1
Rysza d Szadziewski1 , Ma a Gwizdalska-Ken ze 2
Fi s eco ds o Fo cipomyia dich oma a Remm (Dip e a:
Ce a opogonidae) in Poland and Spain
h ps://doi.o g/10.5281/zenodo.17310034
1
Uni e si y o Gdańsk, Depa men o In e eb a e Zoology and Pa asi ology, Wi a S wosza 59,
80-308 Gdańsk, Poland, e-mail: [email p o ec ed], ORCID: 0000-0002-2899-6861
2
Uni e si y o Gdańsk, Educa ion Quali y O ice, Jana Bażyńskiego 8, 80-309 Gdańsk, Poland,
e-mail: [email p o ec ed], ORCID: 0000-0003-3343-0608
Abs ac : Fo cipomyia (subgenus Fo cipomyia) dich oma a Remm, in Remm & Zhogole
1968 is eco ded o he i s ime in Poland and Spain. Ea lie , his Wes e n Palaea c ic
species was epo ed om Eu ope (Uk aine, F ance, No way), Middle Eas (Is ael, Tu key,
Yemen), and Cen al Asia (Tajikis an, Uzbekis an, A ghanis an). The male was diagnosed and
illus a ed. Cu en ly he numbe o bi ing midges (Ce a opogonidae) epo ed om Poland
inc eased o 221 and om Spain o 194 species.
Key wo ds: bi ing midge, No he n Poland, Sie a de Tejeda, aunis ics.
INTRODUCTION
Bi ing midges (Dip e a: Ce a opogonidae) a e small nema oce ous lies which a e
common in all kinds o aqua ic, semiaqua ic and mois e es ial habi a s wo ldwide. Thei
ecen wo ld auna is ela i ely well s udied and comp ises 6276 ex an species (Bo ken
& Dominiak 2020, Bo ken e al. 2022). Almos 600 alid species ha e been epo ed om
Eu ope (Szadziewski e al. 2013 ). The las checklis o bi ing midges o Poland includes
220 species (Szadziewski & Dominiak 2016) and o Spain 193 species (Delécolle 2002).
He e, he p esen ed sho no e p o ides i s eco ds o Fo cipomyia dich oma a Remm om
Poland and Spain.
MATERIAL AND SITES
NE Poland: Pome ania, T ojmiejski Landscape Pa k, Gdynia, Łęg nad Swelinią
ese e (Fig. 1), 54o27’41”N, 18o32’07”E, UTM CF43, 10.09.2011, a ligh , 2 males, leg.
M. Gwizdalska-Ken ze . SW Spain: Andalusia, Malaga p o ince, Sie a de Tejeda, illage
Puen a de don Manuel, 36o53’18”N, 4o08’47”W, 20.04.1984, o es , caugh by hand, 1 male,
leg. P. Su a. The examined specimens a e moun ed on slides in Canada balsam and deposi ed
in he collec ion o ex an in e eb a es a he Depa men o In e eb a e Zoology and
Pa asi ology, Uni e si y o Gdańsk.
ANNALS OF THE UPPER SILESIAN MUSEUM IN BYTOM
ENTOMOLOGY
Vol. 34 (online 009): 1–4 ISSN 0867-1966, eISSN 2544-039X (online) By om, 10.10.2025
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RESULTS AND DISCUSSION
Fo cipomyia (Fo cipomyia) dich oma a Remm, 1968
Fo cipomyia (subg. Fo cipomyia Meigen) dich oma a Remm, in Remm & Zhogole
1968: 826 (male, emale, C imea in Uk aine). Alwin e al. 2016: 363 ( eco ds om Is ael,
Yemen, Tu key); Bo ken & Dominiak 2020: 76 (in ca alogue); Thunes e al. 2021: 19
( eco d om No way); Na ai 2021: 238 ( emale, diagnosis, eco d om A ghanis an).
Fo cipomyia (subg. Mic ohelea Kie e ) dich oma a Remm. Remm 1988: 101 (subgene ic
placemen , eco ds om Uk aine, F ance, Tajikis an, Uzbekis an). Un o una ely, coun y
eco ds om Tajikis an and Uzbekis an a e “emp y eco ds” because hey ha e no de ailed
locali ies.
This mode a ely small species is a ypical membe o he subgenus Fo cipomyia s. s .:
wing leng h o male 1.6-1.7 mm, CR 0.4-0.5; wing memb ane co e ed wi h slende scale-like
mac o ichia; body b own, hume i yellowish, legs pale b ownish, geni alia da k b own; a sal
a io (TR) simila on all legs: TR(1) 0.7, TR(2) 0.6, TR(3) 0.6-0.7; male geni alia ypical o
he subgenus (Fig. 2C), aedeagus iangula (Fig. 2D).
Males o he species a e dis inc in ha ing b oadly used pa ame es, apices o pa ame es
poin ed, s ou (Fig. 2E); hi d palpal segmen wi h small senso y pi loca ed on swollen
base o he segmen (Fig. 2A, B). The species is close o F. uliginosa (Meigen) in ha ing
simila male geni alia and low a sal a ios. They di e mos ly in he shape o hi d palpal
segmen . In F. uliginosa senso y pi is loca ed on dis al hal o hi d palpal segmen while
in F. dich oma a i is loca ed on p oximal hal . Mo eo e , he base o used pa ame es in
F. dich oma a is almos W-shaped while in F. uliginosa U-shaped.
Fig. 1. Collec ing si e in Łęg nad Swelini ese e, N Poland (Ap il 2025).
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Bi ing midges o he subgenus Fo cipomyia s. s . ha e e es ial la ae eeding on ungi
g owing on o ing deb is o plan o igin, o example inside o s ems o oo s, in an nes s,
in cow o ho se dung, unde o ing ba k o ees (Szadziewski e al. 1997). Imma u e s ages
o Fo cipomyia dich oma a ha e no been epo ed and as a esul hei b eeding habi a is
unknown. The species has b oad dis ibu ion in Wes e n Palaea c ic, howe e i is a ely
epo ed. I was epo ed om Eu ope (Uk aine, F ance, No way), Middle Eas (Is ael,
Tu key, Yemen), and Cen al Asia (Tajikis an, Uzbekis an, A ghanis an). F om Poland and
Spain i is eco ded o he i s ime. Cu en ly he numbe o bi ing midges epo ed om
Poland (Szadziewski & Dominiak 2016) inc eased o 221 and om Spain (Delécolle
2002) o 194 species. In addi ion, he numbe o Ce a opogonidae eco ded om he Łęg nad
Swelinią ese e in N Poland (Gwizdalska-Ken ze 2011) inc eased o 50 species.
ACKNOWLEDGMENTS
We a e g a e ul o p o esso Pio Su a om K aków o collec ing in Spain in 1984
a male o Fo cipomyia dich oma a.
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Fig. 2. Male o Fo cipomyia dich oma a Remm. A, B – hi d palpal segmen , C – en al
aspec o geni alia, D – aedeagus, E – pa ame es.
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Accep ed: 27 July 2025; published: 10 Oc obe 2025
Licensed unde a C ea i e Commons A ibu ion License h p://c ea i ecommons.o g/licenses/by/4.0/
