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The type specimens of Keroplatidae and Mycetophilidae (Diptera: Sciaroidea) in the collection of the National Museum of Natural History, Sofia

Kazandzhieva, Svetlozara

Abstract

A catalogue of type specimens of fungus gnats (Diptera: Sciaroidea) in the collection of the National Museum of Natural History, Bulgarian Academy of Sciences, Sofia (NMNHS) is presented. Label information, specimen condition, and preservation details are provided. Primary types include 11 extant and one fossil Mycetophilidae species (Acnemia vratzatica Bechev, Anaclileia beshovskii Bechev, Chalastonepsia vumanhi Kazandzhieva, Ectrepesthoneura arturi Bechev, Ectrepesthoneura ledenikiensis Bechev, Greenomyia tomovi Bechev et Pavlova, Leia graeca Bechev, Mycetophila devioides Bechev, Polylepta meridionalis Bechev (junior syn. of P. zonata), Sciophila muglolutea Bechev et Koç, Sciophila turkolutea Bechev et Koç, Sciophila zaitzevi Bechev) and six Keroplatidae species (Macrocera gemagea Bechev, Macrorrhyncha thracica Bechev, Macrorrhyncha veleka Bechev, Monocentrota matilei Bechev, Orfelia gruevi Bechev, Urytalpa chandleri Bechev et Koç). Paratypes of five Mycetophilidae (A. vratzatica Bechev, A. beshovskii Bechev, Dziedzickia pectinata Ševčík, Bechev et Hippa, M. devioides Bechev, P. meridionalis Bechev) and three Keroplatidae species (Macrorrhyncha anatolica Bechev, M. matilei Bechev, O. gruevi Bechev) are also documented.

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Received: 1 September 2025; accepted: 30 September 2025 · Editor: Snejana Grozeva 347 Historia naturalis bulgarica 47 (11): 347–360 ISSN 2603-3186 (online) | ISSN 0205-3640 (print) · nmnhs.com/historia-naturalis-bulgarica https://doi.org/10.48027/hnb.47.112 Publication date [online]: 3 November 2025 Research article The type specimens of Keroplatidae and Mycetophilidae (Diptera: Sciaroidea) in the collection of the National Museum of Natural History, Sofia Svetlozara B. Kazandzhieva1, 2 (1) National Museum of Natural History, Bulgarian Academy of Sciences, 1 Tsar Osvoboditel Blvd, 1000 Sofia, Bulgaria (2) Regional Museum of Natural History – Plovdiv, 34 Hristo G. Danov Street, 4000 Plovdiv, Bulgaria, [email protected] ✉; https://orcid.org/0000-0001-5018-8951 � Abstract: A catalogue of type specimens of fungus gnats (Diptera: Sciaroidea) in the collection of the National Museum of Natural History, Bulgarian Academy of Sciences, Sofia (NMNHS) is presented. Label information, specimen condition, and preservation details are provided. Primary types include 11 extant and one fossil Mycetophilidae species (Acnemia vratzatica Bechev, Anaclileia beshovskii Bechev, Chalastonepsia vumanhi Kazandzhieva, †Ectrepesthoneura arturi Bechev, Ectrepesthoneura ledenikiensis Bechev, Greenomyia tomovi Bechev & Pavlova, Leia graeca Bechev, Mycetophila devioides Bechev, Polylepta meridionalis Bechev (junior syn. of P. zonata), Sciophila muglolutea Bechev & Koç, Sciophila turkolutea Bechev & Koç, Sciophila zaitzevi Bechev) and six Keroplatidae species (Macrocera gemagea Bechev, Macrorrhyncha thracica Bechev, Macrorrhyncha veleka Bechev, Monocentrota matilei Bechev, Orfelia gruevi Bechev, Urytalpa chandleri Bechev & Koç). Paratypes of five Mycetophilidae (A. vratzatica Bechev, A. beshovskii Bechev, Dziedzickia pectinata Ševčík, Bechev & Hippa, M. devioides Bechev, P. meridionalis Bechev) and three Keroplatidae species (Macrorrhyncha anatolica Bechev, M. matilei Bechev, O. gruevi Bechev) are also documented. Keywords: Bulgaria, fungus gnats, Greece, Oriental Region, Turkey, types Introduction Fungus gnats from the families Mycetophilidae and Keroplatidae (Diptera: Sciaroidea) are a highly diverse group and exhibit substantial morphological variation, yet their taxonomy and distribution remain incompletely known in the Balkans (Bechev, 1997a). This study examines holotypes of seventeen extant Mycetophilidae and Keroplatidae species – eleven from Bulgaria, three from Turkey, two from Greece, and one from Vietnam – along with a fossil species from Poland preserved in Baltic amber. In addition, paratypes of nine species are documented, including seven from Bulgaria, two from Turkey, and one from Thailand. The main part of the collection has been deposited in the National Museum of Natural History, Bulgarian Academy of Sciences, Sofia (NMNHS) by Dimitar Bechev in 2023. These specimens form an important reference for taxonomic verification and comparative studies. Aside from the type material documented in the present article, the NMNHS collections represent a valuable research resource, comprising more than 75,000 ethanol-preserved specimens of Keroplatidae and Mycetophilidae from Bulgaria collected during 2002–2004 and 2018–2019 (Pavlova, 2020). In Bulgaria, the only other substantial collection is at the Regional Natural History Museum – Plovdiv, comprising 308 Palaearctic species, type material of five Oriental species, and additional unsorted specimens (Kazandzhieva, 2025). The work of Dimitar Bechev has been significant in advancing knowledge of fungus gnats. By documenting and analysing these types, this study rein- Svetlozara Kazandzhieva 348 Historia naturalis bulgarica 47 (2025) forces the value of museum collections for future taxonomic research, species identification, and integration of molecular data. Material and methods Type specimens are listed alphabetically, with entries providing family, subfamily, tribe, genus, original combination, taxonomic status, original description, type locality, specimen details (including labels), and preservation notes. Label text is quoted in double quotation marks, specifying paper type and technique (handwritten or printed). Placement hierarchy (from top to bottom) of the label on the pin or vial is noted in round brackets. Individual lines of text are separated by “|”, while larger spaces created by one or more blank lines, or text written on opposite sides of the label, are indicated by “||”. All dry type specimens were photographed using an Olympus TG-7 camera in microscope mode. Figures were assembled in GIMP v3.04, with measurements and scale bars generated in FIJI (v1.54p; Schindelin et al., 2012). Each dry type specimen is illustrated; Fig. 1 shows the full dry type collection. Ethanol-preserved specimens are summarised in Fig. 2. The collection was digitised in the Specify database. Each specimen received a two-sided label: one side with the catalog number and QR code, the reverse reproducing the original label data. Catalog numbers are provided for all specimens. Results Family Keroplatidae Rondani, 1856 Subfamily Keroplatinae Rondani, 1856 Tribe Orfeliini Matile, 1990 Macrorrhyncha Winnertz, 1846 Macrorrhyncha anatolica Bechev, 2010 Bechev, 2010: 336 (original description). Type locality: Turkey, Aydýn, Bozdoğan. Paratype ♂ with labels: (1) printed (on white paper) in large vial “Macrorrhyncha anatolica | Bechev, PARATTYPE Fig. 1. Preservation of dry type specimens in the Entomological collection of NMNHS. Fig. 2. Preservation of ethanol type specimens in the Entomological collection of NMNHS. The type specimens of Keroplatidae and Mycetophilidae in the collection of the National Museum of Natural History, Sofia 349Historia naturalis bulgarica 47 (2025) [sic!], male” (2) printed (on white paper) in small vial “Macrorrhyncha anatolica | Bechev, Parattype [sic!], male” (3) printed (on white paper) “TURKEY: Aydın, Bozdoğan, Yaka | Village, 37°38' N/ 28°22' E, | 115 m a.s.l., 22.04.2005, leg. H. | Koç, A. Karaman & O. Ozgül”. Preservation: In ethanol, in two plastic vials, male genitalia in glycerol microvial within same vial as specimen. Catalog number: BG-NMNHS-ENT000000034435. Paratype ♀ with labels: (1) printed (on white paper) in large vial “Macrorrhyncha anatolica | Bechev, female | det. D. Bechev, 2008 | (handwritten) paratype || TURKEY: Aydın, Çine, Eskiçine | Village, Çine Stream, 37°31' N/ | 28°04' E, 80 m a.s.l., | 21.04.2005, leg. H. Koç, A. | Karaman & O. Ozgül” (2) handrwitten (on white paper) “paratype | female” (3) printed (on white paper) in small vial “Macrorrhyncha anatolica | Bechev, female | det. D. Bechev, 2008”. Preservation: In ethanol in two plastic vials, genitalia not detached. Catalog number: BGNMNHS-ENT-000000034436. Macrorrhyncha thracica Bechev, 1997 Described as Macrorrhyncha geranias thracica in Bechev, 1997b: 105. Raised to species level by Bechev, 2010: 340. Type locality: Greece, Trakia, NE of Sanda. Holotype ♂ with labels (1) printed (on red paper) in large vial “Diptera: Keroplatidae | Macrorrhyncha thracica Bechev, 1997 | HOLOTYPE ♂” (2) printed (on white paper) in small vial “Macrorrhyncha geranis [sic!] | thracica ssp. n. | det. Bechev, 1996 | Holotype, male (3) printed (on white paper) in small vial “Macrorrhyncha thracica Bechev, | 1997 stat. n. | det. Bechev, 2008” (4) printed (on white paper with black borders marked PRIRODOSLOVNI MUZEJ SLOVENJE) in small vial “GREECE: Trakia | NE of Sanda, Sapka Mts. | 41°07'06"N, 25°49'43"E | 24.5.1994 220 m | Leg. I. Sivec, B. Horvat”. Preservation: In ethanol, in two plastic vials, male genitalia in glycerol microvial within same vial as specimen. Catalog number: BGNMNHS-ENT-000000034437. Macrorrhyncha veleka Bechev, 1992 Bechev, 1992: 62 (original description). Type locality: Bulgaria, Sinemorets. Holotype ♂ (Fig. 3) with labels (1) printed (on carton paper) “BG Sinemoretz | 21.6.89 | D. Bechev” (2) printed (on white paper) “Macrorrhyncha | veleka Bechev | HOLOTYPE ♂”. Preservation: Mounted dry on a cardboard point; male genitalia in glycerol-filled microtube pinned between labels 1 and 2. Catalog number: BG-NMNHS-ENT-000000034467. Monocentrota Edwards, 1925 Monocentrota matilei Bechev, 1989 Bechev, 1989: 173 (original description). Type locality: Bulgaria, Osenovlag. Holotype ♂ with labels (1) printed (on pale red paper) in large vial “Bulgaria | W Balkan Range | v. Osenovlag | 6.08.1987 | leg. D. Bechev || Monocentrota | matilei Bechev | Holotype, male” (2) handwritten two-sided (on carton paper) folded in small vial “Monocentrota matilei | holotype” || “Bulgaria, West Balkan Range | v. Osenovlag, 6.08.87, D. Bechev”. Preservation: In ethanol, in two plastic vials, male genitalia in glycerol microvial within same vial as specimen. Catalog number: BG-NMNHS-ENT-000000034440. Paratype ♂ (Fig. 4) with labels (1) handwritten (on carton paper) “Bulgaria | Vratza | 10.7.1982 | D. Bechev” (2) / handwritten (on carton paper) “Monocentrota | matilei Bechev| paratype” (3) printed (on white paper) “Monocentrota matilei | Bechev,1989 | PARATYPE ♂”. Preservation: Mounted dry on a cardboard point, one severed leg, glued on the same card; male genitalia in glycerol-filled microvial pinned between labels 1 and 2. Catalog number: BG-NMNHS-ENT000000034469. Paratype ♂ (Fig. 5) with labels (1) Fig. 3 Habitus of Macrorrhyncha veleka. Holotype ♂ (BGNMNHS-ENT-000000034467). Scale bar 1 mm. Svetlozara Kazandzhieva 350 Historia naturalis bulgarica 47 (2025) handwritten (on carton paper) “Bulgaria | Vratza | 21.07.87 | D. Bechev” (2) / handwritten (on carton paper) “Monocentrota | matilei | paratype” (3) printed (on white paper) “Monocentrota matilei | Bechev,1989 | PARATYPE ♂”. Preservation: Mounted dry on a cardboard point; male genitalia in glycerol-filled microtube pinned above label 1. Catalog number: BG-NMNHS-ENT-000000034470. Orfelia Costa, 1857 Orfelia gruevi Bechev, 2002 Bechev, 2002: 199 (original description). Type locality: Bulgaria, Varna. Holotype ♂ (Fig. 6) with labels (1) handwritten (on carton paper) “BG, Varna | 9.VI.90 | D. Bechev” (2) printed (on white paper) “Orfelia gruevi Bechev | HOLOTYPE ♂”. Preservation: Mounted dry on a cardboard point; male genitalia in glycerol, within a concave transparent celluloid plate, pinned between labels 1 and 2. Catalog number: BG-NMNHS-ENT-000000034460. Paratype ♂ (Fig. 7 A) with labels (1) printed (on carton paper) “Skorpilovtzi | 8.VI.90 | D. Bechev” (2) “Orfelia gruevi Bechev | PARATYPE ♂”. Preservation: Mounted dry on a cardboard point; one leg and the male genitalia in glycerol within a concave transparent celluloid plate, pinned between labels 1 and 2. Catalog number: BG-NMNHS-ENT000000034461. Paratypes 5 ♂♂ (Fig. 7 B–F) with labels (1) handwritten (on carton paper) “BG, Varna | 9.VI.90 | D. Bechev” (2) printed (on white paper) “Orfelia gruevi Bechev | PARATYPE ♂”. Preservation: Mounted dry on a cardboard point; male genitalia in glycerol within a concave transparent celluloid plate, pinned between labels 1 and 2. Catalog numbers: BG-NMNHS-ENT-000000034462 (Fig.7 B); BG-NMNHS-ENT-000000034463 (Fig.7 C); BG-NMNHS-ENT-000000034464 (Fig. 7 D; BGNMNHS-ENT-000000034465 (Fig. 7 E); BGNMNHS-ENT-000000034466 (Fig. 7 F). Urytalpa Edwards, 1929 Urytalpa chandleri Bechev & Koç, 2008 Bechev & Koç, 2008: 31 (original description). Type locality: Turkey, Aydın, Koçarlı. Holotype ♂ with labels (1) printed (on red paper) in large vial Fig. 5. Habitus of Monocentrota matilei. Paratype ♂ (BGNMNHS-ENT-000000034470). Scale bar 1 mm. Fig. 6. Habitus of Orfelia gruevi. Holotype ♂ (BG-NMNHSENT-000000034460). Scale bar 1 mm. Fig. 4. Habitus of Monocentrota matilei. Paratype ♂ (BGNMNHS-ENT-000000034469). Scale bar 1 mm. The type specimens of Keroplatidae and Mycetophilidae in the collection of the National Museum of Natural History, Sofia 351Historia naturalis bulgarica 47 (2025) “Urytalpa chandleri Bechev & | Koç, 2008 Holotype, male | Turkey, Aydın, Koçarlı, Tekeli | Village 37°46' N / 27°38' E, 25 m | a.s.l, 09.04.2005, leg. H. Koç” (2) printed (on white paper) in small vial “Turkey, Aydın, Koçarlı, | Tekeli Village 37°46' N / 27°38' E | 25 m a.s.l, 09.04.2005, leg. H. Koç | Urytalpa chandleri Bechev & | Koç, 2008 Holotype, male”. Preservation: In ethanol, in two plastic vials, male genitalia in glycerol microvial within same vial as specimen. Catalog number: BG-NMNHS-ENT-000000034431. Fig. 7. Habitus of Orfelia gruevi. A – Paratype ♂ (BG-NMNHS-ENT-000000034461) B – Paratype ♂ (BG-NMNHS-ENT000000034462) C – Paratype ♂ (BG-NMNHS-ENT-000000034463) D – Paratype ♂ (BG-NMNHS-ENT-000000034464) E – Paratype ♂ (BG-NMNHS-ENT-000000034465) F – Paratype ♂ (BG-NMNHS-ENT-000000034466). Scale bars 1 mm. Svetlozara Kazandzhieva 352 Historia naturalis bulgarica 47 (2025) Subfamily Macrocerinae [Rondani, 1856] Tribe Macrocerini Rondani, 1856 Macrocera Meigen, 1803 Macrocera gemagea Bechev, 1991 Bechev, 1991: 185 (original description). Type locality: Bulgaria, Veslets. Holotype ♂ (Fig. 8) with labels (1) printed (on pale red paper) “Macrocera | gemagea Bechev | Holotype, male” (2) printed (on white paper) “Bulgaria | W Balkan Range | Vesletz | 20.07.1982 | leg. D. Bechev”. Preservation: Mounted dry on a cardboard point; male genitalia in glycerol within a concave celluloid plate, sealed with a transparent cover, pinned below label 2. Catalog number: BG-NMNHS-ENT-000000034468. Family Mycetophilidae Newman, 1834 Subfamily Gnoristinae Edwards, 1925 Dziedzickia Johannsen, 1909 Dziedzickia pectinata Ševčík, Bechev & Hippa, 2011 Ševčík, Bechev & Hippa, 2011: 691 (original description). Type locality: Thailand, Nakhon Nayok, Khao Yai NP. Paratype ♂ with labels (1) printed (on green paper) in large tube “Diptera: Mycetophilidae | Dziedzickia pectinata Ševčík, Bechev & | Hippa, 2011 PARATYPE ♂” (2) printed (on white paper in large vial “Thailand: Nakhon Nayok, Khao | Khao Yai N.P., Tao Tapoo, | 14.-24.5.2001” (3) printed (on white paper) in small vial “Dziedzickia pectinata | Ševčík, Bechev & Hippa, 2011 | Paratype, male” (4) printed (on white paper) in small vial “Thailand: Nakhon Nayok, Khao | Khao Yai N.P., Tao Tapoo, | 14.-24.5.2001”. Preservation: In ethanol, in two plastic vials, male genitalia in glycerol microvial within same vial as specimen. Catalog number: BGNMNHS-ENT-000000034433. Paratypes 4 ♂♂ with labels (1) printed (on green paper) in large tube “Diptera: Mycetophilidae | Dziedzickia pectinata Ševčík, Bechev & | Hippa, 2011 PARATYPE 4 ♂” (2) printed (on white paper in large vial “Thailand: Nakhon Nayok, Khao | Khao Yai N.P., Tao Tapoo, | 14.-24.5.2001” (3) printed (on white paper) in small vial “Dziedzickia pectinata | Ševčík, Bechev & Hippa, 2011 | Paratype, 4♂” (4) printed (on white paper) in small vial “Thailand: Nakhon Nayok, Khao | Khao Yai N.P., Tao Tapoo, | 14.-24.5.2001”. Preservation: In ethanol in two plastic vials; male genitalia not detached. Catalog number: BGNMNHS-ENT-000000034434. Subfamily Leiinae Edwards, 1925 Ectrepesthoneura Enderlein, 1910 Ectrepesthoneura ledenikiensis Bechev, 1988 Bechev, 1988a: 185 (original description). Holotype ♂ (Fig. 9) with labels (1) printed (on pale red paper) “Ectrepesthoneura | ledenikiensis | Bechev | Holotype, male” (2) “Bulgaria | W Balkan Range | Vrachanska Mts. | Ledenika | 5.04.1986 | leg. D. Bechev”. Preservation: Mounted dry on a cardboard Fig. 8. Habitus of Macrocera gemagea. Holotype ♂ (BGNMNHS-ENT-000000034468). Scale bar 1 mm. Fig. 9. Habitus of Ectrepesthoneura ledenikiensis. Holotype ♂ (BG-NMNHS-ENT-000000034449). Scale bar 1 mm. The type specimens of Keroplatidae and Mycetophilidae in the collection of the National Museum of Natural History, Sofia 353Historia naturalis bulgarica 47 (2025) Ectrepesthoneura arturi sp. n. from Baltic amber (Diptera: Mycetophilidae). Historia naturalis bulgarica, 47, 131-136” (2) printed (on white paper) “Ectrepesthoneura arturi sp. n. | Holotype ♂: Baltic amber inclusion”. Preservation: Piece of amber preserved in a plastic bag inside a plastic vial. Catalog number: BG-NMNHS-ENT-000000034430. Greenomyia Brunetti, 1912 Greenomyia tomovi Bechev & Pavlova, 2012 Bechev & Pavlova, 2012: 109 (original description). Type locality: Bulgaria, Eastern Rhodopes. Holotype ♂ with labels (1) printed (on red paper) in large vial “Diptera: Mycetophilidae | Greenomyia tomovi Bechev & Pavlova, | 2012, HOLOTYPE ♂” (2) printed (on red paper) in small vial “Bulgaria, East Rhodopes Mts., | near Madzharovo Town, | 14.10.2000” (3) printed (on red paper) in small vial “Greenomyia tomovi Bechev & | Pavlova, 2012, HOLOTYPE ♂”. Preservation: In ethanol, in two plastic vials, male genitalia in glycerol microvial within same vial as specimen. Catalog number: BGNMNHS-ENT-000000034432. Leia Meigen, 1818 Leia graeca Bechev, 1997 Bechev, 1997c: 179 (original description). Type locality: Greece, Farsala. Holotype ♂ (Fig. 11) with labels (1) printed (on carton paper) “GREECE | point; male genitalia in glycerol-filled microtube, pinned below label 2. Catalog number: BG-NMNHSENT-000000034449. †Ectrepesthoneura arturi Bechev, 2025 Bechev, 2025: 132 (original description). Type locality: Poland, Gdańsk district. Holotype ♂ (Fig. 10) with labels (1) printed (on white paper) “Ectrepesthoneura arturi sp. n. | urn:lsid:zoobank.org: act: | DF76F0ED-08B9-4124-A1D8-291175AAC268 | Type material. Holotype ♂: Baltic amber inclusion | preserved in 25×13×6 mm piece of amber collected | in the area that includes the villages of Mikoszewo, | Jantar, Stegna and Sztutowo, Gdańsk district, Poland, | leg. Artur Michalski, | in National Museum of Natural History, Sofia, | Bulgaria | Bechev, D. (2025) Fig. 10. Habitus of †Ectrepesthoneura arturi. A – Preservation in the collection of NMNHS. B. Amber inclusion with Holotype ♂ (BG-NMNHS-ENT000000034430). Scale bar 1 mm. Fig. 11. Habitus of Leia graeca. Holotype ♂ (BG-NMNHSENT-000000034448). Scale bar 1 mm. Svetlozara Kazandzhieva 354 Historia naturalis bulgarica 47 (2025) Farsala | 20.04.94 | D. Bechev” (2) handwritten (on carton paper) “Leia | graeca sp. n. | holotype” (3) printed (on white paper) “Leia | graeca Bechev | HOLOTYPE ♂”. Preservation: Mounted dry on a cardboard point; male genitalia in glycerol within a concave plastic plate, pinned below label 2. Catalog number: BG-NMNHS-ENT-000000034448. Subfamily Metanepsiinae Chalastonepsia Søli, 1996 Chalastonepsia vumanhi Kazandzhieva, 2025 Kazandzhieva, 2025 in Kazandzhieva, Langourov, Bechev & Van Nguyen, 2025: e153645 (original description). Type locality: Vietnam, Tam Dao NP. Holotype ♂ with labels (1) printed (on red paper) in large vial “Chalastonepsia vumanhi Kazandzhieva, 2025 | Holotype, ♂ | N. VIETNAM: Vĩnh Phúc Province: Tam Dao NP, Tam Đảo, ab| Chùa Vàng Tam Đảo, 1067 m, 21° 27.6348' N, 105° 38.9184' E, | 16 Oct 2023, Bekchiev, Simov, Langourov leg. (NMNHS)”. (2) printed (on white paper) in small vial with round red sticker “N. VIETNAM: Vĩnh Phúc Province: Tam Dao NP, Tam Đảo, ab| Chùa Vàng Tam Đảo, 1067 m, 21° 27.6348' N, 105° 38.9184' E, | 16 Oct 2023, Bekchiev, Simov, Langourov leg. (NMNHS)” (3) printed (on white paper) “Chalastonepsia vumanhi Kazandzhieva, 2025 | Holotype, ♂” (4) printed (on white paper) in second small vial “Chalastonepsia vumanhi Kazandzhieva, 2025 | holotype♂ Legs”. Note: The specimen is damaged, with several legs detached. These are preserved in a separate microvial and comprise: one fore leg (femur, tibia and tarsus), two mid legs (one partly attached to the specimen but lacking distal tarsomeres), and two hind legs (one lacking tarsus). Preservation: In ethanol in three plastic vials; male genitalia in glycerol microvial within same vial as specimen. Catalog number: BG-NMNHS-ENT000000019865. Subfamily Mycetophilinae Newman, 1834 Tribe Mycetophilini Newman, 1834 Mycetophila Meigen, 1803 Mycetophila devioides Bechev, 1988 Bechev, 1988a: 185 (original description). Type locality: Bulgaria, Balkan Mountains. Holotype ♂ (Fig. 12) with labels (1) printed (on pale red paper) “Mycetophila | devioides Bechev | Holotype, male” (2) printed (on white paper) “Bulgaria | W Balkan Range | h. Bjalata voda | 26.6.1982 | leg. D. Bechev”. Preservation: Mounted dry on a cardboard point; male genitalia in glycerol-filled microtube, pinned below label 2. Catalog number: BG-NMNHS-ENT000000034451. Paratypes 2 ♂♂ (Figs 13–14) with labels (1) handwritten (on carton paper) “Bulgaria | h. B. voda | 26.6.1982 | D. Bechev” (2) “Mycetophila | devioides Bech. | paratypus”. Preservation: Mounted dry on a cardboard point; male genitalia not detached. Catalog numbers: BG-NMNHS-ENT000000034453 (Fig. 13); BG-NMNHS-ENT000000034454 (Fig. 14). Paratype ♂ (Fig. 15) with labels (1) handwritten (on carton paper) “Bulgaria | Fig. 12. Habitus of Mycetophila devioides. Holotype ♂ (BGNMNHS-ENT-000000034451). Scale bar 1 mm. Fig. 13. Habitus of Mycetophila devioides. Paratype ♂ (BGNMNHS-ENT-000000034453). Scale bar 1 mm. The type specimens of Keroplatidae and Mycetophilidae in the collection of the National Museum of Natural History, Sofia 355Historia naturalis bulgarica 47 (2025) h. B. voda | 26.6.1982 | D. Bechev” (2) “Mycetophila | devioides Bech. | paratypus”. Preservation: Mounted dry on a cardboard point; male genitalia in glycerol within a concave transparent celluloid plate, pinned between labels 1 and 2. Catalog number: BGNMNHS-ENT-000000034452. Paratype ♂ (Fig. 16) with labels (1) handwritten (on carton paper) “Bulgaria | h. B. voda | 10.7.1982 | D. Bechev” (2) handwritten (on white paper) “paratype” (3) handwritten (on carton paper) “Mycetophila ♂ | devioides | Bechev”. Preservation: Mounted dry on a cardboard point; male genitalia in glycerol-filled microtube pinned between labels 1 and 2. Catalog number: BG-NMNHS-ENT-000000034455. Polylepta meridionalis Bechev, 1990 Bechev, 1990: 182 (original description), junior synonym of P. zonata (Kurina, 2003: 94). Type locality: Bulgaria, Strandzha. Holotype ♂ (Fig. 17) with labels (1) printed (on carton paper) “Bulgaria | Strandzha mts. | Silkosia res. | 29.05.87 | D. Bechev” (2) printed (on pale red paper) “Polylepta | meridionalis Bechev | Holotype, male” (3) “Bulgaria | Strandzha Mts. | Silkosia res. | 29.5.1987 | leg. D. Bechev”. Preservation: Mounted dry on a cardboard point; male genitalia in glycerol-filled microtube pinned between labels 1 and 2. Catalog number: BGNMNHS-ENT-000000034457. Paratype ♀ (Fig. 18) with labels (1) printed (on carton paper) “BG Slavyanka | Goleshovo 5.6.88 | D. Bechev” (2) printed (printed on green paper) “Polylepta | meridionalis Bechev | Paratype, female”. Preservation: Mounted dry on a cardboard point; female genitalia in glycerol-filled microtube pinned between labels 1 and 2. Catalog number: BGNMNHS-ENT-000000034458. Paratype ♀ (Fig. 19) Fig. 14. Habitus of Mycetophila devioides. Paratype ♂ (BGNMNHS-ENT-000000034454). Scale bar 1 mm. Fig. 15. Habitus of Mycetophila devioides. Paratype ♂ (BGNMNHS-ENT-000000034452). Scale bar 1 mm. Fig. 16. Habitus of Mycetophila devioides. Paratype ♂ (BGNMNHS-ENT-000000034455). Scale bar 1 mm. Fig. 17. Habitus of Polylepta meridionalis. Holotype ♂ (BGNMNHS-ENT-000000034457). Scale bar 1 mm.