h ps://www.biodi e si ylib a y.o g/
Ec opa asi es o Panama.
Chicago,Field Museum o Na u al His o y,1966.
h ps://www.biodi e si ylib a y.o g/bibliog aphy/2633
Fieldiana. Zoology. Special Publica ions.:
h ps://www.biodi e si ylib a y.o g/i em/20461
A icle/Chap e Ti le: Chap e : Checklis o he mammals o Panama
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Hall and Kelson, 1959) a e ma ked wi h a dagge ( ). In ins ances
whe e I was no he ini ial collec o o a species he e epo ed in Panama
o he i s ime, I ha e no ed his ac in he accoun o he species.
Geog aphic dis ibu ion has been de e mined om specimens and
om li e a u e (ci a ions omi ed o b e i y). Mos o he specimens o
Panamanian mammals a e in he Academy o Na u al Sciences o Phila-
delphia, he Ame ican Museum o Na u al His o y, he B i ish Museum
(Na u al His o y), he Museum o Compa a i e Zoology (Ha a d Uni-
e si y), and he Uni ed S a es Na ional Museum, o in collec ions in Pan-
ama. The b ie no es on habi s and habi a a e based en i ely on Pana-
manian obse a ions.
O de MARSUPIALIA
Family Didelphidae
Calu omys de bianus Wa e house
= Philande lanige Goldman
Common a lowe ele a ions (sea le el up o a leas 3200 ee ) , p obably
h oughou Panama. E e g een and deciduous o es ; a bo eal. Se e al
Panamanian subspecies: (1) C. d. de bianus Wa e house. Da ien wes o
Canal Zone and La Cho e a (Panama) ; simila specimens om Guanico
(Los San os), San iago (Ve aguas), and Boque on and Isla Pa ida (Chi i-
qui). (2) C. d. e idus Thomas (=C. d. cen cdis Hollis e ). Ca ibbean
coas o wes e n Panama (mainland, Isla Bas imen os, and Cayo Agua,
Bocas del To o) and San a Fe (Ve aguas). (3) C. d. nau lcus Thomas.
Islands o he Paci ic coas o wes e n Panama (Isla B a a, Chi iqui; Isla
Gobe nado a and Isla Cebaco, Ve aguas) ; simila specimens om Pa i a
(He e a), and Panama Viejo (Panama). (4) C. d. pallidus Thomas. Pa-
ci ic coas o ex eme wes e n Panama (Di ala and Bugaba, Chi iqui) .
Monodelphis adus a Thomas
= Pe amys melanops Goldman
= Monodelphis melanops Hall and Kelson
Ra e a medium ele a ions in ex eme eas e n Panama. Specimens
om Cana, 2000 ee , Guayabo, and Taca cuna Village, 1950 ee (Da ien) .
T apped among boulde s on a d y, pa ially open g a el ba (Taca cuna
Village) and among ocks on a i e bank in hea y o es (Cana). Mono-
delphis adus a occu s om eas e n Panama h ough Colombia and Ecuado
[Begin Page: Page 755]
HANDLEY : CHECKLIST OF MAMMALS 755
o cen al Pe u wi hou exhibi ing geog aphic a ia ion. Pe amys melanops
Goldman and Pe amys pe u ianus Osgood a e synonyms o adus a. Thei
supposed dis inc i e cha ac e is ics a e me ely indi idual a ia ions. On
he o he hand, Monodelphis osgoodi Dou o wes e n Boli ia and sou he n
Pe u is p obably a dis inc species. I has been ega ded as a subspecies
o M. adus a.
* Ma mosa impa ida Tschudi
Ra e. A high ele a ions in ex eme eas e n Panama. 0. P. Pea son
collec ed specimens on Loma Cana, 4900 ee , and Ce o Pi e, 5300 ee
(Da ien). The Panamanian subspecies is M. i. caucae Thomas.
Ma mosa in ic a Goldman
Ra e. A medium ele a ions (1500-4000 ee ), possibly h oughou
Panama, in sui able habi a s. I has been apped on he g ound, and on o
unde logs in mois og o es . Specimens om Cana, Taca cuna Casi a,
and Taca cuna Laguna (Da ien) ; Ce o Azul (Panama) ; and Cylind o
(Bocas del To o).
Ma mosa mexicana Me iam
= Ma mosa mexicana Goldman (pa .)
Uncommon. In he lowland sa annas o wes e n Panama. The sub-
species in Panama is M. m. sa anna um Goldman. The ela ionship o
M. mexicana o M. obinsoni is no clea .
* Ma mosa species
A species esembling Ma mosa phaea Thomas has been aken a Taca -
cuna Casi a, 1500 ee (Da ien) ; A mila (San Bias) ; Salamanca Hyd o-
g aphic S a ion (Canal Zone) ; and Almi an e (Bocas del To o). E e -
g een o es ; a bo eal.
Ma mosa obinsoni Bangs
= Ma mosa ul i en e Goldman
= Ma mosa mexicana Goldman (pa .)
= Ma mosa mi is Hall and Kelson
Abundan h oughou a la ge pa o Panama om sea le el o ele-
a ions o 4000-6000 ee . Semi-a bo eal, bu o en apped on he g ound.
Two subspecies ha e been epo ed in Panama : (1) M. . ul i en e Bangs,
Isla Saboga and Isla del Rey ; (2) M. . is hmica Goldman, mainland. P ac-
ically ubiqui ous. Fo use o he name Ma mosa obinsoni Bangs in place
o M. mi is Bangs, see Cab e a (1958, p. 24).
Philande opossum Linnaeus
= Me achi us opossum Goldman
Locally abundan a lowe ele a ions (up o 2000 ee ) , possibly h ough-
ou Panama. Mos ly e es ial, bu also climbs. E e g een and deciduous
o es . The subspecies in Panama is P. o. uscog iseus J. A. Allen.
[Begin Page: Page 756]
756 ECTOPARASITES OF PANAMA
Me achi us nudicauda us E. Geo oy S .-Hilai e
Locally common a lowe ele a ions (up o 2000 ee ) , possibly h ough-
ou Panama. Te es ial. E e g een and deciduous o es . The subspecies
in Panama is M. n. den aneus Goldman. Whe e Me achi us nudicauda us
and Philande opossum occu oge he one is usually much mo e abundan
han he o he .
Didelphis ma supialis Linnaeus
Abundan h oughou Panama om sea le el up o 5000 ee . Ubiqui-
ous. Te es ial, bu also climbs. Th ee subspecies a e hough o occu
in Panama: (1) D. m. ba yi Thomas, Isla Coiba; (2) D. m. caucae J. A.
Allen (=Z). m. e ensis J. A. Allen) , mainland ; (3) D. m. pa iceps Goldman,
Isla del Rey.
Chi onec es minimus Zimme mann
= Chi onec es panamensis Goldman
= Chi onec es panamensis Hall and Kelson
Locally common o abundan h oughou , om sea le el o 4000 ee , in
sui able s eams. Aqua ic, possibly es ic ed o s eams wi h la ge
popula ions o sh imp. Specimens om Boca de Rio Paya, Cana, 2000
ee , Guayabo, Rio Jaque, and Taca cuna Village, 1950 ee (Da ien) ;
A mila and Mandinga (San Bias) ; Ce o Azul, 2000 ee (Panama) ; Fo
She man, New San Juan, and Ped o Miguel (Canal Zone) ; Ce o Hoya,
1450 ee and Guanico (Los San os) ; Rio Bubi and Rio Viejo, nea Sona
(Ve aguas) ; Rio Colo ado, 4000 ee (Chi iqui) ; and 7 km. SSW. o
Changuinola and Sibube (Bocas del To o) . S udy o specimens om B a-
zil, B i ish Guiana, Venezuela, and Cen al Ame ica shows ha only one
species can be dis inguished. Va ia ions in colo a ion and pe haps in
c anial cha ac e s as well sugges ha he species is poly ypic. The Pana-
manian subspecies is C. m. panamensis Goldman.
O de INSECTIVORA
Family So icidae
* C yp o is ende si Se ze
Ra e. Known by one specimen om Cylind o (Bocas del To o). Fog
o es . Te es ial.
C yp o is nig escens J. A. Allen
= C yp o is me us Goldman
= C yp o is me a Hall and Kelson
= C yp o is e sus Hall and Kelson
= C yp o is ze eki Hall and Kelson
Ra e. A high ele a ions in e e g een o es . Te es ial. Subspecies
in Panama: (1) C. n. me a Goldman, Ce o Pi e, 4500 ee (Da ien) ;
(2) C. n. ze eki Se ze (= C. e sus Goodwin), Ce o Pun a, 6500 ee , and
San a Cla a, 4200 ee (Chi iqui). These subspecies a e poo ly di e en i-
a ed om ypical C. nig escens o Cos a Rica.
[Begin Page: Page 757]
HANDLEY : CHECKLIST OF MAMMALS 757
O de CHIROPTERA
Family Emballonu idae
Rhynchonyc e is naso Wied-Neuwied
= Rhynchiscus naso Goldman
Abundan nea sea le el, p obably h oughou Panama. Flies o e o
nea o es -bo de ed s eams o lakes; oos s on ocks, logs, o unks o
ees o e wa e . Specimens om Boca de Rio Paya, Paya Village, and
Rio Jesuci o (Da ien) ; A mila and Mandinga (San Bias) ; Ba o Colo ado
Island, Co ozal, Fo Gulick, Gamboa, Laga e a, 2 mi. NW. o Ped o
Miguel, Rio Mandinga, and Summi (Canal Zone) ; Almi an e, 7 km. SSW.
o Changuinola, and Sibube (Bocas del To o) ; and San Felix (Chi iqui).
Fo use o he gene ic name Rhynchonyc e is Pe e s in place o Rhynchisciis
Mille , see Husson (1962, p. 35).
Saccop e yx bilinea a Temminck
Abundan nea sea le el, p obably h oughou Panama; occasionally
ound o 2000 ee . Flies o e o nea o es -bo de ed s eams, lakes, o
beaches; oos s on unks o ees, on logs and ocks, and in ca es and
houses. Specimens om Ci u o, Cana, and Ce o Sapo (Da ien) ; A mila
(San Bias) ; Alhajuela, Ancon, Balboa, Ba o Colo ado Island, Culeb a, Fo
Kobbe, Fo San Lo enzo, Fo She man, 3 mi. SE. o Gamboa, Ga un,
Las Cascadas, Madden Dam, O chid Island, Rio Puen e, Summi , Tabe nilla,
Venado Beach (Canal Zone) ; Ce o Azul, Chilib e, Chilib illo Ca es, Isla
Chape a, Isla San Jose, La Cho e a, Panama Vie jo, and Isla Taboga (Pa-
nama) ; Guanico (Los San os) ; and Almi an e (Bocas del To o).
Saccop e yx lep u a Sch ebe
Uncommon. Nea sea le el in he Canal Zone and eas e n Panama.
Flies o e o nea o es -bo de ed s eams and lakes. Specimens om
Boca de Rio Paya and Paya Village (Da ien) ; Paco a (Panama) ; and Alha-
juela, Ba o Colo ado Island, Fo Randolph, and Gamboa (Canal Zone) .
* Co mu a b e i os is Wagne
Uncommon. Lowlands o eas e n Panama and on he Ca ibbean coas
o wes e n Panama. E e g een o es . Roos s unde logs and ocks. Spec-
imens om Cape i (Da ien) ; A mila (San Bias) ; and Almi an e (Bocas
del To o) .
*Pe op e yx kapple i Pe e s
Ra e. Lowlands o eas e n Panama and on he Ca ibbean coas o
wes e n Panama. Roos ing in a ca e a Rio Puen e and ne ed o e a s eam
lowing h ough a pas u e a Almi an e. Specimens om Ya isa (Da ien) ;
Rio Puen e (Panama); and Almi an e (Bocas del To o). The Pana-
manian subspecies is P. k. kapple i Pe e s.
[Begin Page: Page 758]
758 ECTOPARASITES OF PANAMA
Pe op e yx mac o is Wagne
= Pe op e yx canina Goldman
Uncommon. Possibly widesp ead in he lowlands, bu p esen ly e-
co ded only in and nea he Canal Zone. Roos s in ca es and ock c e ices
nea wa e . Specimens om Buena Vis a (Colon) ; Balboa, Ga un, and
San a Rosa (Canal Zone) ; and Rio Puen e (Panama). The Panamanian
subspecies is P. m. mac o is Wagne .
Cen onyc e is maximiliani Fische
= Cen onyc e is cen alis Goldman
Ra e. Lowlands o he Canal Zone and he Paci ic coas o wes e n
Panama. Specimens om Ce o Mali (Da ien) , Ba o Colo ado Island and
Fo Clay on (Canal Zone) , and Bugaba (Chi iqui) . The Canal Zone speci-
mens we e aken in clea ings. The Panamanian subspecies is C. m. cen-
alis Thomas.
Diclidu us i go Thomas
Ra e. Canal Zone and Paci ic coas o wes e n Panama a low and
medium ele a ions. Specimens om Alb ook Field, Fo Gulick, and Gam-
boa (Canal Zone), and Boque e and Pueblo Nue o (Chi iqui).
Family Noc ilionidae
Noc ilio labialis Ke
= Di ias albi en e Goldman
Common a lowe ele a ions, p obably h oughou Panama. Flies o e
s eams and lakes ; oos s in hollow ees and houses. Specimens om Boca
de Rio Paya (Da ien) ; A mila (San Bias) ; Paco a (Panama) ; Nue o
Limon (Colon) ; Ba o Colo ado Island, Empi e, Fo Gulick, Gale a Island,
Gamboa, Juan Mina, and Puma Island (Canal Zone) ; Guanico (Los San os) ;
and Almi an e, 7 km. SSW. o Changuinola, and Sibube (Bocas del To o).
The Panamanian subspecies is N. I. labialis Ke (= Noc ilio mino Osgood.
See Cab e a, 1958, p. 55) .
* Noc ilio lepo inus Linnaeus
Common a lowe ele a ions, p obably h oughou Panama. Flies o e
s eams, lakes, and sea. Roos s in hollow ees. Specimens om Boca de
Rio Paya (Da ien) ; A mila (San Bias) ; Fo She man, Rio Chag es, and
Rio Mandinga (Canal Zone) ; and Almi an e and 7 km. SSW. o Changuinola
(Bocas del To o) . The Panamanian subspecies is 2V. I. mexicanus Goldman.
Family Phyllos omidae
P e ono us pa nellii G ay
= Chilonyc e is ubiginosa Goldman
zz Chilonyc e is pa nellii Hall and Kelson
om Fo Gulick (Canal Zone) and Almi an e and Sibube (Bocas del To o) .
The Panamanian subspecies is M. c. keenani Handley.
* Phyllos omus discolo Wagne
Common a lowe ele a ions, p obably h oughou Panama. E e g een
o es and ui g o es. Specimens om Boca de Rio Paya, Paya Village,
and Taca cuna Village, 1950 ee (Da ien) ; A mila, Mandinga, and Pue o
Obaldia (San Bias) ; Ce o Azul, 2000 ee , and Chilib illo Ca es (Panama) ;
Ba o Colo ado Island (Canal Zone) ; Guanico (Los San os) ; Almi an e, 7
and 14 km. SSW. o Changuinola, and Sibube (Bocas del To o). The Pan-
amanian subspecies is P. d. discolo Wagne .
[Begin Page: Page 762]
762 ECTOPARASITES OF PANAMA
Phyllos omus has a us Pallas
Common in lowlands, up o 2000 ee , h oughou Panama. Fo es and
ui g o es. Roos s in hollow ees, ca es, and houses. Specimens om
Boca de Cupe, Boca de Rio Paya, Boca de Rio Punusa, Cape i, El Real, and
Jaque (Da ien) ; A mila and Pue o Obaldia (San Bias) ; Cabima, Ce o
Azul, 2000 ee , Chepo, Chilib illo Ca es, Paco a, and Panama (Panama) ;
Balboa, Ba o Colo ado Island, Fo Gulick, Juan Mina, Madden Dam,
Pa aiso, and Summi (Canal Zone) ; Guanico (Los San os) ; Boque on
(Chi iqui) ; and Almi an e, Boca del D ago, Cayo Agua, Isla Bas imen os,
and Sibube (Bocas del To o). The Panamanian subspecies is P. h. pana-
mensis J. A. Allen.
* Phyllode ma s enops Pe e s
=. Phyllode ma sep en ionalis Hall & Kelson
Ra e. E e g een o es . Specimens om A mila (San Bias). The
Panamanian specimens combine he cha ac e s o he nominal species P.
s enops Pe e s and P. sep en ionalis Goodwin, bu a e nea e he o me .
Thus, he Panamanian subspecies is P. s. s enops Pe e s.
T achops ci hosus Spix
Common in lowlands and up o 2800 ee , p obably h oughou Panama.
Flies o e o es -bo de ed s eams. Roos s in ca es and houses. Specimens
om Rio Jesuci o and Taca cuna Village, 1950 ee (Da ien) ; A mila,
Mandinga, and Pue o Obaldia (San Bias) ; Ce o Azul, 2000 ee , Chepo,
Chilib illo Ca es, and Paco a (Panama) ; Fo She man (Canal Zone) ;
Guanico and Ce o Hoya, 2800 ee (Los San os) ; and 7-20 km. SSW. o
Changuinola, Isla Bas imen os, and Sibube (Bocas del To o). The Pan-
amanian subspecies is T. c. ci hosus Spix.
* Ch o op e us au i us Pe e s
Ra e. E e g een o es . Specimens om Taca cuna Casi a, 1500 ee
(Da ien), A mila (San Bias), and Ce o Azul, 2000 ee (Panama). I
doub ha subspecies a e ecognizable in C. au i us.
Vampy um spec um Linnaeus
= Vampy us spec um Goldman
Ra e. Ne ed o e o es -bo de ed s eams and in ui g o es. Speci-
mens om Taca cuna Village, 1950 ee (Da ien) ; A mila (San Bias) ;
Boque on (Chi iqui) ; and uppe Rio Changena, 2400 ee , and Sibube (Bo-
cas del To o) . The subspecies V. s. nelsoni Goldman does no appea o be
ecognizable. I ha e compa ed he ype wi h o he specimens om Panama
(7), Venezuela (4), T inidad (10), Ecuado (1), and Pe u (1). Al hough
he Mexican and Cen al Ame ican specimens a e age smalle han hose
om Sou h Ame ica, he di e ence is no g ea . The den al and c anial
cha ac e is ics ha Goldman supposed would dis inguish nelsoni all p o e
o be indi idual a ia ions.
[Begin Page: Page 763]
HANDLEY : CHECKLIST OF MAMMALS 763
* Glossophaga commissa isi Ga dne
Common in he lowlands o eas e n Panama, on he Ca ibbean coas o
wes e n Panama, and a medium ele a ions, possibly h oughou Panama.
E e g een o es . Specimens om Boca de Cupe, Boca de Rio Paya, Paya
Village, and Taca cuna Village, 1950 ee (Da ien) ; A mila and Mandinga
(San Bias) ; Candela ia Hyd og aphic S a ion and Ce o Azul, 2000 ee
(Panama) ; and Almi an e, Boca del D ago, Cayo Agua, and Sibube (Bocas
del To o).
Glossophaga so icina Pallas
Abundan in he lowlands in cen al Panama and on he Paci ic coas
o wes e n Panama and locally common on he Ca ibbean coas . Deciduous
and e e g een o es . Roos s in ca es, cul e s, houses, hollow ees, and
hollow logs. Specimens om Mandinga (San Bias) ; Colon and Po obelo
(Colon) ; Isla Saboga, La Cho e a, Nue a Go gona, Paco a, Panama, and
Panama Viejo (Panama) ; many locali ies h oughou he Canal Zone; El
Cope, Rio Ha o, and San a Cla a (Code) ; Guanico (Los San os) ; Isla Canal
de A ue a (Ve aguas) ; and Almi an e, Boca del D ago, 7 km. SSW. o
Changuinola, Isla Bas imen os, Isla Colon, Isla Escudo de Ve aguas, and
Sibube (Bocas del To o) . The Panamanian subspecies is G. s. leachii G ay.
Lonchophylla mo dax Thomas
= Lonchophylla conca a Goldman
= Lonchophylla conca a Hall and Kelson
Ra e. E e g een o es . Roos s in ca es. Specimens om Cana, 2000
ee , and Taca cuna Village, 1950 ee (Da ien) ; and A mila and Pue o
Obaldia (San Bias) . The Panamanian subspecies is L. m. conca a Goldman,
which is only sligh ly di e en ia ed om ypical L. mo dax.
Lonchophylla obus a Mille
Common. Eas e n Panama and Ca ibbean lowlands o wes e n Pan-
ama. E e g een o es . Roos s in ca es. Specimens om Cana, 2000 ee ,
Taca cuna Casi a, 2600 ee , and Taca cuna Village, 1950 ee (Da ien) ;
A mila and Pue o Obaldia (San Bias) ; Ce o Azul, 2000 ee , Chilib illo
Ca es (Panama) ; Buena Vis a (Colon) ; Ba o Colo ado Island, Salamanca
Hyd og aphic S a ion and Summi (Canal Zone) ; and Almi an e and Isla
Bas imen os (Bocas del To o).
* Lonchophylla homasi J. A. Allen
Ra e. Known in Panama by a single specimen ne ed in a i e side
clea ing a Boca de Rio Paya (Da ien) .
* Lionyc e is spu elli Thomas
Ra e. E e g een o es . Specimens om Cana, 2000 ee (Da ien)
and A mila (San Bias) . Fi s aken in Panama by O. P. Pea son.
[Begin Page: Page 764]
764 ECTOPARASITES OF PANAMA
* Anou a cul a a Handley
Uncommon. E e g een o es . Specimens om Taca cuna Village,
1950 ee (Da ien) ; Ce o Pun a, 5300 ee (Chi iqui) ; and uppe Rio
Changena, 2400 ee (Bocas del To o).
* Anou a geo oyi G ay
Uncommon. E e g een o es . Specimens om Ce o Hoya, 3000 ee
(Los San os) ; Ce o Pun a, 5300 ee (Chi iqui) ; and uppe Rio Changena,
4800 ee (Bocas del To o). The Panamanian subspecies is A. g. lasiopyga
Pe e s.
* Hylonyc e is unde wood! Thomas
Ra e. Known in Panama only by a specimen aken a San a Cla a,
4200 ee (Chi iqui) by F. A. Ha man.
* Lichonyc e is obscu a Thomas
Ra e. E e g een o es and ui g o es. Specimens om Taca cuna
Village, 3200 ee (Da ien) ; A mila (San Bias) ; and Almi an e and uppe
Rio Changena, 2400 ee (Bocas del To o).
Ca ollia cas anea H. Allen
= Hemide ma cas aneum Goldman
= Ca ollia cas anea Hall and Kelson (pa .)
Common a lowe ele a ions h oughou Panama. Fo es and ui
g o es. Appa en ly mos abundan in e e g een o es . Found oos ing
unde o e hanging s eam bank and in mine. Specimens om Boca de
Rio Paya, Cana, 2000 ee , Paya Village, Rio Se egan i, Taca cuna Casi a,
2600 ee , and Taca cuna Village, 3200 ee (Da ien) ; A mila, Mandinga,
and Pue o Obaldia (San Bias) ; Ce o Azul, 2000 ee (Panama) ; F ijoli o
(Colon) ; Ba o Colo ado Island, Chi a Chi a, and Fo Kobbe (Canal
Zone) ; Ce o Hoya, 3000 ee (Los San os) ; and Almi an e, Boca del D ago,
7 km. SSW. o Changuinola, and Sibube (Bocas del To o).
Ca ollia cas anea is mono ypic. I has been con used wi h Ca ollia
sub u a Hahn, bu i can be easily dis inguished om ha species by i s
smalle size ( o ea m usually less han 38 mm.) ; dulle , less sha ply de-
ined banding o do sal hai s ; occlusal ou line o c own o inne lowe in-
ciso usually o oid (wide han long) a he han iangula (longe han
wide) ; i s h ee lowe pos -canine ee h g aded (inc ease in size om he
i s ) a he han subequal ; M almos ' la -c owned (all cusps obsole e) , sig-
ni ican ly lowe han adjacen ee h ; second uppe p emola wi hou pos e o-
ex e nal cusp. Ca ollia cas anea anges om Cos a Rica h ough Panama
o no he n and wes e n Colombia.
Ca ollia pe spicilla a Linnaeus
= Hemide ma pe spicilla um Goldman
Abundan h oughou he lowlands o Panama ; uncommon o a e abo e
[Begin Page: Page 765]
HANDLEY : CHECKLIST OF MAMMALS 765
3000 ee . Ubiqui ous. The Panamanian subspecies is C. p. az eca Saus-
su e.
* Ca ollia sub u a Hahn
= Ca ollia cas anea Hall and Kelson (pa .)
Locally abundan h oughou eas e n Panama and a high ele a ions
and on he Ca ibbean coas o wes e n Panama. Usually he mos abundan
species o Ca ollia abo e 3000 ee . E e g een o es . Roos s in ca es
and in hollow ees. Specimens om Paya Village, Rio Chucunaque, Rio
Se egan i, Taca cuna Casi a, 1500 ee , and Taca cuna Village, 1950 ee
(Da ien) ; A mila, Mandinga, and Pue o Obaldia (San Bias) ; Ce o Azul
(Panama) ; Ce o Pun a, 5300 ee (Chi iqui) ; and Almi an e, Boca del
D ago, Cayo Agua, 7 km. SSW. o Changuinola, Isla Bas imen os, Isla Colon,
uppe Rio Changena, 2400-4800 ee , and Sibube (Bocas del To o) .
Ca ollia sub u a has been con used wi h Ca ollia cas anea, bu ac ually
i is mos like and is mo e closely ela ed o Ca ollia pe spicilla a. F om
C. pe spicilla a, C. sub u a di e s in a e age smalle size ( o ea m 38-42
mm. s. 41-45 mm.) ; a e age hai ie o ea m, ibia, and oo ; a e age
longe , silkie u ; less c owded and pos e io ly mo e di e gen oo h ows ;
educed cusps o cheek ee h (especially p o ocone o M 1 ) ; ou e inciso s
no educed ; highes poin o P 4 well ahead o he mid-leng h o he oo h.
Ca ollia sub u a seems o be mono ypic. I anges om wes e n Mexico o
Colombia and Venezuela.
S u ni a lilium E. Geo ey S .-Hilai e
Abundan a lowe ele a ions (sea le el o abou 4000 ee ) h oughou
Panama. Fo es s and ui g o es. Specimens om Boca de Cupe, Boca
de Rio Paya, Paya Village, and Taca cuna Village, 1950 ee (Da ien) ;
A mila, Mandinga, and Pue o Obaldia (San Bias) ; Ce o Azul, 1800-2000
ee (Panama) ; Queb ada Boni a (Colon) ; Ce o Hoya and Guanico (Los
San os) ; Volcan de Chi iqui (Chi iqui) ; and Almi an e and Isla Bas imen-
os (Bocas del To o). The Panamanian subspecies is S. L pa idens Gold-
man. Specimens om Taca cuna Village a e la ge han hose om lowe
ele a ions in Da ien and elsewhe e in Panama and may ep esen an unde-
sc ibed subspecies o <S. lilium.
- S u ni a ludo ici An hony
Abundan a highe ele a ions. E e g een o es . Specimens om
Ce o Mali, 4700 ee , and Taca cuna Village, 1950 ee (Da ien) ; Ce o
Pun a, 4600-5300 ee (Chi iqui) ; and uppe Rio Changena, 2400-5000 ee
(Bocas del To o).
U ode ma biloba um Pe e s
Abundan a lowe ele a ions h oughou Panama. Usually ound nea
s eams and in ui g o es. Roos s unde palm lea es. Specimens om
Boca de Cupe, Boca de Rio Paya, Cape i, Chepigana, Jaque, and Paya
[Begin Page: Page 766]
766 ECTOPARASITES OF PANAMA
Village (Da ien) ; A mila, Mandinga, and Pue o Obaldia (San Bias) ;
Candela ia Hyd og aphic S a ion, Ce o Azul, 850 ee , Isla Saboga, Isla
San Jose, Isla del Rey, La Cho e a, Paco a, Panama, and Panama Vie jo
(Panama) ; Queb ada Boni a and Colon (Colon) ; Ancon, Ba o Colo ado
Island, F ance Field, Rio Indio, and Summi (Canal Zone) ; Guanico (Los
San os) ; El Cope, 1450 ee , and El Po e o (Code) ; Boque on and Bugaba
(Chi iqui) ; and Almi an e, Boca del D ago, 7 km. SSW. o Changuinola,
Isla Bas imen os, Pun a de Pena, and Sibube (Bocas del To o). The Pan-
amanian subspecies is U. b. biloba um Pe e s (= U. con exum Lyon).
* Vampy ops do salis Thomas
Uncommon. Specimens om Taca cuna Village, 1950 ee (Da ien).
E e g een o es . Ne ed o e o es -bo de ed s eam.
Vampy ops helle i Pe e s
= Pla y hinus helle i Hall and Kelson
Abundan om sea le el o 4000 ee h oughou Panama. Fo es s and
ui g o es. Specimens om Boca de Rio Paya, Cana, 2000 ee , Jaque,
Paya Village, Rio Se egan i, Taca cuna Casi a, 1950 ee , Taca cuna Laguna,
3000 ee , and Taca cuna Village, 1950 ee (Da ien) ; A mila, Mandinga,
and Pue o Obaldia (San Bias) ; Cabima, Ce o Azul, 2000 ee , Isla del Rey,
and Isla San Jose (Panama) ; Bas Obispo, Chi a Chi a, and Pa aiso
(Canal Zone) ; Ce o Hoya and Guanico (Los San os) ; Isla Cebaco (Ve a-
guas) ; and Almi an e, Boca del D ago, Cayo Agua, 7 km. SSW. o Chan-
guinola, Isla Colon, Pun a de Pena, uppe Rio Changena, 2400 ee , and
Sibube (Bocas del To o). Vampy ops za hinus H. Allen is a synonym.
* Vampy ops i a us Pe e s
Enchis henes ha ii Thomas
Uncommon. E e g een and deciduous o es . Specimens om Taca -
cuna Village, 1950 ee (Da ien) ; Qua y Heigh s (Canal Zone) ; and Ce o
Hoya, 2600 ee (Los San os) .
* Cen u io senex G ay
Uncommon. E e g een o es . Specimens om Taca cuna Village,
1950 ee (Da ien) ; Las Cumb es and Nue a Go gona (Panama) ; Ba o
Colo ado Island and C is obal (Canal Zone) ; Ce o Hoya, 1800 ee (Los
San os); and Almi an e and Sibube (Bocas del To o). Fi s aken in
Panama by Ca l Ko o d.
Family Desmodidae
Desmodus o undus E. Geo ey S .-Hilai e
Locally abundan h oughou Panama, and on o -sho e islands, om
sea le el o 5600 ee . Ubiqui ous. Roos s in hollow ees, ca es, and
houses. The Panamanian subspecies is D. . mu inus Wagne .
* Diaemus youngii Jen ink
Ra e. E e g een o es . Specimens om A mila (San Bias) and Isla
Bas imen os (Bocas del To o). The Panamanian subspecies is D. y. cypse-
linus Thomas.
Diphylla ecauda a Spix
= Diphylla cen alis Goldman
Ra e. E e g een o es . Specimens om Boque e (Chi iqui) and
Almi an e and Sibube (Bocas del To o). Diphylla ecauda a is p obably
mono ypic (see Bu and S i on, 1961, p. 37).
Family Na alidae
Na al us s amineus G ay
= Na alus mexicanus Goldman
= Na alus mexicanus Hall and Kelson
Ra e. Roos s in ca es. Specimens om Chilib illo Ca es (Panama) ;
Fo San Lo enzo (Canal Zone) ;Penonome (Code) ;IslaCoiba (Ve aguas).
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770 ECTOPARASITES OF PANAMA
The Panamanian subspecies is N. s. mexicanus Mille (=N.s. sa u a us Dai-
ques and Hall) . Fo he axonomy o N. s amineus see Goodwin (1959) .
Family Fu ip e idae
* Fu ip e us ho ens F. Cu ie
Ra e. E e g een o es . Known in Panama only by specimens ound
oos ing wi h Mimon cozumelae in a small ca e on he banks o he Rio
Changuinola, 20 km. SSW. o Changuinola (Bocas del To o).
Family Thy op e idae
*Thy op e a icolo Spix
Ra e. Roos s in olled Heliconia lea es. Specimens om Rio Jesuci o
and Taca cuna Village (Da ien) ; Ba o Colo ado Island and Co ozal (Canal
Zone) ; and on ail om Chi iquici o o Boque e (Bocas del To o). The
Panamanian subspecies is T. . albi en e Tomes (=T. . albigula G. M.
Allen).
Family Vespe ilionidae
* Myo is albescens E. Geo oy S .-Hilai e
Common. Lowlands o eas e n Panama and Ca ibbean coas o wes e n
Panama. E e g een o es . Specimens om Boca de Rio Paya and Taca -
cuna Village, 1950 ee (Da ien) ; San Pablo and Tabe nilla (Canal Zone) ;
and 7-20 km. SSW. o Changuinola and Sibube (Bocas del To o) .
* Myo is chiloensis Wa e house
Ra e. E e g een o es . Roos s in ca es. Known in Panama only by
specimens om Ce o Pun a, 5300-5800 ee (Chi iqui). Fi s aken in
Panama by Cha les M. Keenan and Ve non J. Tip on. These specimens
ep esen an undesc ibed subspecies.
Myo is nig icans Schinz
Abundan h oughou a lowe ele a ions, om sea le el o a leas 3200
ee ele a ion and pe haps ascending highe in he moun ains. Roos s in
ca es and houses. The Panamanian subspecies is M. n. nig icans Schinz.
I seems likely ha Myo is nig icans, as now cons i u ed, is a composi e
species.
* Myo is simus Thomas
Uncommon. E e g een o es . Specimens om Boca de Rio Paya and
Taca cuna Village, 1950 ee (Da ien) ; A mila (San Bias) ; and Ce o Azul,
2000 ee (Panama) . The Panamanian subspecies is M. s. ipa ius Handley.
Ep esicus b asiliensis Desma es
= Ep esicus p opinquus Goldman
Uncommon, a low ele a ions in cen al and wes e n Panama. E e -
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HANDLEY : CHECKLIST OF MAMMALS 771
g een o es . Roos s in houses. Specimens om Ce o Hoya (Los San os) ;
and 7 km. SSW. o Changuinola and Sibube (Bocas del To o). The Pana-
manian subspecies is E. b. p opinquus Pe e s.
* Ep esicus chi iquinus Thomas
Uncommon, on he Ca ibbean coas o eas e n Panama and a medium
ele a ions h oughou . E e g een o es . Specimens om Taca cuna Vil-
lage, 1950 ee (Da ien) ; A mila (San Bias) ; Ce o Azul, 2000 ee (Pan-
ama) ;Boque e (Chi iqui) ; uppe Rio Changena, 2400 ee (Bocas del To o).
Ep esicus uscus Paliso de Beau ois
Ra e, a high ele a ions in wes e n Panama. E e g een o es . Roos s
in ee holes. Specimens om El Valle (Code) ; Boque e (Chi iqui) ; and
uppe Rio Changena, 4800 ee (Bocas del To o). The Panamanian sub-
species is E. . mi ado ensis H. Allen.
* Ep esicus innoxius Ge ais
Ra e, a low ele a ions on he Paci ic coas al plain. Specimens om
San Pablo (Canal Zone) ; Tocumen (Panama) ; and Boque on (Chi iqui).
Rhogeessa umida H. Allen
=i Rhogeessa pa ula Hall and Kelson (pa .)
Uncommon, a lowe ele a ions. Open coun y. Roos s in houses. Spec-
imens om La Palma (Da ien) ; nume ous locali ies (Canal Zone) ; San a
Cla a (Code) ; Guanico (Los San os) ; Bugaba (Chi iqui) ; and Almi an e, 7
km. SSW. o Changuinola, and Isla Bas imen os (Bocas del To o). The
Panamanian subspecies is R. . umida H. Allen (Goodwin, 1958).
Lasiu us bo ealis Miille
= Nyc e is bo ealis Goldman
Uncommon, al hough widely dis ibu ed bo h geog aphically and al i-
udinally. Ne ed o e s eams. Specimens om Taca cuna Village,
1950 ee (Da ien) ; A mila (San Bias) ; Balboa and Fo Clay on (Canal
Zone) ; Calob e (Ve aguas) ; Ce o Hoya, 1800 ee (Los San os) ; and
Boque e, 3500 ee , and Ce o Pun a, 5300 ee (Chi iqui). The Panaman-
ian subspecies is L. b. an zii Pe e s.
* Lasiu us cas aneus Handley
Ra e. E e g een o es . Specimens om Taca cuna Village, 1950 ee
(Da ien) and A mila (San Bias).
Lasiu us ega Ge ais
= Dasyp e us ega Goldman
= Dasyp e us ega Hall and Kelson
Ra e. Specimens om Fo Clay on (Canal Zone) ; Ce o Hoya, 1800
ee (Los San os) ; and Bugaba, 800 ee (Chi iqui) . The Panamanian sub-
species is L. e. panamensis Thomas. Fo no es on he axonomy o his
species, see Handley (1960, p. 473).
[Begin Page: Page 772]
772 ECTOPARASITES OF PANAMA
* Lasiu us eg egius Pe e s
Ra e. E e g een o es . A specimen om A mila (San Bias) and he
holo ype om San a Ca a ina, B azil, desc ibed in 1870, a e he only known
specimens o his species.
Family Molossidae
Molossops plani os is Pe e s
= Cynomops plani os is Hall and Kelson
Uncommon. Roos s in houses. Specimens om Paco a and Panama
(Panama) ; and Balboa, Cocoli, Fo Amado , Fo Clay on, La Boca, and
Mi a lo es Locks (Canal Zone) . The Panamanian subspecies is M. p. plani-
os is Pe e s. Specimens om Taca cuna Village (Da ien) p obably ep-
esen ano he species o Molossops.
* Tada ida b asiliensis I. Geo ey S .-Hilai e
Uncommon. Roos s in ca es. Specimens om Ce o Pun a, 5700 ee ,
and Rio Chi iqui Viejo a Palo San o, 4200 ee (Chi iqui) ; and uppe Rio
Changena (Bocas del To o). The Panamanian subspecies is T. b. b asilien-
sis I. Geo oy S .-Hilai e.
* Tada ida yuca anica Mille
Uncommon. Roos s in houses. Specimens om Paco a and Panama
(Panama) and Mi a lo es (Canal Zone).
Molossus bondae J. A. Allen
Common a lowe ele a ions in eas e n Panama and on he Ca ibbean
coas o wes e n Panama. Openings in e e g een o es . Roos s in houses.
Specimens om Taca cuna Village (Da ien) ; Escobal and Salud (Colon) ;
Chi a Chi a, Fo Kobbe, Fo She man, and Juan Mina (Canal Zone) ;
and Almi an e, 7 km. SSW. o Changuinola, and Sibube (Bocas del To o).
Molossus coibensis J. A. Allen
= Molossus majo Hall and Kelson (pa .)
Abundan a lowe ele a ions in cen al Panama and on he Paci ic
coas o wes e n Panama. Open coun y. Roos s in houses. Specimens
om Isla Taboga, La Cho e a, Paco a, and Panama (Panama) ; Ancon,
Balboa, Ba o Colo ado Island, Bohio, Chi a Chi a, Co ozal, Culeb a, Cu-
undu, Fo Clay on, Fo She man, Gamboa, Ga iin, Ma ga i a, Pa aiso,
San Pablo, Summi , and Tabe nilla (Canal Zone) ; Escobal, F ijoles, and
Salud (Colon) ; Isla Coiba and Isla Gobe nado a (Ve aguas) ; andBoque on
(Chi iqui). I is possible ha he e is mo e han one species o pygmy
Molossus. The e o e assignmen o M . coibensis o any pa icula one on he
basis o size alone is p ema u e.
* Molossus nig icans Mille
= Molossus u us Hall and Kelson
Uncommon. Roos s in a ics o houses. E. L. Tyson ook specimens a
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HANDLEY: CHECKLIST OF MAMMALS 773
Alanje, 50 ee , Boque on, 600 ee , 2 mi. W. o El Volcan, 4100 ee , and La
Concepcion, 800 ee (Chi iqui). M. a e E. Geo ey S .-Hilai e ( = M.
u us E. Geo ey S .-Hilai e), M. nig icans Mille , and M. p e iosus Mille
a e simila and u he s udy may show hem o be conspeci ic.
Molossus sinaloae J. A. Allen
Uncommon a lowe ele a ions in eas e n Panama and on he Ca ibbean
coas o wes e n Panama. Roos s in houses. Specimens om Boca de Cupe
and El Real (Da ien) ; Fo Amado and Fo Clay on (Canal Zone) ; and
7 km. SSW. o Changuinola and Pun a de Pena (Bocas del To o) . The Pana-
manian subspecies is M. s, ini a us Goodwin, a o m which he e o o e has
been conside ed o be a dis inc species con ined o he island o T inidad.
* P omops cen alis Thomas
Ra e. Roos s in houses. Specimens ha e been aken by Cha les M.
Keenan and Ve non J. Tip on a Co ozal, Fo Amado , and Fo Clay on
(Canal Zone). P omops cen alis Thomas, P. occul us Thomas, and P.
da idsoni Thomas may be conspeci ic. The oldes name is P. cen alis.
* Eumops amazonicus Handley
Ra e. Known in Panama by a single specimen om Taca cuna Village,
1950 ee (Da ien). Ne ed o e o es -bo de ed s eam.
Eumops au ipendulus Shaw
=i Eumops glaucinus Goldman
= Eumops ab asus Hall and Kelson
Uncommon. Roos s in houses. Specimens om F ijoles (Colon) ; Bo-
hio, Empi e, Fo Clay on, Fo Kobbe, Pa aiso, and Summi (Canal Zone) ;
and 7 km. SSW. o Changuinola (Bocas del To o). Fo nomencla u e, see
Goodwin (1960).
Eumops bona iensis Pe e s
= Eumops nanus Goldman
Ra e. Specimens om Boque on and Bugaba (Chi iqui). The Pana-
manian subspecies is E. b. nanus Mille .
* Eumops glaucinus Wagne
Ra e. Roos s in a ics o houses. E. L. Tyson ook specimens a Da id,
150 ee , and La Concepcion, 800 ee (Chi iqui).
Sibube (Bocas del To o). (2) S. b. inci a us Bangs, Isla del Rey. (3)
S. b. messo ius Goldman. Specimens om Boca de Cupe, Cana, 1800 ee ,
Jaque, Taca cuna Casi a, 1500 ee , Taca cuna Village, 1950 ee , and
Tapalisa (Da ien) . Specimens om Ce o Pun a, 5300 ee (Chi iqui) may
ep esen an undesc ibed subspecies.
O de RODENTIA
Family Sciu idae
Sciu us g ana ensis Humbold
= Sciu us ho manni Goldman
= Sciu us ge a di Goldman
Abundan . Th oughou Panama a all ele a ions. Pe haps no o he
Panamanian mammal is so wide- anging. In all so s o o es . A bo eal,
bu spends much ime on he g ound. Subspecies in Panama: (1) S. g.
[Begin Page: Page 777]
HANDLEY : CHECKLIST OF MAMMALS 777
chi iquensis Bangs, wes e n Panama, om 6 mi. E. o El Valle (Panama)
and Guanico (Los San os) wes in o Cos a Rica; (2) S. g. mo ulus Bangs
( = S. g. choco Goldman) , Canal Zone and eas e n Panama. The popula ion
ha has hi he o bo ne he p io name S. g. mo ulus is an in e g ade be ween
he small, plain-colo ed S. g. chi iquensis and he la ge, a icolo ed S, g.
choco, bu is mo e like he la e .
Sciu us a iega oides Ogilby
Abundan a lowe ele a ions in he Canal Zone and wes e n Panama.
F ui g o es, sc ub, and semi-open coun y. A bo eal. Panamanian sub-
species: (1) S. . hel eolus Goldman. Paci ic coas o cen al Panama.
Specimens om Chi a-Chi a, Cocoli, Co ozal, Empi e Range, Fo Clay on,
Paci ic, Pa aiso, and Summi (Canal Zone) ; A aijan, La Cho e a,
Nue o Empe ado , and Panama (=Calidonia) (Panama); Pa i a
(He e a) ; and San iago (Ve aguas). (2) S. . melania G ay. Paci ic
coas o wes e n Panama. Specimens om Guanico (Los San os) ; Pese
(He e a) ; Isla Cebaco, Rio Ma ia o, and Sona (Ve aguas) ; and Boque on,
Boque e, 2000 ee , Bugaba, Da id, Di ala, Isla B a a, Isla Insole a, Isla
Se illa, and Remedies (Chi iqui). (3) S. . homasi Nelson. Ca ibbean
coas o wes e n Panama. Specimens om Almi an e and 7 km. SSW. o
Changuinola (Bocas del To o).
Syn heosciu us b ochus Bangs
Ra e. Fog o es . Known only om he uppe Ca ibbean slope o
he Con inen al Di ide, no h o Boque e, 7000 ee (Bocas del To o).
Mic osciu us al a i J. A. Allen
Uncommon. P obably occu s h oughou he o es ed pa s o Panama.
A bo eal. Diu nal. Subspecies in Panama: (1) M. a. b owni Bangs.
Paci ic coas o wes e n Panama. Specimens om Bugaba, 600 ee , Que-
b ada San a Cla a, 3600-4200 ee , Rio Ga iche, 5300 ee , Se eno, 3600-
3700 ee , and Siolo, 4100-4300 ee (Chi iqui) . (2) M. a. usculus Thomas.
Ex eme sou heas e n Panama, sou h and wes o he Rio Tui a. Speci-
mens om Cana, 2000 ee , Ce o Sapo, and Rio Se egan i, 2600 ee
(Da ien). (3) M. a. enus ulus Goldman. Cen al Panama and he Ca ib-
bean coas o wes e n Panama. Specimens om Mandinga (San Bias) ;
Po obelo (Colon) ; Ce o Azul, 2000-2500 ee , and Rio Pequeni (Pana-
ma) ; Ba o Colo ado Island and Ga un (Canal Zone) ; and Almi an e and
uppe Rio Changena, 5000 ee (Bocas del To o). (4) Specimens om
A mila (San Bias) and Taca cuna Casi a, 1500 ee , and Taca cuna Village,
1950 ee (Da ien) , ep esen an undesc ibed subspecies.
Mic osciu us mimulus Thomas
= Mic osciu us boque ensis Goldman
= Mic osciu us boque ensis Hall and Kelson
= Mic osciu us is hmius Goldman
= Mic osciu us is hmius Hall and Kelson
[Begin Page: Page 778]
778 ECTOPARASITES OF PANAMA
Ra e. In og o es possibly h oughou Panama. Subspecies in Pan-
ama: (1) M.m. boque ensis Nenson. Ce o Mali, 4900 ee , and Taca cuna
Village, 1950 ee (Da ien) ; Ce o Azul, 2500-3000 ee (Panama) ; and
high idges abo e Boque e (Chi iqui and Bocas del To o). (2) M. m.
is hmius Nelson ( = M. m. i a us Goldman). Specimens om Cana, 3500
ee , 20 mi. S. o Cana, 5000 ee , Ce o Sapo, 3000 ee , and Rio Jaque
(Da ien).
Family Geomyidae
Mac ogeomys ca a o Bangs
= Mac ogeomys pansa Goldman
Locally common. Ex eme wes e n Panama. Fosso ial. Mos ly in
ields. Subspecies in Panama: (1) M. c. ca a o Bangs. Boque e, 4800
ee , and Ce o Pun a, 5300-7800 ee (Chi iqui). (2) M. c. pansa Bangs.
Bugaba, 600 ee (Chi iqui).
Mac ogeomys da iensis Goldman
Abundan in he lowe Rio Chucunaque and uppe Rio Tui a basins in
eas e n Panama. Fosso ial. Fields, ui g o es, and o es . Specimens
om Boca de Cupe, 250 ee , Boca de Rio Paya, 500 ee , Cana, 2000-2500
ee , Taca cuna egion, 1500-2500 ee , and Tapalisa (Da ien). Mounds
seen a Ya isa.
Family He e omyidae
Liomys adspe sus Pe e s
Abundan in he semi-a id sa annah coun y o he Paci ic coas o
wes e n Panama. Open ho ny sc ub and weedy ields. Te es ial. Speci-
mens om Ce o Azul, 1500 ee , and Chepo (Panama) ; Alb ook Field,
Balboa, Chi a Chi a, Cocoli, Cu undu, Empi e, Fa an, Fo Clay on, Fo
Kobbe, Madden Fo es , and Summi (Canal Zone) ; Guanico (Los San os) ;
and Pa aco e and San a Fe (Ve aguas) .
He e omys aus alis Thomas
Abundan . Lowe ele a ions in eas e n Panama. Fo es . Te es ial.
Subspecies in Panama: (1) H. a. conscius Goldman. Specimens om
Boca de Rio Paya, 500 ee , Cana, 1800-2000 ee , Rio Se egan i, 2600
ee , Taca cuna Casi a, 1500-1700 ee , and Taca cuna Village, 1950 ee
(Da ien). (2) H. a. paci icus Pea son. Known only om Amagal, 1000
ee , S. o Guayabo Bay (Da ien) .
He e omys desma es ianus G ay
Common a highe ele a ions h oughou , and o sea le el on he Ca ib-
bean coas and in Chi iqui (no specimens ha e been aken elsewhe e on he
Paci ic coas ). Te es ial. Fo es , usually e e g een. Subspecies in
Panama: (1) H, d, chi iquensis Ende s. High ele a ions in ex eme wes -
[Begin Page: Page 779]
HANDLEY : CHECKLIST OF MAMMALS 779
e n Panama. Specimens om Ce o Pando and Ce o Pun a, 6800-7800
ee (Chi iqui). (2) H. d. c assi os is Goldman. High ele a ions in ex-
eme eas e n Panama. Specimens om Ce o Mali (= "Moun Taca -
cuna"), 4900 ee , and Ce o Pi e, 4500-5000 ee (Da ien). (3) H. d.
panamensis Goldman. Medium ele a ions in cen al Panama. Specimens
om Ce o Azul, 2000-3000 ee (Panama) ; and Ce o B uja, 2000 ee
(Colon) . (4) H. d. epens Bangs. Lowe and middle ele a ions in wes e n
Panama. Specimens om Boque on and Boque e, 4000 ee (Chi iqui).
(5) H. d. zonalis Goldman. Lowe ele a ions in cen al Panama and on
he Ca ibbean coas o wes e n Panama. Specimens om Fo She man,
Ga un, Madden Dam, and Rio Indio (Canal Zone) ; Ce o Campana, 6 mi.
E. o El Valle, and Maxon Ranch, Rio T inidad (Panama) ; Salud (Colon) ;
and Almi an e, 7 km. SSW. o Changuinola, and uppe Rio Changena, 2300-
2600 ee (Bocas del To o) . (6) Specimens om he San Bias coas ep e-
sen an undesc ibed subspecies.
Family C ice idae
O yzomys albigula is Tomes
= O yzomys de ius Goldman
= O yzomys de ius Hall and Kelson
= O yzomys pi ensis Goldman
i= O yzomys pi ensis Hall and Kelson
Uncommon. High ele a ions in eas e n and wes e n Panama. Fog
o es . Te es ial. Subspecies in Panama: (1) O. a. de ius Bangs. Speci-
mens om Boque e, 4000-5000 ee , and Ce o Pun a, 6800-7800 ee
(Chi iqui) ; and uppe Rio Changena, 4800-5000 ee (Bocas del To o).
(2) 0. a. pi ensis Goldman. Specimens om Ce o Mali (="M . Taca -
cuna"), 4900 ee , and Ce o Pi e, 4500 ee (Da ien).
O yzomys al a oi J. A. Allen
Uncommon. Highe ele a ions in eas e n and wes e n Panama. E e -
g een o es . Te es ial. Subspecies in Panama: (1) O. a. al a oi J. A.
Allen. Specimens om Boque e, 4000 ee , Ce o Pun a, 5000-5800 ee ,
and El Volcan, 4200 ee (Chi iqui) ; and uppe Rio Changena, 2400 ee
(Bocas del To o). (2) O. a. da iensis Goldman. Specimens om Cana,
2000-2500 ee , Ce o Mali (="M . Taca cuna"), 4800 ee , Taca cuna
Casi a, 1500 ee , and Taca cuna Village, 1950 ee (Da ien).
* O yzomys bicolo Tomes
= O yzomys ende si Hall and Kelson
= O yzomys abea us Hall and Kels'on
Ra e. A lowe ele a ions in he Canal Zone and eas e n Panama. E e -
g een o es . A bo eal. Specimens om Guayabo, Pelisa, 500 ee , and
Rio Jesuci o (Da ien) ; A mila and Pue o Obaldia (San Bias) ; Ce o Azul,
2000 ee (Panama) ; and Ba o Colo ado Island and Fo She man (Canal
[Begin Page: Page 780]
780 ECTOPARASITES OF PANAMA
Zone) . The Panamanian subspecies is O. b. abea us G. M. Allen and Ba -
bou (=O yzomys ende si Goldman). Fo no es on he nomencla u e o
his species see He shko i z (1960, p. 518).
O yzomys bombycinus Goldman
Ra e. Middle ele a ion e e g een o es . Te es ial. Subspecies in
Panama: (1) O. b. alleni Goldman. Uppe Rio Changena, 2400 ee (Bocas
del To o). (2) O. b. bombycinus Goldman. Ce o Azul, 2000-3000 ee ,
and 6 mi. E. o El Valle (Panama) and Ce o B uja, 1000 ee (Colon).
(3) O. b. o inus Pea son. Ce o Pi e (Da ien).
O yzomys caliginosus Tomes
Abundan . Lowe ele a ions in eas e n Panama and on he Ca ibbean
coas o wes e n Panama (seemingly absen om mos o he Paci ic coas
o wes e n Panama). E e g een o es and clea ings. Subspecies in Pan-
ama : (1) O. c. ch ysomelas J. A. Allen. Wes e n Panama. Specimens om
Bugaba, 600 ee (Chi iqui) and Almi an e, Boca del D ago, 7 km. SSW. o
Changuinola, Isla Bas imen os, and uppe Rio Changena, 2400 ee , (Bocas
del To o). (2) O. c. idoneus Goldman. Cen al and eas e n Panama.
Specimens om Cana, 1800-2800 ee , El Real, Taca cuna Casi a, 1500-
1700 ee , Taca cuna Laguna, 3000 ee , and Taca cuna Village, 1950 ee
(Da ien) ; A mila and Mandinga (San Bias) ; Ce o Azul, 2000-2500 ee ,
Ce o Campana, 6 mi. E. o El Valle, and Rio T inidad (Panama) ; and
Empi e Range, Fo She man, and Ga un (Canal Zone).
O yzomys capi o Ol e s
= O yzomys alamancae Goldman
=. O yzomys alamancae Hall and Kelson
= O yzomys la iceps auc o um
Abundan . A lowe ele a ions, p obably h oughou Panama, bu no
eco ded on he Paci ic coas wes o he Azue o Peninsula. Fo es , bo h
e e g een and deciduous. Te es ial. Subspecies in Panama: (1) O. c.
ca ike i J. A. Allen. Ca ibbean coas o wes e n Panama. Specimens om
Almi an e and Boca del D ago (Bocas del To o). (2) O. c. alamancae J.
A. Allen (= O yzomys panamensis Thomas). Cen al and eas e n Pan-
ama. Specimens om Boca de Rio Paya, Cana, 2000 ee , Ci u o, Jaque,
Rio Se egan i, 2600 ee , Taca cuna Casi a, 1500 ee , Taca cuna Village,
3200 ee , and Tapalisa (Da ien) ; A mila and Mandinga (San Bias) ; Ce o
Azul, 2000 ee , Ce o Campana, 6 mi. E. o El Valle, and Rio T inidad (Pan-
ama) ; Ce o B uja (Colon) ; Fo She man, Ga un, Madden Road, and
Mohinga Valley (Canal Zone) ; and Ce o Hoya, 3000 ee (Los San os).
Fo use o he name capi o, see He shko i z (1960, p. 544) and Cab e a
(1961, p. 387).
O yzomys concolo Wagne
= O yzomys ec us Goldman
= O yzomys ec us Hall and Kelson
Uncommon. Lowe ele a ions possibly h oughou Panama. Semi-
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HANDLEY: CHECKLIST OF MAMMALS 781
a bo eal, in all g ass and he baceous g ow h in sa annas and o es open-
ings. Specimens om Cana, 2000 ee , and Taca cuna Village, 1950 ee
(Da ien) ; A mila and Mandinga (San Bias) ; Co ozal and Madden Road
(Canal Zone) ; and Bugaba, 800 ee (Chi iqui). The Panamanian sub-
species is O. c. ec us Thomas (= O. on alis Goldman) . Fo no es on he
nomencla u e o his species see He shko i z (1960, p. 515).
O yzomys ul escens Saussu e
Common. Con ined la gely o he Paci ic slope o cen al and wes e n
Panama. Sa annas and o es openings. Te es ial. Subspecies in Pan-
ama: (1) O. . cos a icensis J. A. Allen. Lowe ele a ions. Specimens om
Ce o Azul, 2000-3000 ee , Ce o Campana, 6 mi. E. o El Valle, La Cho -
e a, Paco a, and Panama Viejo (Panama) ; Alb ook Field, Ba o Colo ado
Island, Cu undu, Fo Clay on, and Fo Kobbe (Canal Zone) ; and Ce o
Pun a, 4800-5600 ee (Chi iqui). (2) O. /. ega us Bangs. Highe ele-
a ions, wes e n Panama. Specimens om Boque e, 3800-4800 ee , and
Ce o Pun a, 6800-7800 ee (Chi iqui) ; and uppe Rio Changena, 4800
ee (Bocas del To o).
O yzomys palus is Ha lan
=i O yzomys ga unensis Goldman
= O yzomys ga unensis Hall and Kelson
Sigmodon hispidus Say and O d
= Sigmodon hispidus Goldman
= Sigmodon aus e ulus Goldman
Abundan . Low ele a ions. Cen al Panama, he Paci ic coas o wes -
e n Panama, and he wes e n ex eme o he Ca ibbean coas . Sa annas
and o es openings. Te es ial. Specimens om he Canal Zone and he
Paci ic coas a e o he subspecies S. h. chi iquensis J. A. Allen : Ce o Azul,
1000 ee and Ce o Campana (Panama) ; many locali ies, h oughou he
Canal Zone; El Valle (Code) ; Ce o Hoya, 3000 ee , and Guanico (Los
San os) ; Rio Ma ia o and San a Fe (Ve aguas) ; and Boque on and Bugaba
(Chi iqui). Specimens om Almi an e and 7 km. SSW. o Changuinola
(Bocas del To o) ep esen an undesc ibed subspecies. The nominal
species "S. aus e u j us Bangs", known only by he holo ype, is now hough
a synonym o S. h. bo ucae J. A. Allen and o ha e come om Cos a Rica, no
he summi o Volcan de Chi iqui as labeled (Ende s, 1953, p. 508) .
* Rheomys ha manni Ende s
Ra e. Known only by wo specimens om he ho sp ings on he Rio
Co i o, 4900 ee (Chi iqui). Aqua ic.
Rheomys ap o Goldman
Ra e. Known only by h ee specimens om Ce o Pi e, 4500 ee
(Da ien). Aqua ic.
* Rheomys unde wood! Thomas
Ra e. One specimen om El Volcan (Chi iqui). Aqua ic.
Family Mu idae
* Ra us no egicus Be kenhou
Uncommon. U ban. Te es ial. Specimens om C is obal and Moun
Hope (Canal Zone). In oduced.
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HANDLEY : CHECKLIST OF MAMMALS 785
Ra us a us Linnaeus
Abundan . Now ound a ound human habi a ions and in some wild
a eas h oughou much, i no all, o Panama. Pa ly e es ial, pa ly
a bo eal. Th ee colo phases a e encoun e ed: black (= "R. . a us Lin-
naeus"), b own wi h g ay belly (="R. . alexand inus E. Geo oy S .-
Hilai e"), and b own wi h whi e belly ("R. . ugi o us Ra inesque").
In oduced.
Mus musculus Linnaeus
Uncommon. In human habi a ions. Specimens om El Valle (Code)
Isla del Rey, Las Cumb es, and Rio A aso (Panama) ; Fo Clay on (Canal
Zone) ; and Ce o Pun a, 5000 ee (Chi iqui). In oduced.
Family E e hizon idae
Coendou mexicanus Ke
= Coendou mexicanum Goldman
In wes e n Panama, om high ele a ions on he Paci ic slope (common)
o nea sea le el on he Ca ibbean coas ( a e). A bo eal. Specimens om
Boque e and Ce o Pun a, 5300-6200 ee (Chi iqui), and Sibube (Bocas
del To o). The Panamanian subspecies is C. m. laena us Thomas.
Coendou o hschildi Thomas
Uncommon. A lowe ele a ions, possibly h oughou Panama, excep on
he Ca ibbean coas o wes e n Panama. A bo eal. Specimens om Boca
de Rio Paya (Da ien) ; A mila (San Bias) ; Ba o Colo ado Island, Ca i a
Road, Fo Da is, Fo Kobbe, Fo She man, F ance Field, Ga un, Rio
Indio, and Tabe nilla (Canal Zone) ; Pa i a (He e a) ; Guanico (Los
San os) ; and Boque on, Isla B a a, and Isla Se illa (Chi iqui).
Family Hyd ochae idae
Hyd ochae is hyd ochae is Linnaeus
= Hyd ochoe us is hmius Goldman
= Hyd ochae is is hmius Hall and Kelson
Uncommon. Eas e n Panama. Semiaqua ic. S eam banks and
ma shes. Specimens o obse a ions om El Real, Ma agan i, and Rio
Se egan i, 2600 ee (Da ien) ; 15 mi. E. o Panama (Panama) ; and Juan
Mina (Canal Zone). The Panamanian subspecies is H. h. is hmius Gold-
man.
Family Dasyp oc idae
Agou i paca Linnaeus
= Cuniculus paca Goldman
Common h oughou Panama excep whe e i has been excessi ely
[Begin Page: Page 786]
786 ECTOPARASITES OF PANAMA
hun ed. Fo es . Te es ial. Specimens om Boca de Rio Paya, Cana,
2000 ee (obse a ion), and Taca cuna Village, 1950 ee (Da ien);
A mila and Mandinga (San Bias) ; Ce o Azul, 2000 ee (Panama) ; Ba o
Colo ado Island, Ga un, and Rio Indio (Canal Zone) ; Ce o Hoya, 2800-
3000 ee (Los San os) ; Ce o Pun a, 5300 ee , and Di ala (Chi iqui) ; and
Almi an e, Boca del D ago, Cayo Agua, Isla Bas imen os, and Sibube (Bocas
del To o). The Panamanian subspecies is A. p. i ga us Bangs.
Dasyp oc a coibae Thomas
Common. Known only on Isla Coiba. Fo es . Te es ial.
Dasyp oc a punc a a G ay
= Dasyp oc a callida Goldman
Common h oughou Panama. A all ele a ions, hough mo e common
in lowlands. Fo es . Te es ial. Subspecies in Panama: (1) D. p. bellula
Kellogg. Isla San Jose. (2) D. p. callida Bangs. Isla Ped o Gonzalez and
Isla del Rey. (3) D. p. da iensis Goldman. Eas e n Panama. Speci-
mens om A uza, Boca de Cupe, Boca de Rio Paya, Cana, Ce o Pi e, 5200
ee , Chepigana, Ci u o, El Real, and Rio Chucunaque (Da ien) ; A mila and
Mandinga (San Bias) ; and Ancon Hill (Canal Zone). (4) D. p. is hmica
Als on. Cen al Panama and a highe ele a ions in wes e n Panama.
Specimens om Ce o Azul, 2000 ee , and Rio T inidad (Panama) ; Colon
(Colon) ; Ba o Colo ado Island, Cocoli, Fo She man, Gamboa, Ga un,
Madden Dam, Pa aiso, and Rio Indio (Canal Zone) ; and Boque e (Chi iqui) .
(5) D, p. nuchalis Goldman. Paci ic lowlands o wes e n Panama, excep
he Azue o Peninsula. Specimens om Bugaba and Di ala (Chi iqui).
(6) D. p. pallidi en is Bole. Azue o Peninsula. Specimens om Ce o
Hoya, 2600-3000 ee , and Guanico (Los San os) ; and Isla Cebaco and
Pa aco e (Ve aguas). (7) D. p. ichmondi Goldman. Ca ibbean coas o
wes e n Panama. Specimens om Almi an e, 7 km. SSW. o Changuinola,
and Sibube.
Family Echimyidae
P oechimys semispinosus Tomes
Abundan h oughou he o es ed po ions o Panama a lowe ele-
a ions. Te es ial. Subspecies in Panama: (1) P. s. bu us Bangs. Isla
del Rey. (2) P. s. goldmani Bole. Azue o Peninsula. Specimens om
Ce o Viejo, Pa aco e and Rio Ma ia o (Ve aguas). (3) P. s. igno us
Kellogg. Isla San Jose. (4) P. s. panamensis Thomas (=P. s. chi iquinus
Thomas). Lowlands h oughou , excep on he Azue o Peninsula. Spec-
imens om Boca de Cupe, Boca de Rio Paya, Cana, 2000 ee , Ci u o, El
Real, Rio Chucunaque, Taca cuna Village, 1950 ee , and Tapalisa (Da ien) ;
A mila and Mandinga (San Bias) ; Ce o Azul, Chiman, and Rio T inidad
(Panama) ; many locali ies h oughou he Canal Zone; Boque on, Bugaba,
800 ee , and Di ala (Chi iqui) ; and Almi an e, Boca del D ago, 7 km.
SSW. o Changuinola, and Isla Bas imen os (Bocas del To o).
[Begin Page: Page 787]
HANDLEY: CHECKLIST OF MAMMALS 787
Hoplomys gymnu us Thomas
Locally uncommon o abundan . Eas e n Panama and he Ca ibbean
coas o wes e n Panama. E e g een o es . Te es ial. Subspecies in
Panama: (1) H. g. goe halsi Goldman. Specimens om Cana, 2000 ee ,
and Taca cuna Village, 1950 ee (Da ien) ; A mila and Mandinga (San
Bias) ; Ce o Azul, 2000-2100 ee (Panama) ; Fo She man, Ga un, and
Rio Indio (Canal Zone), and Almi an e, Boca del D ago, Cayo Agua, 7 km.
SSW. o Changuinola, Isla Bas imen os, Isla Colon, uppe Rio Changena,
2400 ee , and Sibube (Bocas del To o) . (2) H. g. we mo ei Handley. Isla
Escudo de Ve aguas (Bocas del To o) .
Diplomys labilis Bangs
= Diplomys da lingi Goldman
= Diplomys da lingi Hall and Kelson
A low ele a ions in he Canal Zone and eas e n Panama. A bo eal.
Nes s in ee holes. Subspecies in Panama: (1) D. I. da lingi Goldman.
Ra e. Specimens om Ce o Mali, 5000 ee , Ma agan i, and Tapalisa, 400
ee (Da ien) ; A mila and Mandinga (San Bias) ; and Ancon, Fo Kobbe,
and Juan Mina (Canal Zone) . (2) D.I. labilis Bangs. Common on Isla del
Rey (Panama).
O de CETACEA
Family Physe e idae
* Physe e ca odon Linnaeus
Ra e. De ini ely eco ded only o he Gul o Panama, bu p obably
occu s o bo h coas s. Ma ine.
Family Delphinidae
* S enella g a mani Lonnbe g
Common. Gul o Panama, Gul o Chi iqui, and p obably o he Paci ic
coas al wa e s. Ma ine.
* S enella longi os is G ay
Ra e, Paci ic. Specimen aken be ween Panama and Galapagos Islands.
Ma ine. Possibly mo e pelagic han S. g a mani.
* S enella plagiodon Cope
Possibly common. Gul o San Bias and p obably o he Ca ibbean
coas al wa e s. Ma ine.
*Tu siops nuuanu And ews
Paci ic. Specimen aken 70 mi. S. o Panama. Ma ine.
* Globicephala species?
On 29 Ma ch 1962, D . Alexande We mo e obse ed a g oup o small
whales o Isla Cebaco (Ve aguas) which he ook o be Globicephala.
[Begin Page: Page 788]
788 ECTOPARASITES OF PANAMA
Family Balaenop e idae
* Balaenop e a physalus Linnaeus
Possibly uncommon. Ca ibbean. Ma ine.
* Megap e a no aeangliae Bo owski
Uncommon. Paci ic. Obse ed in Gul o Panama. Ma ine.
*Sibbaldus musculus Linnaeus
Possibly a e. Ca ibbean. A dead specimen washed asho e a C is obal
(Canal Zone). Ma ine.
O de CARNIVORA
Family Canidae
* U ocyon cine eoa gen eus Sch ebe
Ra e. Canal Zone and Paci ic coas o wes e n Panama. Semi-a id
sa anna and sc ub coun y. Te es ial. Specimen om 3 mi. W. o Bal-
boa (Canal Zone). The Panamanian subspecies is U. c. u us G. M. Allen
and Ba bou .
Speo hos ena icus Lund
= Ic icyon panamensis Goldman
= Speo hos panamensis Hall and Kelson
Ra e. De ini ely known only in ex eme eas e n Panama, bu epo ed
om Canal Zone, Bocas del To o, and Chi iqui. Fo es . Te es ial. Spec-
imens om Ce o Pi e, 5000 ee (Da ien). The Panamanian subspecies
is S. . panamensis Goldman. Fo use o his name combina ion, see He sh-
ko i z (1957, p. 161).
Family P ocyonidae
Bassa iscus sumich as i Saussu e
Ra e. Known in Panama only by a specimen om Boque e, 6000 ee
(Chi iqui) . A bo eal. The Panamanian subspecies is B. s. no inus Thomas.
P ocyon canc i o us Cu ie
Uncommon. Eas e n Panama and he Ca ibbean coas o wes e n Pan-
ama. I has been epo ed on he Paci ic coas o Cos a Rica. Mos ly e -
es ial. Specimens om Boca de Rio Paya, Cana, 2000 ee , Rio Puc o,
and Taca cuna Village, 1950 ee (obse a ion only) (Da ien) ; A mila (ob-
se a ion only) and Mandinga (San Bias) ; Panama (Panama) ; Po obelo
(Colon) ; Ga un (Canal Zone) ; and 3.7 miles SE. o Almi an e (Bocas del
To o) . The Panamanian subspecies is P. c. panamensis Goldman.
P ocyon lo o Linnaeus
Locally common. Cen al and wes e n Panama. Mos ly a bo eal. Sub-
is T. m. mana us Linnaeus.
O de PERISSODACTYLA
Family Tapi idae
Tapi us bai dii Gill
= Tapi ella bai dii Goldman
Common a all ele a ions in eas e n Panama and on he Ca ibbean coas ,
bu now a e o absen on he Paci ic coas o wes e n Panama. La gely
[Begin Page: Page 792]
792 ECTOPARASITES OF PANAMA
e es ial, al hough o en equen ing s eams and lake sho es. Specimens
om Cana, 2000 ee , and Ce o Pi e, 5000 ee (Da ien) ; Mandinga (San
Bias) ; Ga un and Moun Hope (Canal Zone) ; Boque e, 5000 ee (Chi-
iqui) ; and uppe Rio Changena, 2400 ee (Bocas del To o) .
O de ARTIODACTYLA
Family Tayassuidae
Tayassu peca i Link
Uncommon, bu p obably dis ibu ed locally h oughou he o es ed
pa s o Panama. Te es ial. Specimens om Rio Changuinola (Bocas
del To o). On geog aphical g ounds he subspecies in Panama should be
T. p. spi adens (Goldman) . Fo no es on he use o he name Tayassu peca i
Link a he han Tayassu albi os is Illige as ecommended by Cab e a
(1961, p. 316) see He shko i z (1963, p. 85).
Tayassu ajacu Linnaeus
= Peca i angula us Goldman
Common h oughou Panama excep whe e i has been excessi ely
hun ed. Fo es and o es clea ings. Te es ial. Specimens om Boca
de Cupe, Boca de Rio Paya, El Real, and Taca cuna Village, 1950 ee
(Da ien) ; Ba o Colo ado Island, Gamboa, Ga un, and Madden Dam (Canal
Zone) ;Escobal (Colon) ; Pa aco e (Ve aguas) ; Boque e (Chi iqui) and Al-
mi an e, 7 km. SSW. o Changuinola, Rio Te ibe, and Sibube (Bocas del
To o) . The names T. . bangsi Goldman, Boca de Cupe (Da ien) , and T. .
c usnig um Bangs, Boque e (Chi iqui) , a e a ailable o Panamanian popu-
la ions.
Family Ce idae
Odocoileus i ginianus Zimme mann
= Odocoileus chi iquensis Goldman
=z Odocoileus o hschildi Goldman
= Dama i giniana Hall and Kelson
Uncommon. Paci ic coas o cen al and wes e n Panama, Canal Zone,
and wes e n ex eme o Ca ibbean coas . Appa en ly absen om he e-
mainde o he Ca ibbean coas and eas e n Panama. Te es ial. Usually
in hicke s in sa annas o in pa ches o wild cane along i e s, bu inhabi ing
dense o es on he Azue o Peninsula. Subspecies in Panama: (1) 0. .
chi iquensis J. A. Allen. Specimens om Rio Bayano (Panama) ; Co ozal
and Ga un (Canal Zone) ; Ce o Hoya, 2800 ee (Los San os) ; and Boque-
on and Boque e (Chi iqui). (2) 0. . o hschildi Thomas. Isla Coiba. (3)
O. . uei Me iam (= O. cos a icensis Mille ). Sibube (Bocas del To o).
Mazama ame icana E xleben
= Mazama sa o ii Goldman
Common. P obably h oughou he o es ed pa s o Panama. Te -
es ial. Subspecies in Panama : (1) M. a. ce asina Hollis e . Ca ibbean
[Begin Page: Page 793]
HANDLEY: CHECKLIST OF MAMMALS 793
coas o wes e n Panama. Specimens om Almi an e, 14 km. SSW. o
Changuinola, Rio Te ibe, and Sibube (Bocas del To o) . (2) M. a. epe icia
Goldman. Mainland Panama, excep he wes e n Ca ibbean coas . Speci-
mens om Boca de Cupe, Cana, Chepigana, Ci u o, El Real, Rio Chucu-
naque, Taca cuna Casi a, 1600 ee , Taca cuna Laguna, 3000 ee , Taca -
cuna Village, 3200 ee , and Tapalisa (Da ien) ; Maxon Ranch, Rio T inidad
(Panama) ; Ga un (Canal Zone) ; and.Boque e, 4000-4800 ee (Chi iqui).
* Mazama gouazoubi a Fische
Uncommon. Te es ial. Known in Panama only on Isla San Jose. The
Panamanian subspecies is M. g. pe mi a Kellogg.
[Begin Page: Page 794]
Re e ences ci ed
BURMEISTER, H.
1854. Sys ema ische Uebe sich de Thie e B asiliens. P . 1. Be lin, x + 342 pp.
BURT, W. H., and STIRTON, R. A.
1961. The mammals' o El Sal ado . Misc. Publ. Mus. Zool. Uni . Mich., no. 117,
69 pp., 2 igs.
CABRERA, A.
1958 & 1961. Ca alogo de los mami e os de Ame ica del Su . Re . Mus. A gen ine
Cien. Na . "Be na dino Ri ada ia," Cien. Zool., 4, nos. 1 and 2, pp. xxii + 732, 1 pi.
ENDERS, R. K.
1953. Is Sigmodon aus e ulus a alid species? Jou . Mamm., 34, pp. 508-509.
GOLDMAN, E. A.
1920. Mammals o Panama. Smi hs. Misc. Coll., 69, no. 5, 309 pp., 39 pis., 24 igs.,
map.
GOODWIN, G. G.
1958. Ba s o he genus Rhogeessa. Ame . Mus. No ., no. 1923, 17 pp.
1959. Ba s o he subgenus Na alus. Ame . Mus. No ., no. 1977, 22 pp., 2 igs.
1960. The s a us o Vespe ilio au ipendulus Shaw, 1800, and Molossus a e Geo ey,
1805. Ame . Mus. No ., no. 1994, 6 pp., 1 ig.
1963. Ame ican ba s o he genus Vampy essa, wi h he desc ip ion o a new species.
Ame . Mus. No ., no. 2125, 24 pp., 14 igs.
GOODWIN, G. G., AND GREENHALL, A. M.
1961. A e iew o he ba s o T inidad and Tobago. Bull. Ame . Mus. Na . His .,
122, a . 3, pp. 187-302, 113 igs., 40 pis., 2 maps.
1962. Two new ba s om T inidad, wi h commen s on he s a us o he genus
Mesophylla. Ame . Mus. No ., no. 2080, 18 pp.
HALL, E. R.
1960. O yzomys couesi only subspeci ically di e en om he ma sh ice a , O y-
zomys' palus is. Sou hwes e n Na u alis , 5, pp. 171-173.
HALL, E. R., and KELSON, K. R.
1959. The mammals o No h Ame ica. Ronald P ess, N. Y. 2 ols., 1373 pp., 1216
illus.
HAMLETT, G. W. D.
1939. Iden i y o Dasypus sep emcinc us Linnaeus wi h no es on some ela ed species.
Jou . Mamm., 20, pp. 328-336.
HANDLEY, C. 0., JR.
1960. Desc ip ions o new ba s om Panama. P oc. U. S. Na . Mus'., 112, pp.
459-479.
794
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HANDLEY: CHECKLIST OF MAMMALS 795
HERSHKOVITZ, P.
1957. A synopsis o he wild dogs o Colombia. No ed. Colombianas, Mus. His . Na .
Uni . del Cauca, no. 3, pp. 157-161.
1960. Mammals o no he n Colombia, p elimina y epo no. 8 : A bo eal ice a s,
a sys ema ic e ision o he subgenus Oecomys', genus O yzomys. P oc. U. S. Na .
Mus., 110, pp. 513-568, 6 igs., 12 pis.
1963. The nomencla u e o Sou h Ame ican pecca ies. P oc. Biol. Soc. Wash., 76,
pp. 85-88.
HUSSON, A. M.
1962. The ba s o Su iname. Zool. Ve h., Rijksmus. Van Na . His ., Leiden, no. 58,
282 pp., 39 igs'., 30 pis.
KOOPMAN, K. F.
1955. A new species' o Chilonyc e is om he Wes Indies and a discussion o he
mammals o La Gona e. Jou . Mamm., 36, pp. 109-113, 1 pi.
MILLER, G. S., JR., and KELLOGG, R.
1955. Lis o No h Ame ican ecsn mammals. U. S. Na . Mus. Bull. 205, xii +
954 pp.
SIMPSON, G. G.
1945. The p inciples o classi ica ion and a classi ica ion o mammals. Bull. Ame .
Mus. Na . His ., 85, x i + 350 pp.
WAGNER, J. A.
1855. Die Saug hie e in Abbildungen nach de Na u mi Besch eibungen. Suppl.
5, xx i + 810 pp.
