2019 ● 41 ● 14
adansonia
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cou e u e / co E :
Lec o ype o Calendula g amini olia L. (Commelin 1701: 67, . 34).
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183
ADANSONIA, sé . 3 • 2019 • 41 (14)
© Publica ions scien i iques du Muséum na ional d’His oi e na u elle, Pa is. www.adansonia.com
KEY WORDS
Sou h A ica,
An hemideae,
A c o is,
Calenduleae,
Poi e ,
new lec o ypes,
new neo ype,
new synonym.
McKenzie R. J. & Nepi C. 2019. — Taxonomic eassessmen and ypi ica ion o species names in A c o is L. (As e aceae)
p oposed by Jean Louis Ma ie Poi e . Adansonia, sé . 3, 41 (14): 183-192. h ps://doi.o g/10.5252/adansonia2019 41a14.
h p://adansonia.com/41/14
ABSTRACT
The names A c o is s a ici olia Poi . and A c o is enui olia Poi . we e alidly published by Jean Louis
Ma ie Poi e based on ma e ial collec ed om “Cap de Bonne-Espé ance” and s a ed o be lodged
in he He bie Des on aines. No o iginal ma e ial o A c o is s a ici olia has been aced. The p o-
ologue is inconsis en wi h any known A c o is L. species and p e ious ea men o A. s a ici olia
as a synonym o Gazania linea is (Thunb.) D uce a . linea is is no suppo ed. The p o ologue o
A c o is s a ici olia is mos consis en wi h a na ow-lea ed axon o Dimo pho heca Vaill. ex Moench
and speci ically D. nudicaulis (L.) DC. a . g amini olia (L.) Ha . A neo ype o A c o is s a ici olia
is designa ed consis en wi h ou in e p e a ion and he name is o mally placed in he synonymy o
Dimo pho heca nudicaulis a . g amini olia. A c o is enui olia Poi . has p e iously been ea ed as a
synonym o Dimo pho heca nudicaulis a . g amini olia. O iginal ma e ial was loca ed and he name
A c o is enui olia Poi . is he e lec o ypi ied and is placed in he synonymy o U sinia enui olia (L.)
Poi . subsp. enui olia. A c o is enui olia Poi . is a la e he e o ypic homonym o A c o is enui olia L.
and is he e o e nomencla u ally illegi ima e. A lec o ype is designa ed o Sphenogyne enui olia (L.)
DC. a . he e och oma Ha ., a synonym o U sinia enui olia subsp. enui olia.
Robe J. MCKENZIE
Depa men o Plan Sciences and A omon ane Resea ch Uni ,
Uni e si y o he F ee S a e, Phu hadi jhaba (Sou h A ica)
h p://o cid.o g/0000-0001-7054-3308
[email p o ec ed]
Chia a NEPI
Museo di S o ia Na u ale-Bo anica, Uni e si à di Fi enze,
Via La Pi a 4, 50121, Fi enze (I aly)
[email p o ec ed]
Submi ed on 5 Feb ua y 2019 | accep ed on 24 June 2019 | published on 21 No embe 2019
Taxonomic eassessmen and ypi ica ion
o species names in A c o is L. (As e aceae)
p oposed by Jean Louis Ma ie Poi e
184 ADANSONIA, sé . 3 • 2019 • 41 (14)
McKenzie R. J. & Nepi C.
INTRODUCTION
The genus A c o is L. was es ablished by Linnaeus (1753). In
subsequen gene ic ea men s by Linnaeus and o he au ho s,
di e se species we e desc ibed as membe s o A c o is. Wi h
e inemen o he genus concep and desc ip ion o addi-
ional gene a, nume ous names in A c o is we e subsequen ly
de e mined o apply o elemen s o o he gene a, including
A c o heca J.C.Wendl., Cymbono us Cass. and Haploca pha
Less. (membe s o he ibe A c o ideae, sub ibe A c o idinae
as o A c o is), and also Dimo pho heca Vaill. ex Moench ( ibe
Calenduleae), Gazania Gae n. ( ibe A c o ideae, sub ibe
Go e iinae), and U sinia Gae n. ( ibe An hemideae). Fo a
numbe o hese axa, he epi he o he name in A c o is is
he ea lies legi ima e epi he o he axon a species ank.
As pa o an ongoing axonomic and nomencla u al e i-
sion o he sub ibe A c o idinae, he iden i y o all alidly
published names applicable o axa wi hin he sub ibe e-
qui es eassessmen . Conside able p og ess has been made in
esol ing he species axonomy o A c o idinae (e.g., McKenzie
e al. 2008a, b, 2011; McKenzie & Ba ke 2010, 2013; Mc-
Kenzie 2018, 2019; McKenzie & Be gh 2018). Reapp aisal
o A c o is names known o apply o axa now classi ied in
Dimo pho heca, Gazania and U sinia is no a p ima y ocus o
he cu en e ision o A c o idinae, as he la e h ee gene a
ha e been ho oughly e ised p e iously (No lindh 1943;
Roessle 1959; P assle 1967). Howe e , du ing he cou se
o he p esen e ision, a numbe o alidly published names
in A c o is o p e iously unce ain applica ion ha e been de-
e mined o apply o axa classi ied ou side he A c o idinae.
In he p esen s udy, he iden i y and axonomic s a us o
he names A c o is s a ici olia Poi . and A c o is enui olia Poi .
alidly published by Poi e (1810) we e assessed.
A c o is s a ici olia and A. enui olia Poi . we e based on
ma e ial s a ed o ha e been collec ed om “Cap de Bonne-
MOTS CLÉS
A ique du Sud,
An hemideae,
A c o is,
Calenduleae,
Poi e ,
lec o ypes nou eaux,
néo ype nou eau,
synonyme nou eau.
RÉSUMÉ
Réé alua ion axonomique e ypi ica ion des noms d’espèces chez A c o is L. (As e aceae) p oposés pa
Jean Louis Ma ie Poi e .
Les noms A c o is s a ici olia Poi . e A c o is enui olia Poi . on é é publiés alidemen pa Jean Louis
Ma ie Poi e , à pa i de ma é iaux assemblés au « Cap de Bonne-Espé ance » e décla és ê e p é-
sen s dans l’He bie Des on aines. Aucun ma é iel o iginal n’a cependan é é e ou é pou A c o is
s a ici olia. Le p o ologue es incompa ible a ec ou e espèce connue d’A c o is L. e le ai emen
an é ieu de A. s a ici olia comme synonyme de Gazania linea is (Thunb.) D uce a . linea is n’es pas
alide. Le p o ologue de A c o is s a ici olia es plu ô compa ible a ec un axon à euilles é oi es el
que Dimo pho heca Vaill. ex Moench e plus spéci iquemen D. nudicaulis (L.) DC. a . g amini olia
(L.) Ha . Un néo ype pou A c o is s a ici olia es désigné, con o mémen à no e in e p é a ion, e
le nom es o iciellemen placé en synonymie a ec Dimo pho heca nudicaulis a . g amini olia. A c-
o is enui olia Poi . é ai aupa a an considé é comme un synonyme de Dimo pho heca nudicaulis
a . g amini olia. Le ma é iel o iginal a é é ou é, le nom A c o is enui olia Poi . es ici lec o ypi ié
e considé é comme un synonyme de U sinia enui olia (L.) Poi . subsp. enui olia. A c o is enui olia
Poi . es un homonyme hé é o ypique ul é ieu de A c o is enui olia L. e es donc illégi ime nomen-
cla u alemen . Un lec o ype es désigné pou Sphenogyne enui olia (L.) DC. a . he e och oma Ha .,
synonyme d’U sinia enui olia subsp. enui olia.
Espé ance” [Cape o Good Hope] and lodged in he He bie
Des on aines (Poi e 1810). A c o is enui olia Poi . was in-
cluded in he synonymy o Cas alis nudicaulis a . g amini-
olia (L.) Ha . by No lindh (1943), bu he name has no
been lec o ypi ied. No lindh s a ed ha he did no examine
o iginal ma e ial o he name and his ea men o A c o is
enui olia Poi . ollowed ha o Candolle (1838). The name
A c o is s a ici olia was lis ed unde Gazania ’Species dubia’
by Roessle (1959), and emains un esol ed and un ypi ied,
bu is o en lis ed as a synonym o Gazania linea is (Thunb.)
D uce a . linea is in online nomencla o da abases. Typi ica-
ion and cla i ica ion o he applica ion o alidly published
names is essen ial o he ea men o As e aceae in he e-
Flo a o Sou h A ica p ojec (Le Roux e al. 2017). The e o e,
he objec i e o he p esen s udy was o loca e and examine
o iginal ma e ial and eassess applica ion o he names A c o is
s a ici olia and A. enui olia Poi .
MATERIAL AND METHODS
O iginal ma e ial o A c o is enui olia Poi . lodged in FI was
loca ed and examined. High- esolu ion digi al images o he
lec o ype o A c o is enui olia L. in LINN and specimens
o Dimo pho heca nudicaulis a . g amini olia in P we e ex-
amined. Rele an ma e ial lodged in NBG was examined.
He ba ium ac onyms ollow Thie s (2019). All ele an li -
e a u e was consul ed.
RESULTS AND DISCUSSION
A c o is s A ici oliA Poi .
Poi e (1810) published he name ’A c o is s a ice olia’. The
wo d ’s a ice’, a genus name published by Linnaeus (1753)
bu now in use only as a e nacula name o species o Li-
185
Typi ica ion o A c o is species o Poi e
ADANSONIA, sé . 3 • 2019 • 41 (14)
ig. 1. — Lec o ype o Calendula g amini olia L. (Commelin 1701: 67, . 34). Image om he Biodi e si y He i age Lib a y. Digi ized by Missou i Bo anical Ga den,
Pe e H. Ra en Lib a y (h ps://www.biodi e si ylib a y.o g).
186 ADANSONIA, sé . 3 • 2019 • 41 (14)
McKenzie R. J. & Nepi C.
monium Mill. (Plumbaginaceae), is de i ed om he G eek
s a ikē, eminine o στατικός (s a ikós), meaning ’s a ic,
causing o s and s ill, as ingen ’. Unde he In e na ional
Code o Nomencla u e o algae, ungi, and plan s (ICN;
Tu land e al. 2018) A . 60.10, co ec compounding o
he elemen s ’s a ice’ and ’ olia’ equi es emo al o he case
ending o he ansc ibed G eek -ós and addi ion o he
connec ing owel -i-. Unde he p o isions o ICN A s.
60.10 and 61.4, he epi he ’s a ice olia’ is co ec able o
’s a ici olia’ wi hou a change in he a ibu ed au ho ship
o he name.
Poi e (1810) s a ed ha he species was desc ibed om
ma e ial in he He bie Des on aines collec ed om Cap
de Bonne-Espé ance. No specimen o me ly in he gene al
he ba ium o Des on aines, no ma e ial ha can be linked
o Des on aines in o he he ba ia, and anno a ed wi h he
name “A c o is s a ice olia” has been aced in BM, B-W, C,
FI, LIV, MPU o P (based on he he ba ia holding specimens
linked o Des on aines lis ed by S a leu & Cowan 1976).
Fu he mo e, no shee o any o igin anno a ed wi h he
name has been aced in any he ba ium. In he absence o
o iginal ma e ial, he a ini ies o A c o is s a ici olia mus
be de e mined om he p o ologue o he species.
Poi e (1810) p o ided he diagnosis “A c o is acaulis,
oliis caespi osis, angus issimis, subin eg is, glab is; oliolis
calicinis linea i-lanceola is; lo e soli a io” accompanied by
a desc ip ion o he g ow h habi , lea mo phology, in o-
luc e and lo e colou a ion o A. s a ici olia. A diagnos ic
cha ac e o A c o is is ha he cypselae bea wo o h ee
( a ely i e) do sal wings ha a e used a he dis al and
p oximal ends o c ea e wo o h ee dis inc ’ca i ies’ o
u ows on he do sal side o he cypsela. Un o una ely,
Poi e was unable o examine cypselae o A c o is s a ici o-
lia, bu om he desc ip ion o he in oluc e A. s a ici olia
can be discoun ed as a species o A c o is. The in oluc e o
A c o is is mul ise ia e and he indi idual b ac s s ongly
di e in leng h om he sho es ou e mos b ac s o he
longes inne b ac s, and in shape om iangula -o a e
ou e b ac s o oblong-obo a e inne b ac s. The ou e in-
oluc al b ac s o en ha e a sho linea o obo a e apical
appendage and he inne b ac s ha e a pape y, anslucen
o ed-pigmen ed apex. The b ac s may be glab ous, glan-
dula o spa sely o densely omen ose. Poi e ’s desc ip ion
o he in oluc e o A c o is s a ici olia (“Ce e plan e es
ema quable pa son calice, composé d’un g and nomb e
de olioles é oi es, oblongues, linéai es-lancéolées, aiguës,
légè emen pileuses”) e e s o he in oluc al b ac s as many
in numbe , na ow, oblong, and linea -lanceola e in shape,
wi h an acu e apex, and sligh ly hai y. This desc ip ion
is incompa ible wi h he in oluc e o an A c o is species.
Applica ion o he name A c o is s a ici olia by p e ious
au ho s has a ied. Candolle (1838) lis ed he name A c o is
s a ici olia as a synonym o Gazania subula a R.B . wi hou
commen ing on he eason o he decision. In he mos
ecen e ision o Gazania, Roessle (1959) lis ed he name
unde Gazania ’Species dubiae’. Roessle no ed Candolle’s
ea men o A c o is s a ici olia and s a ed ha he was un-
able o examine he ype and he e o e could no assess he
a ini ies o he species. Roessle classi ied Gazania subula a
as a synonym o G. linea is a . linea is. A c o is s a ici olia
was no men ioned in ea men s o he As e aceae by Lessing
(1832) o Ha ey (1865), no in he mos ecen mono-
g aphs o A c o is (Beau e d 1915; Lewin 1922). Online
plan axonomic da abases (e.g., Ca alogue o Li e, h p://
www.ca alogueo li e.o g/; Global Biodi e si y In o ma ion
Facili y [GBIF], h ps://www.gbi .o g; Global Composi ae
Checklis , h ps://composi ae.landca e esea ch.co.nz; The
Plan Lis , h p://www. heplan lis .o g) p esen ly lis A c o is
s a ici olia as a synonym o Gazania linea is a . linea is.
Poi e ’s desc ip ion o he g ow h habi and he lea mo -
phology and a angemen a e no inconsis en wi h hose o
Gazania linea is a . linea is. The lea es o A. s a ici olia we e
desc ibed as nume ous, ga he ed a he u and all adical,
e y na ow, 5-7.5 cm o mo e long, sca cely b oadened
along he longi udinal axis, ob use, en i e, glab ous, en i e
bu some imes he ma gin was u nished wi h a ew small
dis an ee h. Howe e , Poi e desc ibed he lo e colou a-
ion as “Les leu ons du cen e son d’un jaune-pâle, de
couleu b une à leu somme ; les demi- leu ons lancéolés,
ob us, à ois den s, blancs en deho s, a ec une ein e pu -
pu ine e s leu base, d’un jaune-clai en dessous” [The disc
lo e s o he cen e a e o a pale yellow, he colou b own
a hei summi ; he ay lo e s lanceola e, ob use, wi h
h ee ee h, whi e on ex e io , wi h a pu plish hue owa ds
hei base, ligh yellow unde nea h]. In Gazania linea is
he disc- lo e co olla and uppe su ace o he ay- lo e
limb a e uni o mly and consis en ly b igh yellow. Among
o he Gazania species, in G. ju inei olia DC. he ay- lo e
limb is whi e abo e bu s eaked blackish b own below,
and he disc lo e co olla is yellow bu wi h o wi hou
whi e apices o he lobes. In addi ion, he lea es o Gaza-
nia ju inei olia do no con o m o Poi e ’s desc ip ion o
he lea es o A c o is s a ici olia. I Poi e was desc ibing
a specimen o a Gazania species, i is ema kable ha he
would no no e and commen on he usion o he base o
he in oluc al b ac s o o m a conna e ’cup’ o ’cylinde ’,
which is such a dis inc i e and conspicuous cha ac e is ic
o Gazania. No ably, in a subsequen b ie discussion o he
genus Gazania, Poi e (1811: 712) desc ibed he in oluc e
o ha genus as “Un calice d’une seule pièce, en u à sa
base, en i onné d’écaillés imb iquées, aiguës”. The e o e,
Poi e ’s desc ip ion o A c o is s a ici olia is incompa ible
wi h a Gazania species and conside a ion o he name as
a synonym o G. linea is is no suppo ed. Simila ly, all
o he ecognised gene a in he ibe A c o ideae can be dis-
coun ed owing o incompa ibili y o he in oluc e, lo e
colou a ion and ege a i e cha ac e s wi h he desc ip ion
o A c o is s a ici olia.
Nume ous As e aceae species indigenous o he Cape
egion o Sou h A ica p oduce capi ula wi h whi e ay-
lo e limbs, bu he majo i y a e inconsis en wi h he
g ow h habi , lea mo phology and a angemen , and
o he lo e colou a ion cha ac e s desc ibed o A c o is
s a ici olia. Such axa include Co ula L. spp., Felicia e lexa
187
Typi ica ion o A c o is species o Poi e
ADANSONIA, sé . 3 • 2019 • 41 (14)
ig. 2. — Neo ype o A c o is s a ici olia Poi . (P00126589; le specimen). Rep oduced unde he e ms o he C ea i e Commons A ibu ion License [CC BY 4.0]
(h ps://c ea i ecommons.o g/licenses/by/4.0), © Muséum na ional d’His oi e na u elle, Pa is.
188 ADANSONIA, sé . 3 • 2019 • 41 (14)
McKenzie R. J. & Nepi C.
DC., Ge be a L. spp., Lasiospe mum bipinna um (Thunb.)
D uce, Mai ia c ena a (Thunb.) Nees, Osmi opsis as e -
iscoides (P.J.Be gius) Less., O honna L. and Senecio L. spp.
wi h whi e- adia e capi ula, Poecilolepis G au spp. and
U sinia pili e a (P.J.Be gius) Poi . A o m o Felicia elonga a
O.Ho m. ex Zahlb . p oduces whi e ays ma ked pu plish
ma oon a he base, bu he species is a sh ubby pe ennial
wi h lea es no concen a ed a he base o he plan , and
he plan is coa sely hai y no glab ous.
Se e al Dimo pho heca species indigenous o he Cape e-
gion ha e capi ula wi h whi e ays, including D. plu ialis (L.)
Moench and D. nudicaulis wi h ays ma ked pu ple a he base,
and he abaxial su ace o he ay limb may be whi e, pu ple
o coppe . The in oluc al b ac s o Dimo pho heca axa a e
consis en wi h Poi e ’s desc ip ion o A. s a ici olia. Pa icu-
la ly in he axon Dimo pho heca nudicaulis a . g amini olia,
he lea es a e linea -lanceola e and c owded in a basal u .
This axon is well-illus a ed in a numbe o publica ions
(e.g. Edwa ds 1818; Loddiges & Sons 1818; Hooke 1861;
Ma lo h 1908). Dimo pho heca agus is simila in ege a i e
cha ac e is ics o hose desc ibed o A c o is s a ici olia bu
he ay lo e s a e o ange o yellow abo e and he disc lo e
co olla is blackish pu ple. The dimo phic cypselae o he disc
and ay lo e s o Dimo pho heca nudicaulis a . g amini olia
a e depic ed in he lowe igh hand co ne o he Calendula
g amini olia lec o ype illus a ion (Fig. 1).
O coinciden al in e es , Candolle (1838) lis ed “D. s a i-
ce olia Vaill. ac . pa . 1720 (nomen op .!)” in he synonymy
o Dimo pho heca g amini olia, which was consis en wi h
Linnaeus’s (1753) synonymy o Calendula g amini olia. The e
is no e idence o link Poi e ’s concep o A c o is s a ici olia
wi h he plan ha Vaillan (1720) named “Dimo pho heca
S a ices olio”. Howe e , Poi e ’s (1810) diagnosis o A c-
o is s a ici olia is simila , bu con ains addi ional de ails, o
Linnaeus’s (1753) diagnosis o Calendula g amini olia o
“Calendula oliis linea ibus subin ege imis, caule subnudo”.
No lindh (1943) designa ed as he lec o ype o Calendula
g amini olia an illus a ion in Commelin (1701: . 34) la-
belled wi h he polynomial Bellis A icana lo um pediculis
oliosis oliis angus is e in eg is. The same illus a ion and
polynomial we e ci ed by Vaillan (1720: 280) ollowing
he polynomial Dimo pho heca S a ices olio, oge he wi h
he polynomial Calendula A icana su ec a Ro is ma ini
oliis ha was lis ed and illus a ed by Plukene (1700: 35,
ab. 376, ig. 7). To ou knowledge, he binomial “Dimo -
pho heca s a ici olia” has no been alidly published.
We conclude ha Poi e ’s desc ip ion o A c o is s a ici olia
is no compa ible wi h any membe o he ibe A c o ideae
and is consis en wi h a Dimo pho heca axon, speci i-
cally Dimo pho heca nudicaulis a . g amini olia, based on
he cha ac e s speci ied in he p o ologue. The sea ch o
o iginal ma e ial o A c o is s a ici olia should con inue so
as o cla i y Poi e ’s in ended applica ion o he name. In
he absence o o iginal ma e ial, unde ICN A s 9.8 and
9.13 (Tu land e al. 2018) we he e designa e a neo ype o
A c o is s a ici olia consis en wi h ou in e p e a ion o he
p o ologue. The selec ed neo ype (P ba code P00126589;
Fig. 2) was o me ly in he He bie Poi e . The specimen
i s well wi h Poi e ’s desc ip ion o A c o is s a ici olia and
shows many o he salien cha ac e s desc ibed, namely
lea es na ow, linea , glab ous, en i e o dis an ly den a e,
he lea es inse ed only in he p oximal po ion o he s em,
capi ulum soli a y on a long naked, sho ly glandula -hai y
peduncle, and in oluc e o acu e, lanceola e b ac s wi h
±hyaline ma gins. The specimen was o iginally iden i ied
as Calendula g amini olia L. and subsequen ly was anno-
a ed “a c o is enui olia . poi . ency. sup.”, likely in Poi e ’s
handw i ing. The ga he ing is unmis akably a specimen o
Dimo pho heca nudicaulis a . g amini olia, he e o e A c o is
s a ici olia is he e placed in he synonymy o D. nudicaulis
a . g amini olia. Unde ICN A . 9.19(a), i o iginal ma e ial
o A c o is s a ici olia is loca ed a a u u e da e, he choice
o he neo ype will be supe seded by he o iginal ma e ial.
A c o is enui oliA Poi .
Linnaeus (1771) published he name A c o is enui olia L.
wi h he diagnosis “A c o is losculis adian ibus s e ilibus ?
oliis linea ibus indi isis glab is.” The p o enance o he spe-
cies was s a ed o be “Habi a in Cap. b. spei [Capu bonae
spei] li o e ma i imo.” No specimens o p e iously published
desc ip ions o illus a ions we e ci ed in he p o ologue.
The name is he basionym o he species cu en ly known
as U sinia enui olia (L.) Poi . and was lec o ypi ied o a
specimen in he Linnaean He ba ium (He b. Linn. No.
1036.24; LINN) by P assle (1967). A c o is enui olia L.
was ans e ed o U sinia by Poi e (1808), who epea ed
he abo e-men ioned diagnosis bu wi h ’A c o is’ subs i-
u ed wi h ’U sinia’. When publishing he combina ion,
Poi e ci ed he basionym A c o is enui olia L. (ci ing he
ea men s o ha species by Linnaeus 1771: 288; Linnaeus
1781: 385; Lama ck 1783: 238; Willdenow 1803: 2361).
Poi e (1810: 439) lis ed A c o is enui olia wi h a e y di e -
en diagnosis o ha used by Linnaeus (1771), namely ’A c o is
caule u icoso, glab o; oliis sub ili o mibus, glab is, apice i idis;
pedunculis longissimis, uni lo is’, and no ably speci ied ’(N.)’.
Poi e did no ci e a p e ious publica ion in which he name
A c o is enui olia was used, as he did o o he p e iously named
A c o is species, and speci ically did no ci e A. enui olia L.
no U sinia enui olia (L.) Poi . In addi ion, Poi e ci ed only
unspeci ied ma e ial in he He bie Des on aines and hus
ga e no indica ion o ha ing examined Linnaean specimens.
Gi en ha Poi e (1808) had p e iously ans e ed A c o is
enui olia L. o U sinia, and hus was clea ly amilia wi h he
Linnaean name, i is easonable o in e p e ha Poi e (1810)
in ended o publish a new name o wha he conside ed o
be an A c o is species based on ma e ial di e en o ha o A.
enui olia L. Poi e ’s wo k long p eceded he es ablishmen o a
code o ules o bo anical nomencla u e and, wi h he ans e
o A c o is enui olia L. o U sinia, Poi e migh ha e conside ed
ha he name “A c o is enui olia” was a ailable o use again.
Rega dless, i is concluded ha Poi e (1810) alidly published
a new name, A c o is enui olia Poi ., which is a he e o ypic
homonym o A. enui olia L. and hus is nomencla u ally il-
legi ima e unde ICN A . 53.1 (Tu land e al. 2018).
189
Typi ica ion o A c o is species o Poi e
ADANSONIA, sé . 3 • 2019 • 41 (14)
ig. 3. — Lec o ype o A c o is enui olia Poi . (FI053597). © Sezione di Bo anica, Museo di S o ia Na u ale-Bo anica, Uni e si à di Fi enze, Fi enze.
