu nan) 26°
41'
S 65° 23' W
Buenos Ai es) 36° 42' S 56°
51'
W 323. Ho co Molle, Rio Las Le ias (Ye ba Buena,
297. Gene al M. M. de Gue nes, 30 km SSW (Gen- Tucu nan) 26°
41'
S 65° 23' W
e al Gue nes, Sal a) 24° 40' S 65° 15' W 324. Ho co Molle, Rio Las Pied as (Ye ba Buena,
298. Gene al Pico (Ma a
C6,
La Pampa) 35° 40' S 63° Tucuman) 26°
41'
S 65° 23' W
46'W 325. Ho cones (Rosa io de La F on e a, Sal a) 25° 48'
299. Gene al Roca (Gene al Roca, Rio Neg o) 39° 02' S 64° 55' W
S 67° 33' W 326. Ho el Llao Llao (Ba iloche, Rio Neg o) 41 ° 0 I'
300. Gene al Vedia (Be mejo, Chaco) 26° 56' S 58° S71031'W
41'W 327. Ho el Te mas, Rosa io de La F on e a (Rosa io
30 I. Gi a de (Mo eno, San iago del Es e o) 27° 38' S de La F on e a, Sal a) 25° 50' S 64° 56' W
62° II' W 328. Hoyo de Epuyen (Cusha nen, Chubu ) 42° 03' S
302. Gobe nado Lanusse (Iguazu, Misiones) 25° 58' 71°31'W
S 54°
IT
W 329. Hu nai a (Capi al, Sal a) 24° 55' S 65° 23' W
303. Gobe nado Pied abuena (Bu uyacu, Tucu nan) 330. Huyamampa (La Banda, San iago del Es e o) 27°
26° 45' S 64° 40' W 23' S 64° 18' W
304. Godoy C uz (Godoy C uz, Mendoza) 32° 55' S 331. Icaiio (A ellaneda, San iago del Es e o) 28° 40'
68° 52' W S 62° 53' W
305. Goya (Goya, Co ien es) 29° 08' S 59° IT W 332. Iglesia (Bu uyacu, Tucu nan) 26° 30' S 64° 45' W
306. Goya, 600 m (Goya, Co ien es) 29° 08' S 59° 333. Iguazu (Iguazu, Misiones) 25° 35' S 54° 33' W
IT
W334. Iguazu, ce ca de las Ca a a as del Iguazu (Iguazu,
307. G anja 17 de Ab il (Ma cos Paz, Buenos Ai es) Misiones) 25° 41' S 54°
21'
W
34° 45' S 58° 58' W 335. Ingenio Amalia (Capi al, Tucuman) 26° 52' S 65°
308. Guachipas (Guachipas, Sal a) 25° 31' S 65° 31' W 13'W
309. Guachipas, Iglesia (Guachipas, Sal a) 25° 31' S 336. Ingenio Lules (Lules, Tucuman) 26° 55' S 65°
65° 31' W 20' W
310. Gualeguaychu (Gualeguaychu, En e Rios) 33° 337. Ingenio San Ma in del Tabacal (O an, Sal a) 23°
OI'S 58° 31' W 15' S 64° 15' W
311. Gua ani (Obe a, Misiones) 27°
31'
S 55° 10' W 338. Ingenio San Pablo (Lules, Tucuman) 26° 54' S
312. Gue nes (Gene al Gue nes, Sal a) 24° 31' S 65° 65° 19' W
02'W 339. Ingenio San a Lucia (Mon e os, Tucuman) 27°
313. He e a (A ellaneda, San iago del Es e o) 28° 28' 06' S 65° 31' W
S 63° 04' W 340. Ins i u o Lillo (Capi al, Tucuman) 26° 49' S 65°
314. Hickman (San Ma in, Sal a) 23° 13' S 63° 34' W 13'W
315. Hickman, Luna Mue a (San Ma in, Sal a) 23° 341. Ins i u o Lillo, San Miguel de Tucu nan (Capi-
13' S 63° 34' W al, Tucu nan) 26° 49' S 65° 13' W
316. Ho co Molle (Ye ba Buena, Tucuman) 26° 4
T
S342. Ischigualas o (Valle Fe il, San Juan) 30° 06' S
65° 23' W 67° 55' W
317. Ho co Molle, 15 km W San Miguel de Tucu nan 343. Isla Apipe (I uzaing6, Co ien es) 27° 30' S 56°
(Ye ba Buena, Tucu nan) 26°
41'
S 65° 23' W 53' W
318. Ho co Molle, 900 m (Ye ba Buena, Tucuman) 344. Isla Apipe G ande (I uzaing6, Co ien es) 27° 30'
26° 41'
S 65° 23' W S 56° 53' W
319. Ho co Molle, Biological Rese e (Ye ba Buena, 345. Isla del Ce i o (Be mejo, Chaco) 27° 16' S 58°
Tucuman) 26°
41'
S 65° 23' W 41'W
320. Ho co Molle, Pa que Biol6gico Sie a de San Ja ie 346. Isla Vic o ia (Los Lagos, Neuquen) 41° 00' S 71°
266 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
30'W 372. La Higue a (T ancas, Tucu nan) 26° 23' S 65°
347. Isla Vic o ia, Pied as Blancas (Los Lagos, 26' W
Neuquen) 40° 50' S 71
o
30' W 373. La Jun a, 1600 m (Amba o, Ca ama ca) 28° 08' S
348. Islas del Ibicuy, Pa anaci o (Islas del Ibicuy, En- 65° 54' W
e Rios) 33° 44' S 58° 40' W 374. La Maya, 45
km
SE Bell Ville (Ma cos Jua ez,
349. Islas del Ibicuy (Islas del Ibicuy, En e Rios) 33° C6 doba) 33° 00' S 62° 28' W
44' S 59° 10' W 375. La Me ced (Paclin, Ca ama ca) 28° 10' S 65°
350. I a Iba e (Gene al Paz, Co ien es) 27° 26' S 57° 41' W
20'W 376. La Me ced (Ce illos, Sal a) 24° 58' S 65° 29' W
351. I a i (Ha i, Co ien es) 27° 16' S 58° 15' W 377. La Me ced, 1200 m (Ce illos, Sal a) 24° 58' S
352. I au (Campo La go), Sie as de Ta agal, 1600 m 65° 29' W
(San Ma in, Sal a) 22° 05' S 63° 55' W 378. La Paz (San Ja ie , C6 doba) 32° 15' S 65° 0 I' W
353. I uzaing6 (I uzaing6, Co ien es) 27° 35' S 56° 379. La Pega (Guay nallen, Mendoza) 32° 49' S 68°
41'W 40'W
354. I uzaing6, 25
km
E (I uzaing6, Co ien es) 27° 380. La Pla a (La Pla a, Buenos Ai es) 34° 55' S 57°
36' S 56° 25' W 57' W
355. Je . Hwy 2 and A oyo Pa aiso (Gua ani, 381. La Poblaci6n (C uz del Eje, C6 doba) 30° 50' S
Misiones) 27° II'S 54° 03' W 65
0
21'W
356. Je . Hwy 21 and A oyo O eja Neg a, app ox. 2 382. La Pochi a, Ba i ina (Sob emon e, C6 doba) 29°
km W Pa que P o incial Mocona (Gua ani, 46' S 64° 49' W
Misiones) 27° 08' S 53° 54' W 383. La Pue a (Amba o, Ca ama ca) 28° 10' S 65°
357. Joaquin V. Gonzalez, 8
km
SE, Finca San Ja ie 48'W
(An a, Sal a) 25° 02' S 64° 05' W 384. La Pue a (Rio Hondo, San iago del Es e o) 27°
358. Jujuy ciudad (D . Manuel Belg ano, Jujuy) 24° 38' S 64° 35' W
12' S 65° 19' W 385. La Rinconada (Ye ba Buena, Tucu nan) 26° 49'
359. Jun a de los Rios Ta ija y Be mejo (O an, Sal a) S 65° 20' W
22° 51' S 64° 20' W 386. La Rioja (Capi al, La Rioja) 29° 25' S 66° 51' W
360. Jun as de San An onio (O an, Sal a) 22° 51' S 64° 387. La Toma (Co onel P ingles, San Luis) 33° 04' S
20'W 65° 38' W
361. La A gen ina (Minas, C6 doba) 31
o
14' S 65° 388. La Vi ia (La Vina, Sal a) 25° 28' S 65° 35' W
18'W 389. La Vi ia, iglesia (La Vina, Sal a) 25° 28' S 65°
362. La Banda (La Banda, San iago del Es e o) 27° 35' W
42' S 64° 14' W 390. La e e e (La Ma anza, Buenos Ai es) 34° 45' S
363. La Bande i a (Andalgala, Ca ama ca) 27° 19' S 58° 35' W
65° 58' W 391. Lago Fu alau quen (Fu aleu u, Chubu ) 42° 53'
364. La Bande i a, Ru a 62 camino a Las Es ancias S71°37'W
(Andalgala, Ca ama ca) 27° 19' S 65° 58' W 392. Lago Pe i o Mo eno (Ba iloche, Rio Neg o) 41 °
365. La Bismu ina (Minas, C6 doba) 31 ° 09' S 65° 08' S 71° 26' W
13'W 393. Laguna Blanca (Pilcomayo, Fo mosa) 25° 08' S
366. La Calde a (La Calde a, Sal a) 24° 37' S 65° 23' W 58° 15' W
367. La Cienaga, 2500 m (Ta i del Valle, Tucu nan) 394. Laguna B a a (Capi al, Co ien es) 27° 30' S 58°
26° 46' S 65° 39' W 43'W
368. La Cumb e (Punilla, C6 doba) 30° 59' S 64° 30' W 395. Laguna Chasco nus (Chasco nus, Buenos Ai es)
369. La F agua (Pelleg ini, San iago del Es e o) 26° 35° 34' S 58° 02' W
08' S 64° 20' W 396. Laguna Chasic6 (Villa ino, Buenos Ai es) 38° 38'
370. La Galla e a, Ce i o,
km
60 (Ve a, San a Fe) 29° S 63° 06' W
30' S 60° 30' W 397. Laguna Colo ada G ande, Caleu Caleu (Caleu
371. La Higue a (C uz del Eje, C6 doba) 31
o
0 I' S 65° Caleu, La Pampa) 38° 20' S 63° 45' W
06'W 398. Laguna de las Ca as (San Ma in, Sal a) 22° 16' S

BARQUEZ ET AL.-THE BATS OF ARGENTINA 267
63° 52' W Tucu nan) 26° 39' S 65° 21' W
399. Laguna de los Panzones, Re i o (Ri ada ia, Sal a) 425. Las Tumanas, along Hwy 510 a c ossing o Rio
22° 54' S 63° 10' W Tumanas (Valle Fe il, San Juan) 30° 52' S 67°
400. Laguna Gala za (San o Tome, Co ien es) 28° 06' 20'W
S 56° 40' W 426. Las Tumanas, Rio Tumanas (Valle Fe il, San
401. Laguna G ande (Co pen Aike, San a C uz) 49° Juan) 30° 52' S 67° 20' W
34' S 70° 14' W 427. La alle (Gene al La alle, Buenos Ai es) 36° 30'
402. Laguna La B ea (San a Ba ba a, Jujuy) 23° 56' S S 57° 13' W
64° 28' W 428. La alle (La alle, Mendoza) 32° 43' S 68° 36' W
403. Laguna La B ea, 25 km W Palma Sola (San a 429. La alle, Es ancia El Sal ado (Guasayan,
Ba ba a, Jujuy) 23° 56' S 64° 28' W San iago del Es e o) 28° 12' S 65° 01' W
404. Laguna Mi anda, Zona Zapala (Zapala, Neuquen) 430. Leand o N. Alem (Leand o N. Alem, Misiones)
38° 54' S 70° 16' W 27° 36' S 55° 19' W
405. Laguna Pai a, BoLas Lomas (Capi al, Co ien es) 431. Ledesma (Ledesma, Jujuy) 23° 50' S 64° 41' W
27° 30' S 58° 45' W 432. Leleque (Cushamen, Chubu ) 42° 24' S 71° 04' W
406. Lanus (Lanus, Buenos Ai es) 34° 42' S 58° 23' W 433. Leocadio Paz (T ancas, Tucuman) 26° 08' S 65°
407. Las Chac as (Pocho, Co doba) 31°25' S 65° 00' W 18' W
408. Las Colo adas,
1000
m (Ca an Lil, Neuquen) 39° 434. Libe ad (Iguazu, Misiones) 25° 55' S 54° 36' W
34' S 70° 36' W 435. Libe ado Gene al San Ma in (Ledesma, Jujuy)
409. Las Es ancias (Andalgala, Ca ama ca) 27° 28' S 23° 49' S 64° 41' W
66° 02' W 436. Lib es del Su (Chasco nus, Buenos Ai es) 35°
410. Las Es ancias (Punilla, Co doba) 30° 58' S 64° 44' S 57° 43' W
40'W 437. Lobe ia, 15 km SW (Lobe ia, Buenos Ai es) 38
411. Las Jun as, 22 km W Cho omo o (T ancas, 12' S 58 44' W
Tucuman) 26° 24' S 65° 31' W 438. Lobos (Lobos, Buenos Ai es) 35° 12' S 59° 06' W
412. Las Jun as, 22 km W Cho omo o on Hwy 312, 439. Longchamps (Almi an e B own, Buenos Ai es)
3,500 . (T ancas, Tucuman) 26° 24' S 65° 31' W 34° 52' S 58° 24' W
413. Las Jun as, 50 km NW Ca ama ca Ciudad 440. Los Cisnes (Jua ez Celman, Co doba) 33° 24' S
(Amba o, Ca ama ca) 28° 08' S 65° 54' W 63° 28' W
414. Las Jun as, Es ancia de los Figue oa, 50 km NW 441. Los Helechos, Escuela n° 12 (Obe a, Misiones)
Ca ama ca Ciudad (Amba o, Ca ama ca) 28° 08' 27° 33' S 55° 03' W
S 65° 54' W 442. Los Hoyos (Sob emon e, Co doba) 29° 48' S 64°
415. Las Laji as (An a, Sal a) 24° 41' S 64° 15' W 48'W
416. Las Lomi as (Pa ino, Fo mosa) 24° 43' S 60° 36' W 443. Los Mad ejones, Jun a de los Rios Ta ija e I au,
417. Las Ma ias (San o Tome, Co ien es) 28° 0 I' S 1400 m (San Ma in, Sal a) 22° 18' S 64° 09' W
56° 01' W 444. Los Molles (C uz del Eje, Co doba) 30° 54' S 65°
418. Las Me cedes (Pilcomayo, Fo mosa) 25° 11' S 15'W
57° 54' W 445. Los Nacimien os, u a 40, km 869 (Belen,
419. Las Mesadas (T ancas, Tucu nan) 26° 21' S 65° Ca ama ca) 27° 01' S 66° 42' W
30'W 446. Los Pa lamen os (San Ra ael, Mendoza) 35° 02'
420. Las Pa as (Chicligas a, Tucu nan) 27° 15' S 65° S 69° 20' W
52'W 447. Los Pe ales, San Sal ado de Jujuy (D . Manuel
421. Las Pa as, Aconquija (Chicligas a, Tucuman) 27° Belg ano, Jujuy) 24° II'S 65° 19' W
15' S 65° 52' W 448. Los Sa mien os (Rio Chico, Tucuman) 27° 25' S
422. Las Talas, 4 km al N de Bella Vis a (Leales, 65° 42' W
Tucu nan) 27° 00' S 65° IT W 449. Los Vasquez (Capi al, Tucuman) 26° 50' S 65°
423. Las Te mas (Rio Hondo, San iago del Es e o) 27° 13'W
30' S 64° 50' W 450. Los Vasquez, 445 m (Capi al, Tucuman) 26° 50'
424. Las Ti pas , Pa que Bi olog ico (Ta i Viejo, S 65° 13' W
268 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
45l. Los Yngleses, Ajo (Gene al La alle, Buenos 478. Mo eno (Mo eno, Buenos Ai es) 34° 39' S 58°
Ai es) 36° 31' S 56° 53' W 47' W
452. Luan To o, 25 kIn S (Toay, La Pampa) 36° 20' S 479. Mu iecas (Capi al, Tucuman) 26° 47' S 65° 15' W
65° 07' W 480. Nacuiian (San a Rosa, Mendoza) 34° 03' S 67°
453. Lujan, Quines (Ayacucho, San Luis) 32° 22' S 58'W
65° 56' W 481. Naschel (Chacabuco, San Luis) 32° 56' S 65°
454. Lumb e a (Me an, Sal a) 25° 10' S 64° 56' W 28' W
455. Maipu (Maipu, Buenos Ai es) 36° 52' S 57° 54' W 482. Neuquen (Con luencia, Neuquen) 38° 58' S 68°
456. Malab igo (Gene al Obligado, San a Fe) 29° 20' 03'W
S 59° 59' W 483. Neuquen, 400 m (Con luencia, Neuquen) 38° 58'
457. Mala gue (Mala gue, Mendoza) 35° 28' S 69° S 68° 03' W
35' W 484. Nino Dios, mas
0
menos 1900 m (Minas,
458. Manan iales (Mbu ucuya, Co ien es) 27° 56' S Co doba) 31° 05' S 65° 18' W
58° 06' W 485. Noe inge , E Co doba, 100 m (Union, Co doba)
459. Manuel Elo di, Ve mejo, 500 m (O an, Sal a) 23° 32° 22' S 62° 19' W
16' S 64° 09' W 486. No co, 1200 m (T ancas, Tucuman) 26° 29' S 65°
460. Ma A gen ino (40° 04.5' S 56° 21.5' W) 22'W
46l. Ma del Pla a (Gene al Puey edon, Buenos 487. No co, Vipos, 2500 m (T ancas, Tucuman) 26°
Ai es) 38° 00' S 57° 34' W 29' S 65° 22' W
462. Ma del Pla a, Campo de los Pad es (Gene al 488. Nue a Espe anza (Pelleg ini, San iago del Es e o)
Puey edon, Buenos Ai es) 38° 00' S 57° 34' W 26° 12' S 64° 16' W
463. Ma cos Paz (Ye ba Buena, Tucuman) 26° 49' S 489. Obe a (Obe a, Misiones) 27° 29' S 55° 07' W
65° 17' W 490. Ojo de Agua (Minas, Co doba) 31° 14' S 65° 12' W
464. Ma inez (San Isid o, Buenos Ai es) 34° 30' S 58° 491. Ojo de Agua (Ojo de Agua, San iago del Es e o)
30'W 29° 28' S 63° 39' W
465. Mbu ucuya (Mbu ucuya, Co ien es) 28° 03' S 492. Olega io V. And ade (Candela ia, Misiones) 27°
58° 13' W 34' S 55° 30' W
466. Mendoza (Capi al, Mendoza) 32° 54' S 68° 50' W 493. On Highway 9 a bo de wi h Sal a, a camp-
467. Me cedes (Me cedes, Co ien es) 29° II'S 58° g ound on he way o EI Ca men (EI Ca men,
04'W Jujuy) 24° 28' S 65° 21' W
468. Me an (Me an, Sal a) 25° 30' S 64° 58' W 494. O an (O an, Sal a) 23° 08' S 64° 20' W
469. Me an, 850 m (Me an, Sal a) 25° 30' S 64° 58' W 495. O an, 15 kIn S, 15 k n W along Rio San a Ma ia
470. Mina Pablo, Unchi ne, 25 Ian E Gene al Giie nes, (O an, Sal a) 23° 19' S 64° 14' W
1100 m (Gene al Gue nes, Sal a) 24° 40' S 64° 496. O an, 20 kIn al NW (O an, Sal a) 23° 00' S 64°
55'W 22'W
471. Mina San Ignacio (San Albe o, Co doba) 31° 58' 497. O an, 20 k n al NW de Agua Blanca (O an, Sal a)
S 65° 33' W 22° 33' S 64° 32' W
472. Mision Nue a Pompeya (Gene al Gue nes, 498. Palma Sola (San a Ba ba a, Jujuy) 24° 00' S 64°
Chaco) 24° 50' S 61° 30' W 19'W
473. Molleci o, 30 km SE Anda lga la (Po nan, 499. Palma Sola, 550 m (San a Ba ba a, Jujuy) 24°
Ca ama ca) 27° 46' S 66° 12' W 00' S 64° 19' W
474. Mon e Bello (Rio Chico, Tucuman) 27° 22' S 65° 500. Palma Sola, Mina de Azu e (San a Ba ba a,
45'W Jujuy) 24° 00' S 64° 19' W
475. Mon e Bello, ma gen de echa del Rio Medina 501. Pampa Llaj a (A amisqui, San iago del Es e o)
(Rio Chico, Tucu nan) 27° 22' S 65° 45' W 28° 30' S 63° 50' W
476. Mon eagudo (Simoca, Tucuman) 27° 31' S 65° 502. Pa ada Leis (Capi al, Misiones) 27° 36' S 55°
17'W 50'W
477. Mon eca lo (Mon eca lo, Misiones) 26° 33' S 54° 503. Pa aje Pa aiso, 16 Ian SE, sob e u a 21 (Gua ani,
47' W Misiones) 26° 58' S 54° 10' W
BARQUEZ ET AL.-THE BATS OF ARGENTINA 269
504. Pa ana de Las Palmas y Canal 6 (Campana, 527. Pe i co (EI Ca men, Jujuy) 24° 23' S 65° 07' W
Buenos Ai es) 34° 07' S 59° 02' W 528. Picada Vieja (Obe a, Misiones) 27° 29' S 55° 07' W
505. Pa que 9 de Julio, Tucu nan (Capi al, Tucuman) 529. Pico Salamanca (Escalan e, Chubu ) 45° 25' S 67°
26° 47' S 65° 15' W 25'W
506. Pa que Nacional Ban u, Finca Jakulica, Angos o 530. Pied a Tendida, 12 km V.lNW Bu uyacu along
del Rio Pescado, 650 m (San a Vic o ia, Sal a) Rio Cajon, 2,500 . (Bu uyacu, Tucu nan) 26°
22° 33' S 64° 32' W 30' S 64° 52' W
507. Pa que Nacional Ba i u, Finca Lipeo, a 15 km de 531. Pied a Tendida, 5 km W de Dique El Cajon
Las Jun as, 600 m (San a Vic o ia, Sal a) 22° 27' (Bu uyacu, Tucuman) 26° 30' S 64° 52' W
S 64° 46' W 532. Pied abuena (Bu uyacu, Tucu nan) 26° 44' S 64°
508. Pa que Nacional Ba i u, Las Jun as de Rio Lipeo 40'W
y Be mejo (San a Vic o ia, Sal a) 22° 26' S 64° 533. Pilcomayo (Pilcomayo, Fo mosa) 25° 22' S 57°
32'W 39'W
509. Pa que Nacional Ba i u, nacien e A oyo 534. Pina Vela dez, 1560 m (Ta Viejo, Tucuman)
San elmi a, 900 m (San a Vic o ia, Sal a) 22° 31' 26° 43' S 65° 22' W
S 64° 34' W 535. Pique e (La Capi al, San a Fe) 31° 34' S 60° 43' W
510. Pa que Nacional EI Palma (Colon, En e Rios) 536. Piqui enda Viejo, 30 km N Ta agal (San Ma in,
31° 59' S 58° 18' W Sal a) 22° 21' S 63° 50' W
511. Pa que Nacional EI Rey (An a, Sal a) 24° 42' S 537. Piqui enda Viejo, 6 km W (San Ma in, Sal a)
64° 38' W 22° 21' S 63° 50' W
512. Pa que Nacional EI Rey, A oyo de Los Pues os 538. Piqui enda Viejo, 8 km W (San Ma in, Sal a)
(An a, Sal a) 24° 42' S 64° 38' W 22° 21' S 63° 50' W
513. Pa que Nacional Iguazu (Iguazu, Misiones) 25° 539. Plan a Caimanci o, Pa que Nacional Calilegua
41' S 54° 27' W (Ledesma, Jujuy) 23° 44' S 64° 36' W
514. Pa que Nacional Iguazu, ca a a as (Iguazu, 540. Pla anos (Be aza egui, Buenos Ai es) 34° 47' S
Misiones) 25° 41' S 54° 27' W 58° 10' W
515. Pa que Nacional Iguazu, ex ae opue o (Iguazu, 541. Playa La ga, Rio Los Sosa, Ru a 307 km 19.7
Misiones) 25° 41' S 54° 27' W (Mon e os, Tucuman) 27° 03' S 65° 40' W
516. Pa que Nacional Iguazu, palmi al en u a 107 542. Po nan, 95 km S Andalgala nea balnea io, 4,400
(Iguazu, Misiones) 25° 41' S 54° 27' W . (Po nan, Ca ama ca) 28° 24' S 66° 13' W
517. Pa que Nacional Rio Pilcomayo (Pilcomayo, 543. Posadas (Capi al, Misiones) 27° 22' S 55° 53' W
Fo mosa) 25° 00' S 58° 15' W 544. Posadas, 61 km W, along Hwy 12 (I uzaingo,
518. Pa que P o incial EI Cochuna, km 40 sob e u a Co ien es) 27° 37' S 56° 28' W
47 (Chicligas a, Tucuman) 27° 18' S 65° 54' W 545. Po e o Dike, 13 km N Andalgala (Andalgala,
519. Paso de la C uz (An a, Sal a) 24° 55' S 64° 30' W Ca ama ca) 27° 32' S 66° 20' W
520. Paso del Rey, I km N, along A oyo de la Canada 546. Po e o Ri e Dike, EI Po e o (Andalgala,
Honda (Co onel P ingles, San Luis) 32° 52' S 66° Ca ama ca) 27° 32' S 66° 20' W
OO'W 547. Po e o Ri e Dike, Po e o, App ox. 13 km N
521. Paso del Rey, 9 km N (Co onel P ingles, San Andalgala (Andalgala, Ca ama ca) 27° 32' S 66°
Luis) 32° 50' S 66° 00' W 20'W
522. Paso Flo es (Pilcaniyeu, Rio Neg o) 40° 35' S 70° 548. Pozo del Ga o (Gene al Gue nes, Chaco) 24° 40'
48'W S 61° 58' W
523. Paso Quin ana, Rosa io de La F on e a (Rosa io 549. Pozo del Ga o, 3 km E Pun a del Agua (Gene al
de La F on e a, Sal a) 25° 48' S 64° 58' W Giiemes, Chaco) 24° 40' S 61° 58' W
524. Pa ino (Pa ino, Fo mosa) 24° 30' S 59° 30' W 550. Pozo del Tig e (Pa ino, Fo mosa) 24° 54' S 60°
525. Pa quia (Independencia, La Rioja) 30° 09' S 66° 19'W
57'W 551. Pozo Hondo (Jimenez, San iago del Es e o) 27°
526. Pede nal (Sa mien o, San Juan) 31° 59' S 68° 10' S 64° 30' W
44'W 552. Pozo Hondo, Es ancia EI Guapo (Jimenez,
270 SPECIAL PUBLICATIONS, MUSEUM OF TEXAS TECH UNIVERSITY
San iago del Es e o) 27° 10' S 64° 30' W 576. Queb ada de Lules, 11 kIn SW San Pablo (Lules,
553. P ime o de Mayo (U uguay, En e Rios) 32° 15' Tucuman) 26° 56' S 65° 21' W
S 58° 25' W 577. Queb ada del A oyo As ica, 8 km W As ica
554. P o undidad (Candela ia, Misiones) 27° 35' S 55° (Valle Fe il, San Juan) 30° 54' S 67° IT W
4T W 578. Queb ada del To o (T ancas, Tucuman) 26° 18' S
555. P onunciamien o (U uguay, En e Rios) 32° 21' 65° 41' W
S 58° 26' W 579. Quequen (Necochea, Buenos Ai es) 38° 32' S 58°
556. P o incia de Fo mosa, 24° 56' 59° 12' (Pa ino, 42'W
Fo mosa) 24° 56' S 59° 12' W 580. Rancho Los Yngleses, Ajo (Gene al La alle,
557. Puen e sob e e1Rio Be mejo (San Ma in, Chaco) Buenos Ai es) 36° 31' S 56° 53' W
26° 20' S 59° 22' W 581. Rayo Co ado-Ce o Colo ado (Rio Seco,
558. Pue o Agui e (=Pue o Iguazu) (Iguazu , Co doba) 30° 04' S 63° 50' W
Misiones) 25° 35' S 54° 33' W 582. Reconquis a (Gene al Obligado, San a Fe) 29° 10'
559. Pue o Be mejo (Be mejo, Chaco) 26° 56' S 58° S 59° 50' W
30'W 583. Rese a Eco lo gica Nacu ian (San a Rosa,
560. Pue o Pilcomayo (Pilcomayo, Fo mosa) 25° 22' Mendoza) 34° 03' S 67° 58' W
S 57° 39' W 584. Rese a P o incial "Aguas Chiqui as" EI Cadillal
561. Pue o Pi ay, Ru a 12 (Mon eca lo, Misiones) 26° (Bu uyacu, Tucuman) 26° 37' S 65° 12' W
28' S 54° 43' W 585. Rese a P o incial San a Ana, Remanso del
562. Pue o Rico (Libe ado Gene al San Ma in, Gallego (Rio Chico, Tucuman) 27° 26' S 65° 46' W
Misiones) 26° 48' S 55° 02' W 586. Rese a Tel eca (La alle, Mendoza) 32° 20' S 67°
563. Pue o San a Fe (La Capi al, San a Fe) 3 1°40' S 45'W
60° 40' W 587. Resis encia (San Fe nando, Chaco) 27° 2T S 59°
564. Pue o Velaz (Laishi, Fo mosa) 26° 40' S 58° OO'W
32' W 588. Resis encia, 20 km N (P ime o de Mayo, Chaco)
565. Pues o A oyo Cabeza de Vaca, 30 km N 27° 16' S 58° 58' W
Apolina io Sa a ia (O an, Sal a) 23° 55' S 64° 589. Resis encia, ma genes del Rio Pa ana (San
02'W Fe nando, Chaco) 27° 27' S 58° 5T W
566. Pues o Co ade as (Bu uyacu, Tucu nan) 26° 18' 590. Rincon de Luna (Concepcion, Co ien es) 28° 38'
S 64° 37' W S58°10'W
567. Pues o Di isade o, 35 km S, 5 km E lng. 591. Rincon de Papagayos, 2 km E Papagayos
Guille mo N. Jua ez (Ma acos, Fo mosa) 24° 14' (Chacabuco, San Luis) 32° 40' S 64° 5T W
S 61° 56' W 592. Rio Be mejo, 10 mi S Colonia km 503 (Pa ino,
568. Pues o F on e izo 6 km W La Bande i a Fo mosa) 25° 26' S 60° 15' W
(Andalgala, Ca ama ca) 27° 19' S 65° 58' W 593. Rio Blanco, 35 k n SW Sal a (Rosa io de Le ma,
569. Pues o Viejo, 16 km N Guemes, Rio Las Pa as Sal a) 24° 59' S 65° 35' W
(EI Ca men, Jujuy) 24° 30' S 65° 08' W 594. Rio Ca abe1as (Campana, Buenos Ai es) 34° 07'
570. Punilla, Valle He moso (Punilla, Co doba) 31° S 59° 02' W
08' S 64° 30' W 595. Rio Ceballos (Colon, Co doba) 31° II'S 64° 20' W
571. Pun a La a (Ensenada, Buenos Ai es) 34° 49' S 596. Rio Cua o (Rio Cua o, Co doba) 33° 08' S 64°
57° 59' W 18'W
572. Pun a Loyola, S. Pa agonia (Gue Aike, San a 597. Rio de O o, Gene al Vedia (Be mejo, Chaco) 26°
C uz)51°37'S69°01' W 56' S 58° 41' W
573. Pun a Rasa, Cabo San An onio (Gene al La alle, 598. Rio de Zo a y c uce con u a 34 (Ledesma, Jujuy)
Buenos Ai es) 36° 23' S 56° 43' W 23° 44' S 64° 40' W
574. Queb ada de Acambuco, 5 km W Dique I iyu o 599. Rio del Valle (An a, Sal a) 24° 38' S 64° 16' W
(San Ma in, Sal a) 22° 10' S 63° 53' W 600. Rio Dulce, 5 km E Las Te mas, camino a Vina a (Rio
575. Queb ada de Lopez, San F ancisco del Mon e de Hondo, San iago del Es e o) 27° 30' S 64° 46' W
O o (Ayacucho, San Luis) 32° 40' S 66° 07' W 601. Rio E1 Na anjo, 14 km W de Ru a p o incial 5
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